Family-Group Names In Coleoptera (Insecta)

1Zoological nomenclature affects the work of all zoologists, yet only a minuscule fraction of one percent of [the craziest] zoologists deal directly with problems associated with scientific names of animals.

2The accurate use of scientific names in zoology is necessary to minimize nomenclatural instability and allow the maximal retrieval of scientific information from the ever increasing body of literature. Akin to other scientific names such as species- and genus-group names, family-group names are extremely important in the exchange of information about the world we live in. Rules about how to treat family-group names, as determined by the International Commission on Zoological Nomenclature [henceforth the Commission], have been an integral part of zoological nomenclature since the early 1900s (for a historical review of family-group names refer to ).

3The first family-group names based on the stem of their type genus appeared in zoological literature in the early 19th Century (see ). The first authors inconsistently used a variety of endings (–ides, –ites, –ida, –i, –ide, etc.) for suprageneric divisions at various levels. The ending –idae was apparently first suggested by as a characteristic ending for names at the rank of family. Although many ranks have been used in the past for suprageneric names, only the following suffixes are currently regulated by the International Code of Zoological Nomenclature [henceforth the Code]: superfamily (–oidea), family (–idae), subfamily (–inae), tribe (–ini) and subtribe (–ina).

4The last twenty five years has seen an increase in the number of studies on family-group names of entire groups in zoology (e.g., for Hemiptera: Nepomorpha; for Aves; for Mammalia; for Pisces: Siluriformes; for Diptera; for Isoptera; for Lepidoptera: Noctuoidea; for Gastropoda; for Hymenoptera: Apoidea; for Dermaptera; for Bivalvia). The contents (i.e., whether they include fossils or not) and presentation of the data (i.e., in a classification scheme, in chronological order or in alphabetic order) in those works vary greatly. The most extensive treatments of animal family-group names to date are those of Diptera and Gastropoda which covered names based on approximately 2000 and 2400 distinct type genera respectively. Some of the reasons given by authors of these works for providing comprehensive lists of family-group names include avoiding the unnecessary proposal of new names, facilitating decisions on priority and promoting long-term stability of the classification. We believe that such catalogues are especially important as the number of studies dealing with the higher relationships of major clades will continue to increase as new algorithms and data sets (e.g., molecular data) become available.

5Given the importance of family-group names in the scientific literature, it is rather surprising that still many authors do not cite the author or year of publication of those names correctly, or at all. According to Recommendation 51A of the Code “The original author and date of a name should be cited at least once in each work dealing with the taxon denoted by that name.” In addition to satisfying Recommendation 51A of the Code, citation of authorities of scientific names can also have a positive impact on modern world taxonomists. Adding to further confusion and errors in the literature is the fact that the Principle of Coordination (Art. 36.1) is still overlooked by some authors.

6Coleoptera are currently the most species-rich group of organisms on this planet with approximately 360 000 described species. The great morphological diversity of beetles has led to the proliferation of suprageneric taxa at various ranks. was apparently the first to introduce the concept of family-level taxa (see: 244) but it is only a few years later that he proposed available names for these groupings, including several in the order Coleoptera. More recent evidence suggests that available family-group names in some groups of animals other than Coleoptera (e.g., Chordata: Sauropsida) appeared in the late 18th Century (see ), even earlier than in.

7The nomenclature of family-group names in Coleoptera did not receive much attention until the treatment of Geadephaga names by. Similar lists, based on rules from two different editions of the Code, have now been published in Dytiscidae, Staphyliniformia, Cucujoidea, Curculionoidea, Buprestoidea, Tenebrionidae, Scarabaeoidea and Cerambycidae. Another important contribution was that of which included a review of the nomenclature of all beetle family-group names for the rank of subfamily and above.

8The vast body of scientific literature dealing with beetles has been a deterrent to producing a complete review of all Coleoptera family-group names in the past. This publication was only made possible by the collaboration of several coleopterists.

9The specific objectives of this study are to: 1) establish the first comprehensive list of all Coleoptera family-group names with information on type genus, author(s), year of publication and complete bibliographical references; 2) assess the availability and validity of each name using rules laid out in the most recent Code of Zoological Nomenclature, 3) summarize Priority and Homonymy problems with currently used names and, 4) propose or implement solutions to these problems in order to promote stability. We include family-group names that were published on or before December 31, 2010.

10This publication should be seen as the starting point of what could eventually serve as the basis for a submission of Part of a List of Available names in Zoology (Art. 79). With this goal in mind, we ask all coleopterists to either send us, or to publish, corrections or differences of opinions in the months and years to come. It is our intention to update our list of family-group names as new data are published and to provide a second edition of this work in approximately five years.

11Decisions about the availability and validity of each Coleoptera family-group name in our catalogue were made following the process outline in Figure 1. First we established whether the name was available or unavailable based on the criteria of availability summarized below. If a name first proposed on a particular type genus was determined to be unavailable then searches were conducted to establish if a family-group name based on the same type genus was made available subsequently. If so then this available name was entered in our catalogue. From the pool of available names, we removed those that are permanently invalid (Art. 39). All the remaining available names were then separated into those that are valid based on the classification used here (at any rank from subtribe to superfamily) and those that are invalid (i.e., synonyms).

12In order to be available, a family-group name proposed before 1931 must to be a scientific name (i.e., in latinized form) in the nominative plural based on the stem of an available genus name then used as valid in the new suprageneric taxon (Art. 11.7).

13In addition to the criteria of availability mentioned above, new family-group names proposed between 1931 and 1999 had to be described in words, or be associated with a bibliographic reference to such a description, in order to be considered available (Arts 13, 15). In works published after 1930 which contained more than one use of a new family-group name, we selected a page where a description was clearly associated with the new taxon for the catalogue. It should be noted that replacement names proposed in that time period are available without description (Art. 13.1.3). Additionally, “a family-group name first published after 1930 and before 1961 which does not satisfy the provisions of Article 13.1 is available from its original publication only if it was used as valid before 2000, and also was not rejected by an author who, after 1960 and before 2000, expressly applied Article 13 of the then current editions of the Code” (Art. 13.2.1). Names proposed between 1931 and 1960 can also be considered available if the description of the new family-group name and a single new genus-group name is combined (Art. 13.5). Finally, family-group names proposed since 2000 have to be explicitly indicated as new and the name of the type genus has to be clearly cited in order to be available (Art. 16). Based on our interpretation of Article 16.2, any new family-group name proposed after 1999 in a paper in which the name of the type genus is clearly cited in connection with it (although maybe not explicitly with a formula such as “Type genus = Aus Doe, 2010”) is available.

14One of the most difficult tasks while working on this review was to decide on the most consistent and objective way to apply Article 11.7.2, which deals with the availability of names that were originally proposed in a vernacular form. Vernacular names are generally not treated as scientific names in zoological nomenclature (see Recommendation 11A). However, the International Commission on Zoological Nomenclature has made a single, but rather restricted, exception that applies to family-group names. According to the Code, a vernacular name “…is available with its original author and date only if it has been latinized by later authors and has been generally accepted as valid by authors interested in the group concerned and as dating from that first publication in vernacular form” (Art. 11.7.2).

15We have interpreted names proposed in the following languages to be vernacular: all German names with the suffix “–en”, all Spanish names with the suffix “–os” or “–as” and all French names with the suffix “–iens”. The most important issue was to determine the correct status of names originally proposed with the suffix “–es.” We have used the principle that all family-group names proposed by non-French writers with the suffix “–es” were in fact latinized names with an ending that differs from those regulated by the Commission (these are mostly older names proposed before rules of zoological nomenclature became well-established). For each name with the suffix “–es” first proposed by French workers (e.g., Latreille, Lacordaire, Lameere) we went through the entire work containing those names and established if these authors consistently used either vernacular or latinized names in their work. French vernacular names often have accents in them (“é” or “è”) while latinized names do not. We have found that determining whether an author used vernacular or latinized names in a particular work was fairly straightforward. Latreille proposed new family-group names with the suffix “–es” in several of his works. We noticed that he most often used both the vernacular (listed first, with accents when required by French language) and latinized (listed second, always in italics) forms of each name together in the same heading. We have listed the latinized form of the name in each of those cases.

16As in, we have interpreted the requirements listed in Art. 11.7.2. as three separate conditions to be met. For each vernacular name we determined if it had 1) been subsequently latinized, 2) been generally accepted as valid by authors interested in the group and 3) been attributed to the author and date of original publication. Failure to fulfil any of the requirements resulted in the treatment of that name as unavailable. In those cases, we provide a comment explaining why this taxon was treated as unavailable (e.g., “original vernacular name unavailable (Art. 11.7.2): not subsequently latinized”). Many recent authors have treated family-group names first proposed in vernacular form as available if they were latinized by later authors but we do not believe that this practice is in line with the requirements of the Code. We have accepted as available all vernacular names published before 1900 that have subsequently been used in latinized form, while being used as valid, and credited to the publication in their vernacular form. For every vernacular name that meets the requirement of availability, we have added a comment in the format of the following example “original vernacular name available (Art. 11.7.2): first used in latinized form by, as Meracanthini), generally accepted as in, as Meracanthinae)”.

17Article 29.2 includes a list of suffixes to be used for groups at the superfamily (-oidea), family (-idae), subfamily (-inae), tribe (-ini) and subtribe (-ina) names. The suffix of other categories is not regulated by the Code. The rank of supertribe (-itae) was used here for a small number of groups in which recent changes in classification required an additional rank between subfamily and tribe (e.g., Staphylinoidea).

18The correct spelling of family-group names depends on the stem of its type genus (Art. 29.3). As pointed out by the family-group name stems of most generic names that can be regarded as Latin nouns are determined easily by dropping the following nominative case ending –us, –um, –es and –a. Examples of such straightforward case in Coleoptera include Carabidae (type genus Carabus), Adeliini (type genus Adelium), Trechodina (type genus Trechodes) and Anthiini (type genus Anthia). A summary of other commonly encountered generic endings, along with their corresponding stems, is given in Table 1. Note that generic names ending with -gaster (Greek for stomach) can have either -gaster- or -gastr- as their correct stem and therefore we have accepted the stem formation of the first author of the family-group name based on such genera as correct. Stems were reviewed for all type genera included in our catalogue.

19It should be noted that if a family-group name was not formed in accordance with Art. 29.3 but its original spelling is in prevailing usage then the current spelling is to be maintained (Art. 29.5). We have conserved the spelling of several family-group names currently used as valid however we did not do so for names that are listed as synonyms. For names based on incorrect stems proposed after 1999, we have considered that prevailing usage cannot be used to conserve the original spellings because too few references using these names could be found. We have therefore corrected the stems of such names unless the name of the type genus was an arbitrary combination of letters (Art. 29.4).

20As stated in Art. 35.4.1 “A family-group name based upon an unjustified emendation … or an incorrect spelling of the name of the type genus must be corrected, unless it is preserved under Article 29.5 or unless the spelling of the genus-group name used to form the family-group name is preserved under Articles 33.2.3.1 or 33.3.1.” When an unjustified emendation or an incorrect subsequent spelling of the type genus is in prevailing usage and is attributed to the original author and date (Art. 33.2.3.1; 33.3.1), the correct spelling of the type genus is that in current usage.

21In the glossary of the Code, a name in prevailing usage is defined as a “name which is adopted by at least a substantial majority of the most recent authors concerned with the relevant taxon, irrespective of how long ago their work was published”. The unfortunate subjectivity in this definition, as pointed out by, left us with no choice but determine prevailing usage in an ad hoc fashion throughout.

22The Principle of Coordination (Art. 36.1) is, unfortunately, still overlooked by some authors. In some instances authors who propose new ranks for previously established suprageneric names are sometimes treated as the author of those names when in fact only a change of rank was presented. Based on the Code “A name established at any rank in the family-group is deemed to have been simultaneously established for nominal taxa at all ranks in the family-group; all these taxa have the same type genus, and their names are formed from the stem of the name of the type genus [Art. 29.3] with appropriate change of suffix [Art. 34.1]. The name has the same authorship and date at every rank.”

23As for species- and genus-group names, the oldest available name for a family-group taxon should be considered as valid (Art. 23). However four important exceptions need further discussion. Firstly, when a little-known family-group name was discovered to be older than a name currently used as valid for a particular taxon, we used the Reversal of Precedence to conserve usage of the younger name if the conditions of Art. 23.9.2 could be fulfilled. Younger names conserved using Reversal of Precedence are listed in Appendix 1. In some instances we could not fulfill all conditions of Art. 23.9.2 although we considered that using the newly discovered older name as valid would threaten stability or cause confusion. In those cases we maintained usage of the younger name as valid and either submitted an application to the Commission to conserve the younger name or made a recommendation that such submission should be submitted in the near future (Art. 23.9.3).

24According to Art. 35.5 “If after 1999 a name in use for a family-group taxon (e.g., for a subfamily) is found to be older than a name in prevailing usage for a taxon at higher rank in the same family-group taxon (e.g., for the family within which the older name is the name of a subfamily) the older name is not to displace the younger name.” Indeed, we encountered a small number of cases in which the replacement of a name at the higher rank (e.g., Lymexyloidea Fleming, 1821) by the discovery of an older name for a taxon at a lower rank (e.g., Hylecoetoidea Germar, 1818) would not have served the stability of well-established names. In such cases usage of the younger name at the higher rank was conserved.

25In cases where a family-group name was replaced before 1961 because of the synonymy of its type genus (e.g., Lepiceridae Hinton, 1936 instead of Cyathoceridae Sharp, 1882), the substitute name is to be maintained if it is in prevailing usage (Art. 40.2). In such cases, the valid family-group name retains its own author but takes the priority of the replaced name. Based on Recommendation 40A, we have cited those names with their original author and date, followed by the date of its priority enclosed in parentheses, as determined by Art. 40.2.1 (e.g., Lepiceridae Hinton, 1936 (1882)).

26Lastly, if the stability and continuity of the meaning of a family-group name is threatened by the discovery that the type genus was originally misidentified, or that the type genus was based on a misidentified type species, or that an older type species of the type genus had been overlooked, then the case is to be referred to the Commission for a ruling (Art. 65.2.1). In cases where the oldest available name for a family-group taxon is based on misidentified type genus, or an altered concept of the type genus (e.g., see Scydmaenini Reitter, 1882), we have preferred to consider this name as invalid and to use instead the family-group name which is in prevailing usage (e.g., Cyrtoscydmini L. W. Schaufuss, 1889) until an application is submitted and a ruling is rendered by the Commission.

27A list of problem cases based on the Principle of Priority, with comments on implementation of solutions or necessary actions to be taken in the future, is given in Appendix 2.

28Based on the Principle of Homonymy “when two or more names are homonyms, only the senior, as determined by the Principle of Priority…, may be used as the valid name…” (Art. 55.2). Here we report several instances in which family-group names in Coleoptera are identical to other family-group names in zoological nomenclature. These names are either based on identical type genera or on type genera that are similar but not identical. In the first instance the family-group name based on the preoccupied type genus is permanently invalid (Art. 39) but available.

29When family-group names are homonyms because their type genera are similar but not identical, the case must be referred to the Commission for a ruling to remove homonymy (Art. 55.3.1). Such is the case with Adeliini Kirby, 1825 (type genus Adelium Kirby, 1819) and the hymenopteran name Adeliini Viereck, 1918 (type genus Adelius Haliday, 1833) which are both correctly formed from the stem of their type genus. More than thirty such cases were encountered during our research on Coleoptera family-group names.

30Junior homonyms conserved using Reversal of Precedence are listed in Appendix 1. A list of problem cases based on the Principle of Homonymy, with comments on implementation of solutions or necessary actions to be taken in the future, is given in Appendix 3.

31As mentioned above, we have encountered several cases which require an application to the Commission because of problems with priority and/or homonymy. Some of these cases have been submitted recently and others will be submitted in the near future (see summary in Appendices 2 and 3). However, it is not our intention to submit applications for all cases. Since we intend to submit this work as Part of the List of Available Names in Zoology in approximately five years, we hope the coleopterist community will take this opportunity to submit applications to the Commission in order to resolve some of the remaining problems outlined here.

32: 282, 290) and: 157) reported that the important works on Elateridae classification by and were published in May and June of 1857 respectively. Since then, when the same new name appeared in both of these works, Candèze’s names have been given priority over Lacordaire’s. We have discovered that Lacordaire’s work was in fact published before 25 May 1857 (as recorded by the Académie des Sciences de France) which would make the names in his work the oldest based on strict adherence to the Code (Art. 21.3). In order to avoid unnecessary changes in authorship in the future and maintain prevailing usage, we have opted to continue attributing elaterid family-group names to.

33Authorship of the three volumes on Coleoptera of the series Encyclopédie d’histoire naturelle is debated. The title pages on the volumes list Jean Charles Chenu as the author of the Encyclopédie d’histoire naturelle with the assistance of Eugène Desmarest for the Coleoptera section. However, the Société Entomologique de France recorded the livraisons received for 1851 in their Bulletin (p. cxxv) as “Encyclopédie d’histoire naturelle, ou traité complet de cette science, sous la direction de M. le docteur Chenu. Coléoptères, par M.E. Desmarest.” This was likely the title on the wrapper and clearly suggests that Desmarest was responsible alone for the three volume series on Coleoptera. This is also substained by Desmarest himself who in the Bulletin de la Société Entomologique de France for 1860 (p. lxiii) speaks of the three volumes series as being his work, published under the direction of Chenu. Based on the above fact, we credit the three-volume series to Desmarest alone.

34The articles presented by Mulsant (and Rey) in the “Opuscules Entomologiques” are considered here to be reprints (as a compilation) of original publications in various Annales, mostly printed at Lyon, for the following reasons: In the dedication to the first volume Mulsant wrote “ces Opuscules, publiés déjà ça et là dans nos Recueils académiques”. The dates of the dedications in the Opuscules are always later than the dates on which they were presented to the various Sociétés, frequently in the following year. Although published at Paris, the Opuscules (see verso of title pages) were printed at Lyon by F. Dumoulin and others, the same printers which published the Mémoires de l’Académie... des Sciences, Belles-Lettres et Arts de Lyon, etc. Rey himself, in later publications such as the continuation of the “Histoire Naturelle des Coléoptères de France” (e.g., Annales de la Société Linnéenne de Lyon (N. S.) 32[1885]: 1-186 + [4], pls. 1-2 [see p. 106]) cited the Annales de la Société Linnéenne de Lyon article before the Opuscules version. E. A.: 46-47), in his obituary of Mulsant, indicated that both the Histoire Naturelle des Coléoptères de France and the Opuscules Entomologiques «appeared originally... in the Annals» and «later republished in separate form at Paris». These concepts were reiterated by: 352).: 20) wrote, “Tous les ouvrages de Mulsant, sauf quelques-uns, ont été publiés dans les Annales des trois sociétés... et l’Académie des Sciences. Tous ont été reproduits séparément soit dans l’Histoire Naturelle des Coléoptères et des Punaises de France, soit dans les Opuscules.»

35However, the first fascicles of the Histoire Naturelle series up to the “Pectinipèdes” were not reprinted in journals. It seems that most of the succeeding fascicles were submitted to the various Sociétés, were printed by Dumoulin, Barret, Pinier, etc. at Lyon, the dedications were then added and they were sent to Paris for separate publication with the same typesetting, and with changes only to the pagination and some titles. Since the issuing of the journal volumes was frequently delayed by the compilation of many papers (usually in the following year), the separate edition often appeared first, as shown by records of other journals and, in particular, the Bibliographie de la France. Therefore we have given date priority to the Histoire Naturelle versions in most instances, only those page numbers are cited with the scientific names in the catalogue unless evidence was found to the contrary, and the alternate versions are cited only in the bibliography, following each reference.

36This paper is organized into two main parts: 1) a synoptic classification containing only the names which we consider as valid herein and 2) a catalogue of all family-group names organized in the same order as in the synoptic classification. It should be noted that our publication is first and foremost a nomenclatural treatment of family-group names. We want to emphasize that the classification used here is not based on newly generated phylogenetic data. Furthermore, we do not necessarily endorse all parts of the classification presented, particularly those which were not based on phylogenetic approaches. For the most part, we follow currently accepted concepts in recent taxon-specific catalogues as well as comprehensive syntheses such as the Handbook of Zoology, American Beetles and the Catalogue of Palaearctic Coleoptera. Although we recognize that the classification we use will likely become outdated in the near future, we believe that this is the best way to present the assembled data.

37Only family-group names that are based on genera are included in our catalogue. For each family-group name the original spelling, author, year of publication, page number, correct stem and type genus are given. Complete data and comments regarding a particular family-group name are presented with the lowest-rank name when the same stem is used in more than one rank, since the same criteria apply in accordance with the Principle of Coordination (Art. 36.1). The correct stem to be used, which is given in square brackets (e.g., [stem: Uralocole-]), is especially important for synonyms, since the status of a name may be changed from synonym to valid in the future. Subsequent alternative spellings of family-group names (i.e., simple changes in suffix depending on change in rank) are not given here. Both available and unavailable names are listed together in the catalogue in order to enhance information retrieval. Unavailable names are preceded by an asterisk “*” and are identified as such in the “Comments” section along with the reason for this status. As is common practice in publications of the International Commission on Zoological Nomenclature, we use small capital letters for family-group names. Names in the catalogue are organized in a “phylogenetic” framework, as accepted in recent publications on the group, down to subfamily level. Valid tribes and subtribes are listed in alphabetical order within each subfamily. Synonyms are listed in chronological order under each valid taxon.

38Known works of Johannes Gistel (who also published under the name Johannes Gistl and as G. Tilesius; see ) were included in this paper. Although Gistel’s works were largely ignored by contemporaries (e.g., ) and continue to be ignored by some workers (e.g., ) they are nevertheless broadly available and represent a significant contribution to the nomenclature of family-group and genus-group names in Coleoptera. In order to promote stability, some names proposed for the first time by Gistel (but ignored until now) that threaten names in current usage are treated here as nomina oblita (when conditions of Art. 23.9 could be met) or are subject to appeals to the Commission. According to the glossary of the Code, a nomen oblitum is a “Latin term applied after 1 January 2000 to a name, unused since 1899, which as a result of an action taken under Article 23.9.2 does not take precedence over a younger synonym or homonym in prevailing usage. The term nomen oblitum was also applied to a disused senior synonym rejected between 6 November 1961 and 1 January 1973 under Art. 23b of the Code editions then in force (see Art. 23.12.2). Nomina oblita are available names; see Articles 23.9 and 23.12 for conditions controlling their use as valid names.”

39Because of the importance of the Principle of Priority, we have tried to find the most accurate date of publication (given in square brackets in the References section) for works cited in the manuscript. The date of publication was determined either from the original publication itself, from the date of “stamps” when received in libraries of natural history institutions or from secondary literature sources. A list of natural history institutions and secondary literature sources is given at the beginning of the References section. Data on dates of publication were provided by AFN, PB, AED, MAAZ, CHCL, ABTS and YB. References for type genera are not included here to conserve space and because they are for the most part available in other recent publications (e.g.,,,,,, ).

40Names on the Official List of Family-Group Names in Zoology and type genera on the Official List of Generic Names in Zoology are included in our catalogue and are summarized in Appendix 4 and 5 respectively. Cases involving family-group names and/or their type genera that are awaiting a ruling by the Commission are summarized in Appendix 6.

41A total of 4887 family-group names are included in our catalogue (see Figure 1). Of the names recorded, 4492 are available, of which 183 are permanently invalid because they are based on a preoccupied type genus or a genus which has been suppressed by the Commission. The majority of names were proposed in the suborder Polyphaga (4314) followed by Adephaga (531), Myxophaga (16), Archostemata (17) and Protocoleoptera (9). Within Adephaga the family Carabidae contains the highest number of names by far (441) followed by Dytiscidae (47). The five superfamilies with the highest number of names in Polyphaga are the Curculionoidea (862), Chrysomeloidea (794), Staphylinoidea (594), Tenebrionoidea (579) and Scarabaeoidea (436). Overall, the five families with the highest number of family-group names proposed are Curculionidae (555), Staphylinidae (493), Cerambycidae (468), Carabidae (441) and Tenebrionidae (323).

42The number of Coleoptera family-group names that appeared in the 19th and 20th Centuries are almost identical (2331 and 2556 respectively) while the last decade saw an increase of 5% (246 names) in the total number of names in the literature (Figure 2). A large number of names proposed in the middle of the 19th Century are unavailable. Those names were generally proposed in vernacular form and were not made available subsequently (Art. 11.7.2). Unavailable names proposed after 1930 generally lacked a description or bibliographic reference to such a description (Arts 13, 15). Several names proposed in the last decade were either not explicitly introduced as new taxa or did not include necessary information about the type genus and are therefore unavailable (Art. 16).

43A summary of the most significant contributions, in terms of the total number of names proposed by author, is shown in Table 2. Eighteen authors proposed 50 or more family-group names in Coleoptera. Lacordaire was the most prolific with 353 new names. Although the majority of Lacordaire’s names were proposed in vernacular form, a large proportion of them (90%) were subsequently made available and many are used as valid today.

44The authors with the lowest percentage of available family-group names (Mulsant, Chapuis, Rey and Blanchard) originally proposed their names in vernacular form and a significant proportion of those names were not made available subsequently. All names proposed by LeConte, Reitter, Casey, Horn, Legalov, C. G. Thomson and Laporte are considered available.

45Thirteen of the authors in Table 2 were active exclusively during the 19th Century while two authors, Reitter and Casey, published their works containing new family-group names in both the 19th and 20th Centuries. Two authors published family-group names exclusively during the 20th Century (Jeannel and Voss). Legalov is our only contemporary colleague in this list; he proposed more than 50 new family-group names in the last decade. It should be noted that the recent proliferation of new scientific names proposed by Legalov is treated by some authors as “extreme splitting” and “based on numerous spurious characters of doubtful phylogenetic value”.

46The broad taxonomic interests and expertise of Lacordaire, Gistel, LeConte, Mulsant, Latreille and C. G. Thomson are exemplified by the fact that they introduced new family-group names in 25 or more families. This was done either in catalogues (e.g., Gistel, C. G. Thomson) and/or in large scale taxonomic treatments (e.g., Lacordaire, LeConte, Latreille, Mulsant). On the other hand, the taxonomic expertise of J. Thomson, Chapuis, Jeannel, Voss, Burmeister and Legalov is much more targeted towards a small number of families (fewer than 10).

47The following synonymies are recorded here for the first time: Agronomina Gistel, 1848 syn. nov. of Amarina Zimmermann, 1832 (Carabidae), Hylepnigalioini Gistel, 1856 syn. nov. of Melandryini Leach, 1815 (Melandryidae), Polycystophoridae Gistel, 1856 syn. nov. of Malachiinae Fleming, 1821 (Melyridae), Sclerasteinae Gistel, 1856 syn. nov. of Ptilininae Shuckard, 1839 (Ptinidae), Phloeonomini Ádám, 2001 syn. nov. of Omaliini MacLeay, 1825 (Staphylinidae), Sepedophilini Ádám, 2001 syn. nov. of Tachyporini MacLeay, 1825 (Staphylinidae), Phibalini Gistel, 1856 syn. nov. of Cteniopodini Solier, 1835 (Tenebrionidae); Agronoma Gistel 1848 (type species Carabus familiaris Duftschmid, 1812, designated herein) syn. nov. of Amara Bonelli, 1810 (Carabidae), Hylepnigalio Gistel, 1856 (type species Chrysomela caraboides Linnaeus, 1760, by monotypy) syn. nov. of Melandrya Fabricius, 1801 (Melandryidae), Polycystophorus Gistel, 1856 (type species Cantharis aeneus Linnaeus, 1758, designated herein) syn. nov. of Malachius Fabricius, 1775 (Melyridae), Sclerastes Gistel, 1856 (type species Ptilinus costatus Gyllenhal, 1827, designated herein) syn. nov. of Ptilinus Geoffroy, 1762 (Ptinidae), Paniscus Gistel, 1848 (type species Scarabaeus fasciatus Linnaeus, 1758, designated herein) syn. nov. of Trichius Fabricius, 1775 (Scarabaeidae), Phibalus Gistel, 1856 (type species Chrysomela pubescens Linnaeus, 1758, by monotypy) syn. nov. of Omophlus Dejean, 1834 (Tenebrionidae). The following replacement name is used here for the first time: Gompeliina Bouchard, 2011 nom. nov. for Olotelina Báguena Corella, 1948 (Aderidae).

48Reversal of Precedence (Article 23.9) is used to conserve usage of the following names (family-group names followed by genus-group names): Perigonini Horn, 1881 nom. protectum over Trechicini Bates, 1873 nom. oblitum (Carabidae), Anisodactylina Lacordaire, 1854 nom. protectum over Eurytrichina LeConte, 1848 nom. oblitum (Carabidae), Smicronychini Seidlitz, 1891 nom. protectum over Desmorini LeConte, 1876 nom. oblitum (Curculionidae), Bagoinae Thomson, 1859 nom. protectum over Lyprinae Gistel 1848 nom. oblitum (Curculionidae), Aterpina Lacordaire, 1863 nom. protectum over Heliomenina Gistel, 1848 nom. oblitum (Curculionidae), Naupactini Gistel, 1848 nom. protectum over Iphiini Schönherr, 1823 nom. oblitum (Curculionidae), Cleonini Schönherr, 1826 nom. protectum over Geomorini Schönherr, 1823 nom. oblitum (Curculionidae), Magdalidini Pascoe, 1870 nom. protectum over Scardamyctini Gistel, 1848 nom. oblitum (Curculionidae), Agrypninae/-ini Candèze, 1857 nom. protecta over Adelocerinae/-ini Gistel, 1848 nom. oblita and Pangaurinae/-ini Gistel, 1856 nom. oblita (Elateridae), Prosternini Gistel, 1856 nom. protectum over Diacanthini Gistel, 1848 nom. oblitum (Elateridae), Calopodinae Costa, 1852 nom. protectum over Sparedrinae Gistel, 1848 nom. oblitum (Oedemeridae), Adesmiini Lacordaire, 1859 nom. protectum over Macropodini Agassiz, 1846 nom. oblitum (Tenebrionidae), Bolitophagini Kirby, 1837 nom. protectum over Eledonini Billberg, 1820 nom. oblitum (Tenebrionidae), Throscidae Laporte, 1840 nom. protectum over Stereolidae Rafinesque, 1815 nom. oblitum (Throscidae) and Lophocaterini Crowson, 1964 over Lycoptini Casey, 1890 nom. oblitum (Trogossitidae); Monotoma Herbst, 1799 nom. protectum over Monotoma Panzer, 1792 nom. oblitum (Monotomidae); Pediacus Shuckard, 1839 nom. protectum over Biophloeus Dejean, 1835 nom. oblitum (Cucujidae), Pachypus Dejean, 1821 nom. protectum over Pachypus Billberg, 1820 nom. oblitum (Scarabaeidae), Sparrmannia Laporte, 1840 nom. protectum over Leocaeta Dejean, 1833 nom. oblitum and Cephalotrichia Hope, 1837 nom. oblitum (Scarabaeidae).

49Order Coleoptera

50†Suborder PROTOCOLEOPTERA

51†Superfamily Tshekardocoleoidea Rohdendorf, 1944

52†Family Tshekardocoleidae Rohdendorf, 1944

53†Family Labradorocoleidae Ponomarenko, 1969

54†Family Oborocoleidae Kukalová, 1969

55†Superfamily Permocupedoidea Martynov, 1933

56†Family Permocupedidae Martynov, 1933

57†Family Taldycupedidae Rohdendorf, 1961

58†Superfamily Permosynoidea Tillyard, 1924

59†Family Ademosynidae Ponomarenko, 1968

60†Family Permosynidae Tillyard, 1924

61Suborder ARCHOSTEMATA

62Family Crowsoniellidae Iablokoff-Khnzorian, 1983

63Family Cupedidae Laporte, 1836

64Subfamily Priacminae Crowson, 1962

65†Subfamily Mesocupedinae Ponomarenko, 1969

66Subfamily Cupedinae Laporte, 1836

67Family Micromalthidae Barber, 1913

68Family Ommatidae Sharp and Muir, 1912

69†Subfamily Brochocoleinae Hong, 1982

70Subfamily Tetraphalerinae Crowson, 1962

71Subfamily Ommatinae Sharp and Muir, 1912

72†Tribe Lithocupedini Ponomarenko, 1969

73†Tribe Notocupedini Ponomarenko, 1966

74Tribe Ommatini Sharp and Muir, 1912

75Family Jurodidae Ponomarenko, 1985

76†Family Triadocupedidae Ponomarenko, 1966

77†Family Magnocoleidae Hong, 1998

78†Family Obrieniidae Zherikhin and Gratshev, 1994

79†Subfamily Kararhynchinae Zherikhin and Gratshev, 1994

80†Tribe Kararhynchini Zherikhin and Gratshev, 1994

81†Tribe Kenderlykaini Legalov, 2009

82†Subfamily Obrieniinae Zherikhin and Gratshev, 1994

83Suborder MYXOPHAGA

84†Superfamily Asiocoleoidea Rohdendorf, 1961

85†Family Asiocoleidae Rohdendorf, 1961

86†Family Tricoleidae Ponomarenko, 1969

87†Superfamily Rhombocoleoidea Rohdendorf, 1961

88†Family Rhombocoleidae Rohdendorf, 1961

89†Superfamily Schizophoroidea Ponomarenko, 1968

90†Family Schizophoridae Ponomarenko, 1968

91†Family Catiniidae Ponomarenko, 1968

92†Family Schizocoleidae Rohdendorf, 1961

93Superfamily Lepiceroidea Hinton, 1936 (1882)

94Family Lepiceridae Hinton, 1936 (1882)

95Superfamily Sphaeriusoidea Erichson, 1845

96Family Torridincolidae Steffan, 1964

97Subfamily Torridincolinae Steffan, 1964

98Subfamily Deleveinae Endrödy-Younga, 1997

99Family Hydroscaphidae LeConte, 1874

100Family Sphaeriusidae Erichson, 1845

101Suborder ADEPHAGA

102†Family Tritarsidae Hong, 2002

103Family Gyrinidae Latreille, 1810

104Subfamily Spanglerogyrinae Folkerts, 1979

105Subfamily Gyrininae Latreille, 1810

106Tribe Enhydrini Régimbart, 1882

107Subtribe Dineutina Desmarest, 1851

108Subtribe Enhydrina Régimbart, 1882

109Tribe Gyrinini Latreille, 1810

110Subtribe Gyrinina Latreille, 1810

111Subtribe Heterogyrina Brinck, 1956

112Tribe Orectochilini Régimbart, 1882

113Family Trachypachidae Thomson, 1857

114†Subfamily Eodromeinae Ponomarenko, 1977

115Subfamily Trachypachinae Thomson, 1857

116Family Rhysodidae Laporte, 1840

117Tribe Leoglymmiini Bell and Bell, 1978

118Tribe Dhysorini Bell and Bell, 1978

119Tribe Medisorini Bell and Bell, 1987

120Tribe Rhysodini Laporte, 1840

121Tribe Clinidiini Bell and Bell, 1978

122Tribe Omoglymmiini Bell and Bell, 1978

123Tribe Sloanoglymmiini Bell and Bell, 1991

124Family Carabidae Latreille, 1802

125†Subfamily Protorabinae Ponomarenko, 1977

126†Subfamily Conjunctiinae Ponomarenko, 1977

127Subfamily Nebriinae Laporte, 1834

128Tribe Nebriini Laporte, 1834

129Tribe Notiokasiini Kavanaugh and Nègre, 1983

130Tribe Notiophilini Motschulsky, 1850

131Tribe Opisthiini Dupuis, 1912

132Tribe Pelophilini Kavanaugh, 1996

133Subfamily Cicindinae Csiki, 1927

134Subfamily Cicindelinae Latreille, 1802

135Tribe Amblycheilini Csiki, 1903

136Tribe Cicindelini Latreille, 1802

137Subtribe Apteroessina Rivalier, 1971

138Subtribe Cicindelina Latreille, 1802

139Subtribe Dromicina Thomson, 1859

140Subtribe Iresiina Rivalier, 1971

141Subtribe Theratina Horn, 1893

142Tribe Collyridini Brullé, 1834

143Subtribe Collyridina Brullé, 1834

144Subtribe Tricondylina Naviaux, 1991

145Tribe Ctenostomatini Laporte, 1834

146Tribe Manticorini Laporte, 1834

147Tribe Megacephalini Laporte, 1834

148Subfamily Carabinae Latreille, 1802

149Tribe Carabini Latreille, 1802

150Tribe Ceroglossini Lapouge, 1927

151Tribe Cychrini Perty, 1830

152Tribe Pamborini Hope, 1838

153Subfamily Loricerinae Bonelli, 1810

154Subfamily Omophroninae Bonelli, 1810

155Subfamily Elaphrinae Latreille, 1802

156Subfamily Migadopinae Chaudoir, 1861

157Tribe Amarotypini Erwin, 1985

158Tribe Migadopini Chaudoir, 1861

159Subtribe Aquilicina Moret, 2005

160Subtribe Migadopina Chaudoir, 1861

161Subfamily Hiletinae Schiødte, 1848

162Subfamily Scaritinae Bonelli, 1810

163Tribe Carenini MacLeay, 1887

164Tribe Clivinini Rafinesque, 1815

165Subtribe Ardistomina Putzeys, 1867

166Subtribe Clivinina Rafinesque, 1815

167Subtribe Forcipatorina Bänninger, 1938

168Tribe Dalyatini Mateu, 2002

169Tribe Dyschiriini Kolbe, 1880

170†Tribe Palaeoaxinidiini McKay, 1991

171Tribe Pasimachini Putzeys, 1867

172Tribe Promecognathini LeConte, 1853

173Tribe Salcediini Alluaud, 1930 (1929)

174Subtribe Androzelmina Bell, 1998

175Subtribe Salcediina Alluaud, 1930 (1929)

176Subtribe Solenogenyina Bell, 1998

177Tribe Scaritini Bonelli, 1810

178Subtribe Acanthoscelina Csiki, 1927

179Subtribe Corintascarina Basilewsky, 1973

180Subtribe Dyscherina Basilewsky, 1973

181Subtribe Ochyropina Basilewsky, 1973

182Subtribe Oxylobina Andrewes, 1929

183Subtribe Scapterina Putzeys, 1867

184Subtribe Scaritina Bonelli, 1810

185Subtribe Storthodontina Jeannel, 1946

186Subfamily Broscinae Hope, 1838

187Tribe Broscini Hope, 1838

188Subtribe Axonyina Roig-Juñent, 2000

189Subtribe Baripodina Jeannel, 1941

190Subtribe Broscina Hope, 1838

191Subtribe Creobiina Jeannel, 1941

192Subtribe Nothobroscina Roig-Juñent, 2000

193Subfamily Apotominae LeConte, 1853

194Subfamily Siagoninae Bonelli, 1813

195Tribe Enceladini Horn, 1881

196Tribe Lupercini Lecordier, 1977

197Tribe Siagonini Bonelli, 1813

198Subfamily Melaeninae Csiki, 1933

199Subfamily Gehringiinae Darlington, 1933

200Tribe Gehringiini Darlington, 1933

201Subtribe Gehringiina Darlington, 1933

202Subtribe Helenaeina Deuve, 2007

203Subfamily Trechinae Bonelli, 1810

204Tribe Bembidiini Stephens, 1827

205Subtribe Anillina Jeannel, 1937

206Subtribe Bembidiina Stephens, 1827

207Subtribe Tachyina Motschulsky, 1862

208Subtribe Xystosomina Erwin, 1994

209Tribe Horologionini Jeannel, 1949

210Tribe Pogonini Laporte, 1834

211Tribe Trechini Bonelli, 1810

212Subtribe Aepina Fowler, 1887

213Subtribe Cnidina Jeannel, 1958

214Subtribe Perileptina Sloane, 1903

215Subtribe Plocamotrechina Jeannel, 1960

216Subtribe Trechina Bonelli, 1810

217Subtribe Trechodina Jeannel, 1926

218Tribe Zolini Sharp, 1886

219Subtribe Chalteniina Roig-Juñent and Cicchino, 2001

220Subtribe Sinozolina Deuve, 1997

221Subtribe Zolina Sharp, 1886

222Subfamily Patrobinae Kirby, 1837

223Tribe Lissopogonini Zamotajlov, 2000

224Tribe Patrobini Kirby, 1837

225Subtribe Deltomerina Chaudoir, 1871

226Subtribe Deltomerodina Zamotajlov, 2002

227Subtribe Patrobina Kirby, 1837

228Subtribe Platidiolina Zamotajlov and Lafer, 2001

229Subfamily Psydrinae LeConte, 1853

230Tribe Amblytelini Blackburn, 1892

231Tribe Mecyclothoracini Jeannel, 1940

232Tribe Meonini Sloane, 1898

233Tribe Moriomorphini Sloane, 1890

234Tribe Psydrini LeConte, 1853

235Tribe Tropopterini Sloane, 1898

236Subfamily Nototylinae Bänninger, 1927

237Subfamily Paussinae Latreille, 1806

238Tribe Metriini LeConte, 1853

239Tribe Mystropomini Horn, 1881

240Tribe Ozaenini Hope, 1838

241Tribe Paussini Latreille, 1806

242†Subtribe Arthropteritina Luna de Carvalho, 1961

243Subtribe Carabidomemnina Wasmann, 1928

244Subtribe Cerapterina Billberg, 1820

245†Subtribe Eopaussina Luna de Carvalho, 1951

246Subtribe Heteropaussina Janssens, 1950

247Subtribe Homopterina Wasmann, 1920

248Subtribe Paussina Latreille, 1806

249Subtribe Pentaplatarthrina Jeannel, 1946

250Tribe Protopaussini Gestro, 1892

251Subfamily Brachininae Bonelli, 1810

252Tribe Brachinini Bonelli, 1810

253Subtribe Aptinina Gistel, 1848

254Subtribe Brachinina Bonelli, 1810

255Subtribe Mastacina Erwin, 1970

256Subtribe Pheropsophina Jeannel, 1949

257Tribe Crepidogastrini Jeannel, 1949

258Subfamily Harpalinae Bonelli, 1810

259Tribe Abacetini Chaudoir, 1873

260Tribe Amorphomerini Sloane, 1923

261Tribe Anthiini Bonelli, 1813

262Tribe Atranini Horn, 1881

263Tribe Bascanini Basilewsky, 1953

264Tribe Calophaenini Jeannel, 1948

265Tribe Catapieseini Bates, 1882

266Tribe Chaetodactylini Tschitschérine, 1903

267Tribe Chaetogenyini Emden, 1958

268Tribe Chlaeniini Brullé, 1834

269Subtribe Callistina Laporte, 1834

270Subtribe Chlaeniina Brullé, 1834

271Tribe Cnemalobini Germain, 1911

272Tribe Cratocerini Lacordaire, 1854

273Tribe Ctenodactylini Laporte, 1834

274Tribe Cuneipectini Sloane, 1907

275Tribe Cyclosomini Laporte, 1834

276Subtribe Cyclosomina Laporte, 1834

277Subtribe Masoreina Chaudoir, 1871

278Tribe Dercylini Sloane, 1923

279Tribe Drimostomatini Chaudoir, 1872

280Tribe Dryptini Bonelli, 1810

281Tribe Enoicini Basilewsky, 1985

282Tribe Galeritini Kirby, 1825

283Subtribe Galeritina Kirby, 1825

284Subtribe Planetina Jedlička, 1941

285Tribe Geobaenini Péringuey, 1896

286Tribe Ginemini Ball and Shpeley, 2002

287Tribe Glyptini Horn, 1881

288Tribe Graphipterini Latreille, 1802

289Tribe Harpalini Bonelli, 1810

290Subtribe Anisodactylina Lacordaire, 1854 nomen protectum

291Subtribe Harpalina Bonelli, 1810

292Subtribe Pelmatellina Bates, 1882

293Subtribe Stenolophina Kirby, 1837

294Tribe Helluonini Hope, 1838

295Subtribe Helluonina Hope, 1838

296Subtribe Omphrina Jedlička, 1941

297Tribe Hexagoniini Horn, 1881 (1834)

298Tribe Idiomorphini Bates, 1891

299Tribe Lachnophorini LeConte, 1853

300Subtribe Lachnophorina LeConte, 1853

301Subtribe Selinina Jeannel, 1948

302Tribe Lebiini Bonelli, 1810

303Subtribe Actenonycina Bates, 1871

304Subtribe Agrina Kirby, 1837

305Subtribe Apenina Ball, 1983

306Subtribe Calleidina Chaudoir, 1873

307Subtribe Celaenephina Habu, 1982

308Subtribe Cymindidina Laporte, 1834

309Subtribe Demetriadina Bates, 1886

310Subtribe Dromiusina Bonelli, 1810

311Subtribe Gallerucidiina Chaudoir, 1872

312Subtribe Lebiina Bonelli, 1810

313Subtribe Metallicina Basilewsky, 1984

314Subtribe Nemotarsina Bates, 1883

315Subtribe Pericalina Hope, 1838

316Subtribe Pseudotrechina Basilewsky, 1984

317Subtribe Sugimotoina Habu, 1975

318Subtribe Trichina Basilewsky, 1984

319Tribe Licinini Bonelli, 1810

320Subtribe Dicaelina Laporte, 1834

321Subtribe Dicrochilina Ball, 1992

322Subtribe Lestignathina Ball, 1992

323Subtribe Licinina Bonelli, 1810

324Tribe Melanchitonini Jeannel, 1948

325Tribe Microcheilini Jeannel, 1948

326Tribe Morionini Brullé, 1835

327Tribe Odacanthini Laporte, 1834

328Tribe Omphreini Ganglbauer, 1891

329Tribe Oodini LaFerté-Sénectère, 1851

330Tribe Orthogoniini Schaum, 1857

331Tribe Panagaeini Bonelli, 1810

332Subtribe Brachygnathina Basilewsky, 1946

333Subtribe Panagaeina Bonelli, 1810

334Subtribe Tefflina Basilewsky, 1946

335Tribe Peleciini Chaudoir, 1880

336Subtribe Agonicina Sloane, 1920

337Subtribe Peleciina Chaudoir, 1880

338Tribe Pentagonicini Bates, 1873

339Tribe Perigonini Horn, 1881 nomen protectum

340Tribe Physocrotaphini Chaudoir, 1863

341Tribe Platynini Bonelli, 1810

342Tribe Pseudomorphini Hope, 1838

343Tribe Pterostichini Bonelli, 1810

344Subtribe Abacomorphina Tschitschérine, 1902

345Subtribe Euchroina Chaudoir, 1874

346Subtribe Metiina Straneo, 1951

347Subtribe Microcephalina Tschitschérine, 1898

348Subtribe Pterostichina Bonelli, 1810

349Tribe Sphodrini Laporte, 1834

350Subtribe Atranopsina Baehr, 1982

351Subtribe Calathina Laporte, 1834

352Subtribe Dolichina Brullé, 1834

353Subtribe Pristosiina Lindroth, 1956

354Subtribe Sphodrina Laporte, 1834

355Subtribe Synuchina Lindroth, 1956

356Tribe Xenaroswellianini Erwin, 2007

357Tribe Zabrini Bonelli, 1810

358Subtribe Amarina Zimmermann, 1832

359Subtribe Zabrina Bonelli, 1810

360Tribe Zuphiini Bonelli, 1810

361Subtribe Dicrodontina Machado, 1992

362Subtribe Leleupidiina Basilewsky, 1951

363Subtribe Metazuphiina Mateu, 1992

364Subtribe Mischocephalina Mateu, 1992

365Subtribe Patriziina Basilewsky, 1953

366Subtribe Zuphiina Bonelli, 1810

367Family Haliplidae Aubé, 1836

368†Family Triaplidae Ponomarenko, 1977

369†Family Colymbotethidae Ponomarenko, 1994

370†Family Parahygrobiidae Ponomarenko, 1977

371†Family Coptoclavidae Ponomarenko, 1961

372†Subfamily Necronectinae Ponomarenko, 1977

373†Subfamily Charonoscaphinae Ponomarenko, 1977

374†Subfamily Coptoclavinae Ponomarenko, 1961

375†Subfamily Coptoclaviscinae Soriano, Ponomarenko and Delclos, 2007

376†Subfamily Hispanoclavinae Soriano, Ponomarenko and Delclos, 2007

377†Family Liadytidae Ponomarenko, 1977

378Family Meruidae Spangler and Steiner, 2005

379Family Noteridae Thomson, 1860

380Subfamily Noterinae Thomson, 1860

381Tribe Neohydrocoptini Zalat, Saleh, Angus and Kaschef, 2000

382Tribe Noterini Thomson, 1860

383Tribe Pronoterini Nilsson, 2005

384Tribe Tonerini Miller, 2009

385Subfamily Notomicrinae Zimmermann, 1919

386Subfamily Phreatodytinae Uéno, 1957

387Family Amphizoidae LeConte, 1853

388Family Aspidytidae Ribera, Beutel, Balke and Vogler, 2002

389Family Hygrobiidae Régimbart, 1879 (1837)

390Family Dytiscidae Leach, 1815

391Subfamily Agabinae Thomson, 1867

392Subfamily Colymbetinae Erichson, 1837

393Tribe Anisomeriini Brinck, 1948

394Tribe Carabdytini Pederzani, 1995

395Tribe Colymbetini Erichson, 1837

396Subfamily Copelatinae Branden, 1885

397Subfamily Coptotominae Branden, 1885

398Subfamily Dytiscinae Leach, 1815

399Tribe Aciliini Thomson, 1867

400Tribe Aubehydrini Guignot, 1942

401Tribe Cybisterini Sharp, 1880

402Tribe Dytiscini Leach, 1815

403Tribe Eretini Crotch, 1873

404Tribe Hydaticini Sharp, 1880

405Tribe Hyderodini Miller, 2000

406Subfamily Hydrodytinae Miller, 2001

407Subfamily Hydroporinae Aubé, 1836

408Tribe Bidessini Sharp, 1880

409Tribe Carabhydrini Watts, 1978

410Tribe Hydroporini Aubé, 1836

411Tribe Hydrovatini Sharp, 1880

412Tribe Hygrotini Portevin, 1929

413Tribe Hyphydrini Gistel, 1848

414Tribe Laccornini Wolfe and Roughley, 1990

415Tribe Methlini Branden, 1885

416†Tribe Schistomerini Palmer, 1957

417Tribe Vatellini Sharp, 1880

418Subfamily Laccophilinae Gistel, 1848

419Tribe Agabetini Branden, 1885

420Tribe Laccophilini Gistel, 1848

421Subfamily Lancetinae Branden, 1885

422Subfamily Matinae Branden, 1885

423†Subfamily Palaeogyrininae Schlechtendal, 1894

424†Subfamily Liadytiscinae Prokin and Ren, 2010

425Suborder POLYPHAGA

426Series STAPHYLINIFORMIA

427Superfamily Hydrophiloidea Latreille, 1802

428Family Hydrophilidae Latreille, 1802

429Subfamily Helophorinae Leach, 1815

430Subfamily Epimetopinae Zaitzev, 1908

431Subfamily Georissinae Laporte, 1840

432Subfamily Hydrochinae Thomson, 1859

433Subfamily Spercheinae Erichson, 1837

434Subfamily Horelophinae Hansen, 1991

435Subfamily Horelophopsinae Hansen, 1997

436Subfamily Hydrophilinae Latreille, 1802

437Tribe Anacaenini Hansen, 1991

438Tribe Berosini Mulsant, 1844

439Tribe Chaetarthriini Bedel, 1881

440Tribe Hydrophilini Latreille, 1802

441Subtribe Acidocerina Zaitzev, 1908

442Subtribe Globuloseina García, 2001

443Subtribe Hydrobiusina Mulsant, 1844

444Subtribe Hydrophilina Latreille, 1802

445Tribe Laccobiini Houlbert, 1922

446Tribe Sperchopsini Hansen, 1991

447Subfamily Sphaeridiinae Latreille, 1802

448Tribe Andotypini Hansen, 1991

449Tribe Borborophorini Hansen, 1991

450Tribe Coelostomatini Heyden, 1891 (1890)

451Tribe Megasternini Mulsant, 1844

452Tribe Omicrini Smetana, 1975

453Tribe Protosternini Hansen, 1991

454Tribe Rygmodini Orchymont, 1916

455Tribe Sphaeridiini Latreille, 1802

456Tribe Tormissini Hansen, 1991

457Family Sphaeritidae Shuckard, 1839

458Family Synteliidae Lewis, 1882

459Family Histeridae Gyllenhal, 1808

460Subfamily Niponiinae Fowler, 1912

461Subfamily Abraeinae MacLeay, 1819

462Tribe Abraeini MacLeay, 1819

463Tribe Acritini Wenzel, 1944

464Tribe Acritomorphini Wenzel, 1944

465Tribe Plegaderini Portevin, 1929

466Tribe Teretriini Bickhardt, 1914

467Subfamily Trypeticinae Bickhardt, 1913

468Subfamily Trypanaeinae Marseul, 1857

469Subfamily Saprininae Blanchard, 1845

470Subfamily Dendrophilinae Reitter, 1909

471Tribe Anapleini Olexa, 1982

472Tribe Bacaniini Kryzhanovskij, 1976

473Tribe Dendrophilini Reitter, 1909

474Tribe Paromalini Reitter, 1909

475Subfamily Onthophilinae MacLeay, 1819

476Subfamily Tribalinae Bickhardt, 1914

477Subfamily Histerinae Gyllenhal, 1808

478Tribe Exosternini Bickhardt, 1914

479Tribe Histerini Gyllenhal, 1808

480Tribe Hololeptini Hope, 1840

481Tribe Omalodini Kryzhanovskij, 1972

482Tribe Platysomatini Bickhardt, 1914

483Subfamily Haeteriinae Marseul, 1857

484Tribe Haeteriini Marseul, 1857

485Tribe Nymphistrini Tishechkin, 2007

486Tribe Synoditulini Tishechkin, 2007

487Subfamily Chlamydopsinae Bickhardt, 1914

488Superfamily Staphylinoidea Latreille, 1802

489Family Hydraenidae Mulsant, 1844

490Subfamily Orchymontiinae Perkins, 1997

491Subfamily Prosthetopinae Perkins, 1994

492Tribe Coelometoponini Perkins, 2005

493Tribe Nucleotopini Perkins, 1994

494Tribe Parasthetopini Perkins, 1994

495Tribe Prosthetopini Perkins, 1994

496Tribe Protosthetopini Perkins, 1994

497Tribe Pterosthetopini Perkins, 1994

498Subfamily Hydraeninae Mulsant, 1844

499Tribe Hydraenidini Perkins, 1980

500Tribe Hydraenini Mulsant, 1844

501Tribe Limnebiini Mulsant, 1844

502Tribe Madagastrini Perkins, 1997

503Tribe Parhydraenini Perkins, 1997

504Subfamily Ochthebiinae Thomson, 1859

505Tribe Ochthebiini Thomson, 1859

506Subtribe Enicocerina Perkins, 1997

507Subtribe Meropathina Perkins, 1997

508Subtribe Neochthebiina Perkins, 1997

509Subtribe Ochthebiina Thomson, 1859

510Subtribe Protochthebiina Perkins, 1997

511Tribe Ochtheosini Perkins, 1997

512Family Ptiliidae Erichson, 1845

513Subfamily Ptiliinae Erichson, 1845

514Tribe Discheramocephalini Grebennikov, 2009

515Tribe Nanosellini Barber, 1924

516Tribe Ptenidiini Flach, 1889

517Tribe Ptiliini Erichson, 1845

518Tribe Ptinellini Reitter, 1906 (1891)

519Subfamily Cephaloplectinae Sharp, 1883

520Subfamily Acrotrichinae Reitter, 1909 (1856)

521Family Agyrtidae Thomson, 1859

522Subfamily Agyrtinae Thomson, 1859

523Subfamily Necrophilinae Newton, 1997

524Subfamily Pterolomatinae Thomson, 1862

525Family Leiodidae Fleming, 1821

526Subfamily Camiarinae Jeannel, 1911

527Tribe Agyrtodini Jeannel, 1936

528Tribe Camiarini Jeannel, 1911

529Tribe Neopelatopini Jeannel, 1962

530Subfamily Catopocerinae Hatch, 1927 (1880)

531Tribe Catopocerini Hatch, 1927 (1880)

532Tribe Glacicavicolini Westcott, 1968

533Subfamily Leiodinae Fleming, 1821

534Tribe Agathidiini Westwood, 1838

535Tribe Estadiini Portevin, 1914

536Tribe Leiodini Fleming, 1821

537Tribe Pseudoliodini Portevin, 1926

538Tribe Scotocryptini Reitter, 1884

539Tribe Sogdini Lopatin, 1961

540Subfamily Coloninae Horn, 1880 (1859)

541Subfamily Cholevinae Kirby, 1837

542Tribe Anemadini Hatch, 1928

543Subtribe Anemadina Hatch, 1928

544Subtribe Eocatopina Jeannel, 1936

545Subtribe Eunemadina Newton, 1998

546Subtribe Nemadina Jeannel, 1936

547Subtribe Paracatopina Jeannel, 1936

548Tribe Cholevini Kirby, 1837

549Subtribe Catopina Chaudoir, 1845

550Subtribe Cholevina Kirby, 1837

551Tribe Eucatopini Jeannel, 1921

552Tribe Leptodirini Lacordaire, 1854 (1849)

553Subtribe Anthroherponina Jeannel, 1910

554Subtribe Bathysciina Horn, 1880

555Subtribe Bathysciotina Guéorguiev, 1974

556Subtribe Leptodirina Lacordaire, 1854 (1849)

557Subtribe Pholeuina Reitter, 1886

558Subtribe Platycholeina Horn, 1880

559Subtribe Spelaeobatina Guéorguiev, 1974

560Tribe Oritocatopini Jeannel, 1936

561Tribe Ptomaphagini Jeannel, 1911

562Subtribe Baryodirina Perreau, 2000

563Subtribe Ptomaphagina Jeannel, 1911

564Subtribe Ptomaphaginina Szymczakowski, 1964

565Tribe Sciaphyini Perreau, 2000

566Subfamily Platypsyllinae Ritsema, 1869

567Family Silphidae Latreille, 1806

568Subfamily Silphinae Latreille, 1806

569Subfamily Nicrophorinae Kirby, 1837

570Family Staphylinidae Latreille, 1802

571Subfamily Glypholomatinae Jeannel, 1962

572Subfamily Microsilphinae Crowson, 1950

573Subfamily Omaliinae MacLeay, 1825

574Tribe Anthophagini Thomson, 1859

575Tribe Aphaenostemmini Peyerimhoff, 1914

576Tribe Corneolabiini Steel, 1950

577Tribe Coryphiini Jakobson, 1908

578Subtribe Boreaphilina Zerche, 1990

579Subtribe Coryphiina Jakobson, 1908

580Tribe Eusphalerini Hatch, 1957

581Tribe Hadrognathini Portevin, 1929

582Tribe Omaliini MacLeay, 1825

583Subfamily Empelinae Newton and Thayer, 1992

584Subfamily Proteininae Erichson, 1839

585Tribe Anepiini Steel, 1966

586Tribe Austrorhysini Newton and Thayer, 1995

587Tribe Nesoneini Steel, 1966

588Tribe Proteinini Erichson, 1839

589Tribe Silphotelini Newton and Thayer, 1995

590Subfamily Micropeplinae Leach, 1815

591Subfamily Neophoninae Fauvel, 1905

592Subfamily Dasycerinae Reitter, 1887

593Subfamily Protopselaphinae Newton and Thayer, 1995

594Subfamily Pselaphinae Latreille, 1802

595Supertribe Batrisitae Reitter, 1882

596Tribe Amauropini Jeannel, 1948

597Tribe Batrisini Reitter, 1882

598Subtribe Ambicocerina Leleup, 1970

599Subtribe Batrisina Reitter, 1882

600Subtribe Leupeliina Jeannel, 1954

601Subtribe Stilipalpina Jeannel, 1954

602Tribe Thaumastocephalini Poggi, Nonveiller, Colla, Pavićević and Rada, 2001

603Supertribe Clavigeritae Leach, 1815

604Tribe Clavigerini Leach, 1815

605Subtribe Apoderigerina Jeannel, 1954

606Subtribe Clavigerina Leach, 1815

607Subtribe Clavigerodina Schaufuss, 1882

608Subtribe Disarthricerina Jeannel, 1949

609Subtribe Hoplitoxenina Célis, 1969

610Subtribe Lunillina Célis, 1969

611Subtribe Mastigerina Jeannel, 1954

612Subtribe Miroclavigerina Jeannel, 1949

613Subtribe Neocerina Jeannel, 1954

614Subtribe Radamina Jeannel, 1954

615Subtribe Thysdariina Jeannel, 1954

616Tribe Colilodionini Besuchet, 1991

617Tribe Tiracerini Besuchet, 1986

618Supertribe Euplectitae Streubel, 1839

619Tribe Bythinoplectini Schaufuss, 1890

620Subtribe Bythinoplectina Schaufuss, 1890

621Subtribe Pyxidicerina Raffray, 1904

622Tribe Dimerini Raffray, 1908

623Tribe Euplectini Streubel, 1839

624Tribe Jubini Raffray, 1904

625Tribe Mayetiini Winkler, 1925

626Tribe Metopiasini Raffray, 1904

627Subtribe Metopiasina Raffray, 1904

628Subtribe Rhinoscepsina Bowman, 1934

629Tribe Trichonychini Reitter, 1882

630Subtribe Bibloporina Park, 1951

631Subtribe Panaphantina Jeannel, 1950

632Subtribe Trichonychina Reitter, 1882

633Subtribe Trimiina Bowman, 1934

634Tribe Trogastrini Jeannel, 1949

635Subtribe Phtegnomina Park, 1951

636Subtribe Rhexiina Park, 1951

637Subtribe Trogastrina Jeannel, 1949

638Supertribe Faronitae Reitter, 1882

639Supertribe Goniaceritae Reitter, 1882 (1872)

640Tribe Arnylliini Jeannel, 1952

641Tribe Barrosellini Leleup, 1973

642Tribe Brachyglutini Raffray, 1904

643Subtribe Baradina Park, 1951

644Subtribe Brachyglutina Raffray, 1904

645Subtribe Decarthrina Park, 1951

646Subtribe Eupseniina Park, 1951

647Tribe Bythinini Raffray, 1890

648Subtribe Bythinina Raffray, 1890

649Subtribe Machaeritina Jeannel, 1950

650Subtribe Xenobythina Jeannel, 1950

651Tribe Cyathigerini Schaufuss, 1872

652Tribe Goniacerini Reitter, 1882 (1872)

653Tribe Imirini Jeannel, 1949

654Tribe Iniocyphini Park, 1951

655Subtribe Iniocyphina Park, 1951

656Subtribe Natypleurina Newton and Thayer, 1992

657Tribe Machadoini Jeannel, 1951

658Tribe Proterini Jeannel, 1949

659Tribe Pygoxyini Reitter, 1909

660Tribe Speleobamini Park, 1951

661Tribe Tychini Raffray, 1904

662Tribe Valdini Park, 1953

663Supertribe Pselaphitae Latreille, 1802

664Tribe Arhytodini Raffray, 1890

665Tribe Attapseniini Bruch, 1933

666Tribe Ctenistini Blanchard, 1845

667Tribe Hybocephalini Raffray, 1890

668Tribe Odontalgini Jeannel, 1949

669Tribe Pachygastrodini Leleup, 1969

670Tribe Phalepsini Jeannel, 1949

671Tribe Pselaphini Latreille, 1802

672Tribe Schistodactylini Raffray, 1890

673Tribe Tmesiphorini Jeannel, 1949

674Tribe Tyrini Reitter, 1882

675Subtribe Centrophthalmina Jeannel, 1949

676Subtribe Janusculina Cerruti, 1970

677Subtribe Somatipionina Jeannel, 1949

678Subtribe Tyrina Reitter, 1882

679Subfamily Phloeocharinae Erichson, 1839

680Subfamily Olisthaerinae Thomson, 1858

681Subfamily Tachyporinae MacLeay, 1825

682Tribe Deropini Smetana, 1983

683Tribe Megarthropsini Cameron, 1919

684Tribe Mycetoporini Thomson, 1859

685Tribe Tachyporini MacLeay, 1825

686Tribe Vatesini Seevers, 1958

687Subfamily Trichophyinae Thomson, 1858

688Subfamily Habrocerinae Mulsant and Rey, 1876

689Subfamily Aleocharinae Fleming, 1821

690Tribe Actocharini Bernhauer and Schubert, 1911

691Tribe Aenictoteratini Kistner, 1993

692Tribe Akatastopsisini Pace, 2000

693Tribe Aleocharini Fleming, 1821

694Subtribe Aleocharina Fleming, 1821

695Subtribe Compactopediina Kistner, 1970

696Subtribe Hodoxenina Kistner, 1970

697Tribe Athetini Casey, 1910

698Subtribe Athetina Casey, 1910

699Subtribe Coptotermoeciina Kistner and Pasteels, 1970

700Subtribe Microceroxenina Kistner, 1970

701Subtribe Nasutiphilina Kistner, 1970

702Subtribe Schistogeniina Fenyes, 1918

703Subtribe Taxicerina Lohse, 1989

704Subtribe Termitotelina Kistner, 1970

705Subtribe Thamiaraeina Fenyes, 1921

706Tribe Autaliini Thomson, 1859

707Tribe Cordobanini Bernhauer, 1910

708Tribe Corotocini Fenyes, 1918

709Subtribe Abrotelina Seevers, 1957

710Subtribe Corotocina Fenyes, 1918

711Subtribe Eburniogastrina Jacobson, Kistner and Pasteels, 1986

712Subtribe Nasutitellina Jacobson, Kistner and Pasteels, 1986

713Subtribe Sphuridaethina Pace, 1988

714Subtribe Termitocharina Seevers, 1957

715Subtribe Termitocupidina Jacobson, Kistner and Pasteels, 1986

716Subtribe Termitogastrina Bernhauer and Scheerpeltz, 1926

717Subtribe Termitoiceina Jacobson, Kistner and Pasteels, 1986

718Subtribe Termitopithina Jacobson, Kistner and Pasteels, 1986

719Subtribe Termitoptochina Fenyes, 1921

720Subtribe Timeparthenina Fenyes, 1921

721Tribe Crematoxenini Mann, 1921

722Tribe Cryptonotopseini Pace, 2003

723Tribe Deinopsini Sharp, 1883

724Tribe Diestotini Mulsant and Rey, 1871

725Tribe Diglottini Jakobson, 1909

726Tribe Digrammini Fauvel, 1900

727Tribe Dorylogastrini Wasmann, 1916

728Tribe Dorylomimini Wasmann, 1916

729Tribe Drepanoxenini Kistner and Watson, 1972

730Tribe Ecitocharini Seevers, 1965

731Tribe Ecitogastrini Fenyes, 1918

732Tribe Eusteniamorphini Bernhauer and Scheerpeltz, 1926

733Tribe Falagriini Mulsant and Rey, 1873

734Tribe Feldini Kistner, 1972

735Tribe Gymnusini Heer, 1839

736Tribe Himalusini Klimaszewski, Pace and Center, 2010

737Tribe Homalotini Heer, 1839

738Subtribe Bolitocharina Thomson, 1859

739Subtribe Dinardopsina Bernhauer and Scheerpeltz, 1926

740Subtribe Gyrophaenina Kraatz, 1856

741Subtribe Homalotina Heer, 1839

742Subtribe Silusina Fenyes, 1918

743Tribe Hoplandriini Casey, 1910

744Subtribe Hoplandriina Casey, 1910

745Subtribe Platandriina Hanley, 2002

746Subtribe Pseudoplandriina Hanley, 2002

747Tribe Hygronomini Thomson, 1859

748Subtribe Hygronomina Thomson, 1859

749Subtribe Saphoglossina Bernhauer and Scheerpeltz, 1926

750Tribe Hypocyphtini Laporte, 1835

751Tribe Leucocraspedini Fenyes, 1921

752Tribe Liparocephalini Fenyes, 1918

753Tribe Lomechusini Fleming, 1821

754Subtribe Aenictobiina Kistner, 1997

755Subtribe Lomechusina Fleming, 1821

756Subtribe Myrmedoniina Thomson, 1867

757Subtribe Termitozyrina Seevers, 1957

758Tribe Masuriini Cameron, 1939

759Tribe Mesoporini Cameron, 1959

760Tribe Mimanommatini Wasmann, 1912

761Subtribe Dorylophilina Fenyes, 1921

762Subtribe Mimanommatina Wasmann, 1912

763Tribe Mimecitini Wasmann, 1917

764Subtribe Labidopullina Jacobson and Kistner, 1991

765Subtribe Leptanillophilina Fenyes, 1918

766Subtribe Mimecitina Wasmann, 1917

767Subtribe Mimonillina Bernhauer and Scheerpeltz, 1926

768Tribe Myllaenini Ganglbauer, 1895

769Tribe Oxypodini Thomson, 1859

770Subtribe Aphytopodina Bernhauer and Scheerpeltz, 1926

771Subtribe Blepharhymenina Klimaszewski and Peck, 1986

772Subtribe Dinardina Mulsant and Rey, 1873

773Subtribe Meoticina Seevers, 1978

774Subtribe Oxypodina Thomson, 1859

775Subtribe Tachyusina Thomson, 1859

776Tribe Oxypodinini Fenyes, 1921

777Tribe Paglini Newton and Thayer, 1992

778Tribe Paradoxenusini Bruch, 1937

779Tribe Pediculotini Ádám, 1987

780Tribe Philotermitini Seevers, 1957

781Tribe Phyllodinardini Wasmann, 1916

782Tribe Phytosini Thomson, 1867

783Tribe Placusini Mulsant and Rey, 1871

784Tribe Pronomaeini Mulsant and Rey, 1873

785Tribe Pseudoperinthini Cameron, 1939

786Tribe Pygostenini Fauvel, 1899

787Tribe Sahlbergiini Kistner, 1993

788Tribe Sceptobiini Seevers, 1978

789Tribe Skatitoxenini Kistner and Pasteels, 1969

790Tribe Termitodiscini Wasmann, 1904

791Subtribe Athexeniina Pace, 2000

792Subtribe Termitodiscina Wasmann, 1904

793Tribe Termitohospitini Seevers, 1941

794Subtribe Hetairotermitina Seevers, 1957

795Subtribe Termitohospitina Seevers, 1941

796Tribe Termitonannini Fenyes, 1918

797Subtribe Perinthina Bernhauer and Scheerpeltz, 1926

798Subtribe Termitonannina Fenyes, 1918

799Tribe Termitopaediini Seevers, 1957

800Tribe Termitusini Fenyes, 1918

801Subtribe Termitospectrina Seevers, 1957

802Subtribe Termitusina Fenyes, 1918

803Tribe Trichopseniini LeConte and Horn, 1883

804Tribe Trilobitideini Fauvel, 1899

805Subfamily Trigonurinae Reiche, 1866

806Subfamily Apateticinae Fauvel, 1895

807Subfamily Scaphidiinae Latreille, 1806

808Tribe Cypariini Achard, 1924

809Tribe Scaphidiini Latreille, 1806

810Tribe Scaphiini Achard, 1924

811Tribe Scaphisomatini Casey, 1893

812Subfamily Piestinae Erichson, 1839

813Subfamily Osoriinae Erichson, 1839

814Tribe Eleusinini Sharp, 1887

815Tribe Leptochirini Sharp, 1887

816Tribe Osoriini Erichson, 1839

817Subtribe Osoriina Erichson, 1839

818Subtribe Parosoriina Bernhauer and Schubert, 1911

819Tribe Thoracophorini Reitter, 1909

820Subtribe Clavilispinina Newton and Thayer, 1992

821Subtribe Glyptomina Newton and Thayer, 1992

822Subtribe Lispinina Bernhauer and Schubert, 1910

823Subtribe Thoracophorina Reitter, 1909

824Subfamily Oxytelinae Fleming, 1821

825Tribe Blediini Ádám, 2001

826Tribe Coprophilini Heer, 1839

827Tribe EuphaniiniReitter, 1909

828Tribe Oxytelini Fleming, 1821

829Tribe Planeustomini Jacquelin du Val, 1857

830Subfamily Oxyporinae Fleming, 1821

831Subfamily Megalopsidiinae Leng, 1920

832Subfamily Scydmaeninae Leach, 1815

833†Supertribe Hapsomelitae Poinar and Brown, 2004

834Supertribe Mastigitae Fleming, 1821

835Tribe Clidicini Casey, 1897

836Tribe Leptomastacini Casey, 1897

837Tribe Mastigini Fleming, 1821

838Supertribe Scydmaenitae Leach, 1815

839Tribe Cephenniini Reitter, 1882

840Tribe Chevrolatiini Reitter, 1882

841Tribe Cyrtoscydmini Schaufuss, 1889

842Tribe Eutheiini Casey, 1897

843Tribe Leptoscydmini Casey, 1897

844Tribe Plaumanniolini Costa Lima, 1962

845Tribe Scydmaenini Leach, 1815

846Subfamily Steninae MacLeay, 1825

847Subfamily Euaesthetinae Thomson, 1859

848Tribe Alzadaesthetini Scheerpeltz, 1974

849Tribe Austroesthetini Cameron, 1944

850Tribe Euaesthetini Thomson, 1859

851Tribe Fenderiini Scheerpeltz, 1974

852Tribe Nordenskioldiini Bernhauer and Schubert, 1911

853Tribe Stenaesthetini Bernhauer and Schubert, 1911

854Subfamily Solieriinae Newton and Thayer, 1992

855Subfamily Leptotyphlinae Fauvel, 1874

856Tribe Cephalotyphlini Coiffait, 1963

857Tribe Entomoculiini Coiffait, 1957

858Tribe Leptotyphlini Fauvel, 1874

859Tribe Metrotyphlini Coiffait, 1963

860Tribe Neotyphlini Coiffait, 1963

861Subfamily Pseudopsinae Ganglbauer, 1895

862Subfamily Paederinae Fleming, 1821

863Tribe Paederini Fleming, 1821

864Subtribe Astenina Hatch, 1957

865Subtribe Cryptobiina Casey, 1905

866Subtribe Cylindroxystina Bierig, 1943

867Subtribe Dolicaonina Casey, 1905

868Subtribe Echiasterina Casey, 1905

869Subtribe Lathrobiina Laporte, 1835

870Subtribe Lithocharina Casey, 1905

871Subtribe Medonina Casey, 1905

872Subtribe Paederina Fleming, 1821

873Subtribe Scopaeina Mulsant and Rey, 1878

874Subtribe Stilicina Casey, 1905

875Subtribe Stilicopsina Casey, 1905

876Tribe Pinophilini Nordmann, 1837

877Subtribe Pinophilina Nordmann, 1837

878Subtribe Procirrina Bernhauer and Schubert, 1912

879Subfamily Staphylininae Latreille, 1802

880Tribe Arrowinini Solodovnikov and Newton, 2005

881Tribe Diochini Casey, 1906

882Tribe Maorothiini Assing, 2000

883Tribe Othiini Thomson, 1859

884Tribe Platyprosopini Lynch Arribálzaga, 1884

885Tribe Staphylinini Latreille, 1802

886Subtribe Amblyopinina Seevers, 1944

887Subtribe Anisolinina Hayashi, 1993

888Subtribe Eucibdelina Sharp, 1889

889Subtribe Hyptiomina Casey, 1906

890Subtribe Philonthina Kirby, 1837

891Subtribe Quediina Kraatz, 1857

892Subtribe Staphylinina Latreille, 1802

893Subtribe Tanygnathinina Reitter, 1909

894Subtribe Xanthopygina Sharp, 1884

895Tribe Xantholinini Erichson, 1839

896†Subfamily Protactinae Heer, 1847

897Series SCARABAEIFORMIA

898Superfamily Scarabaeoidea Latreille, 1802

899Family Pleocomidae LeConte, 1861

900Subfamily Pleocominae LeConte, 1861

901†Subfamily Cretocominae Nikolajev, 2002

902†Subfamily Archescarabaeinae Nikolajev, 2010

903Family Geotrupidae Latreille, 1802

904Subfamily Taurocerastinae Germain, 1897

905Subfamily Bolboceratinae Mulsant, 1842

906Tribe Athyreini Lynch Arribálzaga, 1878

907Tribe Bolbelasmini Nikolajev, 1996

908Tribe Bolboceratini Mulsant, 1842

909Tribe Bolbochromini Nikolajev, 1970

910Tribe Eubolbitini Nikolajev, 1970

911Tribe Eucanthini Nikolajev, 2003

912Tribe Gilletinini Nikolajev, 1990

913Tribe Odonteini Shokhin, 2007

914Tribe Stenaspidiini Nikolajev, 2003

915Subfamily Geotrupinae Latreille, 1802

916Tribe Ceratotrupini Zunino, 1984

917Tribe Enoplotrupini Paulian, 1945

918†Tribe Cretogeotrupini Nikolajev, 1996

919Tribe Geotrupini Latreille, 1802

920Tribe Lethrini Oken, 1843

921Family Belohinidae Paulian, 1959

922Family Passalidae Leach, 1815

923Subfamily Aulacocyclinae Kaup, 1868

924Tribe Aulacocyclini Kaup, 1868

925Tribe Ceracupedini Boucher, 2006

926Subfamily Passalinae Leach, 1815

927Tribe Leptaulacini Kaup, 1871

928Tribe Macrolinini Kaup, 1871

929Tribe Passalini Leach, 1815

930Tribe Proculini Kaup, 1868

931Tribe Solenocyclini Kaup, 1871

932Family Trogidae MacLeay, 1819

933†Subfamily Avitortorinae Nikolajev, 2007

934Subfamily Troginae MacLeay, 1819

935Subfamily Omorginae Nikolajev, 2005

936Family Glaresidae Kolbe, 1905

937Family Diphyllostomatidae Holloway, 1972

938Family Lucanidae Latreille, 1804

939†Subfamily Protolucaninae Nikolajev, 2007

940Subfamily Aesalinae MacLeay, 1819

941Tribe Aesalini MacLeay, 1819

942Tribe Ceratognathini Sharp, 1899

943Tribe Nicagini LeConte, 1861

944†Subfamily Ceruchitinae Nikolajev, 2006

945Subfamily Syndesinae MacLeay, 1819

946Subfamily Lampriminae MacLeay, 1819

947Tribe Lamprimini MacLeay, 1819

948Tribe Streptocerini Kikuta, 1986

949Subfamily Lucaninae Latreille, 1804

950Tribe Chiasognathini Burmeister, 1847

951Tribe Lucanini Latreille, 1804

952Tribe Platycerini Mulsant, 1842

953Tribe Platyceroidini Paulsen and Hawks, 2008

954†Subfamily Paralucaninae Nikolajev, 2000

955Family Ochodaeidae Mulsant and Rey, 1871

956†Subfamily Cretochodaeinae Nikolajev, 1995

957Subfamily Ochodaeinae Mulsant and Rey, 1871

958Tribe Enodognathini Scholtz, 1988

959Tribe Ochodaeini Mulsant and Rey, 1871

960Subfamily Chaetocanthinae Scholtz, 1988

961Tribe Chaetocanthini Scholtz, 1988

962Tribe Pseudochodaeini Scholtz, 1988

963Tribe Synochodaeini Scholtz, 1988

964Family Hybosoridae Erichson, 1847

965†Subfamily Mimaphodiinae Nikolajev, 2007

966Subfamily Anaidinae Nikolajev, 1996

967Subfamily Ceratocanthinae Martínez, 1968

968Tribe Ceratocanthini Martínez, 1968

969Tribe Ivieolini Howden and Gill, 2000

970Tribe Scarabatermitini Nikolajev, 1999

971Subfamily Hybosorinae Erichson, 1847

972Subfamily Liparochrinae Ocampo, 2006

973Subfamily Pachyplectrinae Ocampo, 2006

974Family Glaphyridae MacLeay, 1819

975Subfamily Glaphyrinae MacLeay, 1819

976Subfamily Amphicominae Blanchard, 1845

977†Subfamily Cretoglaphyrinae Nikolajev, 2005

978Family Scarabaeidae Latreille, 1802

979†Subfamily Lithoscarabaeinae Nikolajev, 1992

980Subfamily Chironinae Blanchard, 1845

981Subfamily Aegialiinae Laporte, 1840

982Subfamily Eremazinae Iablokoff-Khnzorian, 1977

983Subfamily Aphodiinae Leach, 1815

984Tribe Aphodiini Leach, 1815

985Subtribe Aphodiina Leach, 1815

986Subtribe Didactyliina Pittino, 1985

987Subtribe Proctophanina Stebnicka and Howden, 1995

988Tribe Corythoderini Schmidt, 1910

989Tribe Eupariini Schmidt, 1910 nomen protectum

990Tribe Odontolochini Stebnicka and Howden, 1996

991Tribe Odochilini Rakovič, 1987

992Tribe Psammodiini Mulsant, 1842

993Subtribe Phycocina Landin, 1960

994Subtribe Psammodiina Mulsant, 1842

995Subtribe Rhyssemina Pittino and Mariani, 1986

996Tribe Rhyparini Schmidt, 1910

997Tribe Stereomerini Howden and Storey, 1992

998Tribe Termitoderini Tangelder and Krikken, 1982

999Subfamily Aulonocneminae Janssens, 1946

1000Subfamily Termitotroginae Wasmann, 1918

1001Subfamily Scarabaeinae Latreille, 1802

1002Tribe Ateuchini Perty, 1830

1003Subtribe Ateuchina Perty, 1830

1004Subtribe Scatimina Vaz-de-Mello, 2008

1005Tribe Coprini Leach, 1815

1006Tribe Deltochilini Lacordaire, 1856

1007Tribe Eucraniini Burmeister, 1873

1008Tribe Gymnopleurini Lacordaire, 1856

1009Tribe Oniticellini Kolbe, 1905

1010Subtribe Drepanocerina van Lansberge, 1875

1011Subtribe Eurysternina Vulcano, Martínez and Pereira, 1961

1012Subtribe Helictopleurina Janssens, 1946

1013Subtribe Oniticellina Kolbe, 1905

1014Tribe Onitini Laporte, 1840

1015Tribe Onthophagini Burmeister, 1846

1016Tribe Phanaeini Hope, 1838

1017Tribe Scarabaeini Latreille, 1802

1018Tribe Sisyphini Mulsant, 1842

1019†Subfamily Prototroginae Nikolajev, 2000

1020†Subfamily Cretoscarabaeinae Nikolajev, 1995

1021Subfamily Dynamopodinae Arrow, 1911

1022Tribe Dynamopodini Arrow, 1911

1023Tribe Thinorycterini Semenov and Reichardt, 1925

1024Subfamily Phaenomeridinae Erichson, 1847

1025Subfamily Orphninae Erichson, 1847

1026Tribe Aegidiini Paulian, 1984

1027Tribe Orphnini Erichson, 1847

1028Subfamily Allidiostomatinae Arrow, 1940

1029Subfamily Aclopinae Blanchard, 1850

1030Tribe Aclopini Blanchard, 1850

1031†Tribe Holcorobeini Nikolajev, 1992

1032Tribe Phaenognathini Iablokoff-Khnzorian, 1977

1033Subfamily Melolonthinae Leach, 1819

1034Tribe Ablaberini Blanchard, 1850

1035Tribe Automoliini Britton, 1978

1036Tribe Chasmatopterini Lacordaire, 1856

1037Tribe Colymbomorphini Blanchard, 1850

1038Tribe Comophorinini Britton, 1957

1039†Tribe Cretomelolonthini Nikolajev, 1998

1040Tribe Dichelonychini Burmeister, 1855

1041Tribe Diphucephalini Laporte, 1840

1042Tribe Diphycerini Medvedev, 1952

1043Tribe Diplotaxini Kirby, 1837

1044Tribe Euchirini Hope, 1840

1045Tribe Heteronychini Lacordaire, 1856

1046Tribe Hopliini Latreille, 1829

1047Subtribe Hopliina Latreille, 1829

1048Subtribe Pachycnemina Laporte, 1840

1049Tribe Lichniini Burmeister, 1844

1050Tribe Liparetrini Burmeister, 1855

1051Tribe Macrodactylini Kirby, 1837

1052Tribe Maechidiini Burmeister, 1855

1053Tribe Melolonthini Leach, 1819

1054Subtribe Enariina Dewailly, 1950

1055Subtribe Heptophyllina Medvedev, 1951

1056Subtribe Leucopholina Burmeister, 1855

1057Subtribe Melolonthina Leach, 1819

1058Subtribe Pegylina Lacroix, 1989

1059Subtribe Rhizotrogina Burmeister, 1855

1060Subtribe Schizonychina Burmeister, 1855

1061Tribe Oncerini LeConte, 1861

1062Tribe Pachypodini Erichson, 1840

1063Tribe Pachytrichini Burmeister, 1855

1064Tribe Phyllotocidiini Britton, 1957

1065Tribe Podolasiini Howden, 1997

1066Tribe Scitalini Britton, 1957

1067Tribe Sericini Kirby, 1837

1068Subtribe Phyllotocina Burmeister, 1855

1069Subtribe Sericina Kirby, 1837

1070Subtribe Trochalina Brenske, 1898

1071Tribe Sericoidini Erichson, 1847

1072Tribe Systellopini Sharp, 1877

1073Tribe Tanyproctini Erichson, 1847

1074Subtribe Macrophyllina Burmeister, 1855

1075Subtribe Tanyproctina Erichson, 1847

1076Subfamily Rutelinae MacLeay, 1819

1077Tribe Adoretini Burmeister, 1844

1078Subtribe Adoretina Burmeister, 1844

1079Subtribe Adorrhinyptiina Arrow, 1917

1080Subtribe Pachyrhinadoretina Ohaus, 1912

1081Subtribe Prodoretina Ohaus, 1912

1082Subtribe Trigonostomusina Ohaus, 1912

1083Tribe Alvarengiini Frey, 1975

1084Tribe Anatistini Lacordaire, 1856

1085Tribe Anomalini Streubel, 1839 nomen protectum

1086Subtribe Anisopliina Burmeister, 1844

1087Subtribe Anomalina Streubel, 1839 nomen protectum

1088Subtribe Isopliina Péringuey, 1902

1089Subtribe Leptohopliina Potts, 1974

1090Subtribe Popilliina Ohaus, 1918

1091Tribe Anoplognathini MacLeay, 1819

1092Subtribe Anoplognathina MacLeay, 1819

1093Subtribe Brachysternina Burmeister, 1844

1094Subtribe Phalangogoniina Ohaus, 1918

1095Subtribe Platycoeliina Burmeister, 1844

1096Subtribe Schizognathina Ohaus, 1918

1097Tribe Geniatini Burmeister, 1844

1098Tribe Rutelini MacLeay, 1819

1099Subtribe Areodina Burmeister, 1844

1100Subtribe Desmonychina Arrow, 1917

1101Subtribe Didrepanephorina Ohaus, 1918

1102Subtribe Heterosternina Bates, 1888 nomen protectum

1103Subtribe Lasiocalina Ohaus, 1918

1104Subtribe Oryctomorphina Burmeister, 1847

1105Subtribe Parastasiina Burmeister, 1844

1106Subtribe Rutelina MacLeay, 1819

1107Subfamily Dynastinae MacLeay, 1819

1108Tribe Agaocephalini Burmeister, 1847

1109Tribe Cyclocephalini Laporte, 1840

1110Tribe Dynastini MacLeay, 1819

1111Tribe Hexodontini Lacordaire, 1856

1112Tribe Oryctini Mulsant, 1842

1113Tribe Oryctoderini Endrödi, 1966

1114Tribe Pentodontini Mulsant, 1842

1115Subtribe Cheiroplatina Carne, 1957

1116Subtribe Dipelicina Carne, 1957

1117Subtribe Pentodontina Mulsant, 1842

1118Subtribe Pseudoryctina Carne, 1957

1119Tribe Phileurini Burmeister, 1847

1120Subtribe Cryptodina Burmeister and Schaum, 1840

1121Subtribe Phileurina Burmeister, 1847

1122Subfamily Cetoniinae Leach, 1815

1123Tribe Cetoniini Leach, 1815

1124Subtribe Cetoniina Leach, 1815

1125Subtribe Euphoriina Horn, 1880

1126Subtribe Leucocelina Kraatz, 1882

1127Tribe Cremastocheilini Burmeister and Schaum, 1841

1128Subtribe Aspilina Krikken, 1984

1129Subtribe Coenochilina Burmeister, 1842

1130Subtribe Cremastocheilina Burmeister and Schaum, 1841

1131Subtribe Cymophorina Krikken, 1984

1132Subtribe Genuchina Krikken, 1984

1133Subtribe Goliathopsidina Krikken, 1984

1134Subtribe Heterogeniina Krikken, 1984

1135Subtribe Lissogeniina Krikken, 1984

1136Subtribe Macromina Burmeister and Schaum, 1840

1137Subtribe Nyassinina Krikken, 1984

1138Subtribe Oplostomina Krikken, 1984

1139Subtribe Pilinurgina Krikken, 1984

1140Subtribe Spilophorina Krikken, 1984

1141Subtribe Telochilina Krikken, 1984

1142Subtribe Trichoplina Krikken, 1984

1143Subtribe Trogodina Krikken, 1984

1144Tribe Diplognathini Burmeister, 1842

1145Tribe Goliathini Latreille, 1829

1146Subtribe Coryphocerina Burmeister, 1842

1147Subtribe Dicronocephalina Krikken, 1984

1148Subtribe Goliathina Latreille, 1829

1149Subtribe Ichnestomatina Burmeister, 1842

1150Tribe Gymnetini Kirby, 1827

1151Subtribe Blaesiina Schoch, 1895

1152Subtribe Gymnetina Kirby, 1827

1153Tribe Phaedimini Schoch, 1894

1154Tribe Schizorhinini Burmeister, 1842

1155Subtribe Lomapterina Burmeister, 1842

1156Subtribe Schizorhinina Burmeister, 1842

1157Tribe Stenotarsiini Kraatz, 1880

1158Subtribe Anochiliina Krikken, 1984

1159Subtribe Coptomiina Schenkling, 1921

1160Subtribe Chromoptiliina Krikken, 1984

1161Subtribe Doryscelina Schenkling, 1921

1162Subtribe Euchroeina Paulian and Descarpentries, 1982

1163Subtribe Heterophanina Schoch, 1894

1164Subtribe Heterosomatina Krikken, 1984

1165Subtribe Pantoliina Krikken, 1984

1166Subtribe Parachiliina Krikken, 1984

1167Subtribe Stenotarsiina Kraatz, 1880

1168Tribe Taenioderini Mikšić, 1976

1169Subtribe Chalcotheina Mikšić, 1976

1170Subtribe Taenioderina Mikšić, 1976

1171Tribe Trichiini Fleming, 1821

1172Subtribe Cryptodontina Lacordaire, 1856

1173Subtribe Incina Burmeister, 1842

1174Subtribe Osmodermatina Schenkling, 1922

1175Subtribe Platygeniina Krikken, 1984

1176Subtribe Trichiina Fleming, 1821

1177Tribe Valgini Mulsant, 1842

1178Subtribe Microvalgina Kolbe, 1904

1179Subtribe Valgina Mulsant, 1842

1180Tribe Xiphoscelidini Burmeister, 1842

1181†Family Coprinisphaeridae Genise, 2004

1182†Family Pallichnidae Genise, 2004

1183Series ELATERIFORMIA

1184Superfamily Scirtoidea Fleming, 1821

1185Family Decliniidae Nikitsky, Lawrence, Kirejtshuk and Gratshev, 1994

1186Family Eucinetidae Lacordaire, 1857

1187Family Clambidae Fischer von Waldheim, 1821

1188Subfamily Calyptomerinae Crowson, 1955

1189Subfamily Acalyptomerinae Crowson, 1979

1190Subfamily Clambinae Fischer von Waldheim, 1821

1191Family Scirtidae Fleming, 1821

1192Subfamily Scirtinae Fleming, 1821

1193Subfamily Nipponocyphoninae Lawrence and Yoshitomi, 2007

1194Subfamily Stenocyphoninae Lawrence and Yoshitomi, 2007

1195†Family Elodophthalmidae Kirejtshuk and Azar, 2008

1196†Family Mesocinetida Kirejtshuk and Ponomarenko, 2010

1197Superfamily Dascilloidea Guérin-Méneville, 1843 (1834)

1198Family Dascillidae Guérin-Méneville, 1843 (1834)

1199Subfamily Dascillinae Guérin-Méneville, 1843 (1834)

1200Tribe Cinnabariini Pic, 1914

1201Tribe Dascillini Guérin-Méneville, 1843 (1834)

1202Subfamily Karumiinae Escalera, 1913

1203Tribe Emmitini Escalera, 1914

1204Tribe Escalerinini Paulus, 1972

1205Tribe Genecerini Pic, 1914

1206Tribe Karumiini Escalera, 1913

1207Family Rhipiceridae Latreille, 1834

1208Superfamily Buprestoidea Leach, 1815

1209Family Schizopodidae LeConte, 1859

1210Subfamily Schizopodinae LeConte, 1859

1211Tribe Dystaxiini Théry, 1929

1212†Tribe Electrapatini Iablokoff-Khnzorian, 1962

1213Tribe Schizopodini LeConte, 1859

1214Family Buprestidae Leach, 1815

1215Subfamily Julodinae Lacordaire, 1857

1216Subfamily Polycestinae Lacordaire, 1857

1217Tribe Acmaeoderini Kerremans, 1893

1218Subtribe Acmaeoderina Kerremans, 1893

1219Subtribe Acmaeoderoidina Cobos, 1955

1220Subtribe Nothomorphina Cobos, 1955

1221Tribe Astraeini Cobos, 1980

1222Tribe Bulini Bellamy, 1995

1223Tribe Haplostethini LeConte, 1861

1224Tribe Paratracheini Cobos, 1980

1225Tribe Perucolini Cobos, 1980

1226Tribe Polycestini Lacordaire, 1857

1227Subtribe Polycestina Lacordaire, 1857

1228Subtribe Xenopseina Volkovitsh, 2008

1229Tribe Polyctesini Cobos, 1955

1230Tribe Prospherini Cobos, 1980

1231Tribe Ptosimini Kerremans, 1903

1232Tribe Thrincopygini LeConte, 1861

1233Tribe Tyndaridini Cobos, 1955

1234Subtribe Mimicoclytrinina Bellamy, 2003

1235Subtribe Pseudacherusiina Cobos, 1980

1236Subtribe Tylaucheniina Cobos, 1959

1237Subtribe Tyndaridina Cobos, 1955

1238Tribe Xyroscelidini Cobos, 1955

1239Subfamily Galbellinae Reitter, 1911

1240Subfamily Chrysochroinae Laporte, 1835

1241Tribe Chrysochroini Laporte, 1835

1242Subtribe Chalcophorina Lacordaire, 1857 (1848)

1243Subtribe Chrysochroina Laporte, 1835

1244Subtribe Eucallopistina Bellamy, 2003

1245Tribe Dicercini Gistel, 1848

1246Subtribe Dicercina Gistel, 1848

1247Subtribe Haplotrinchina Holyński, 1993

1248Subtribe Hippomelanina Holyński, 1993

1249Subtribe Pseudoperotina Tôyama, 1987

1250Tribe Evidini Tôyama, 1987

1251Tribe Paraleptodemini Cobos, 1975

1252Subtribe Euchromatina Holyński, 1993

1253Subtribe Euplectaleciina Holyński, 1993

1254Subtribe Hypoprasina Holyński, 1993

1255Subtribe Paraleptodemina Cobos, 1975

1256Subtribe Pristipterina Holyński, 1993

1257Tribe Paratassini Bílý and Volkovitsh, 1996

1258Tribe Poecilonotini Jakobson, 1913

1259Subtribe Poecilonotina Jakobson, 1913

1260Subtribe Nesotrinchina Bílý, Kubáň and Volkovitsh, 2009

1261Tribe Sphenopterini Lacordaire, 1857

1262Tribe Vadonaxiini Descarpentries, 1970

1263Subfamily Buprestinae Leach, 1815

1264Tribe Actenodini Gistel, 1848

1265Tribe Anthaxiini Gory and Laporte, 1839

1266Tribe Bubastini Obenberger, 1920

1267Tribe Buprestini Leach, 1815

1268Subtribe Agaeocerina Nelson, 1982

1269Subtribe Buprestina Leach, 1815

1270Subtribe Lamprocheilina Holyński, 1993

1271Subtribe Trachykelina Holyński, 1988

1272Tribe Chrysobothrini Gory and Laporte, 1836

1273Tribe Coomaniellini Bílý, 1974

1274Tribe Curidini Holyński, 1988

1275Subtribe Anilarina Bílý, 2000

1276Subtribe Curidina Holyński, 1988

1277Subtribe Neocuridina Holyński, 1988

1278Tribe Epistomentini Levey, 1978

1279Tribe Exagistini Tôyama, 1987

1280Tribe Julodimorphini Kerremans, 1903

1281Tribe Kisanthobiini Richter, 1949

1282Tribe Maoraxiini Holyński, 1984

1283Tribe Melanophilini Bedel, 1921

1284Tribe Melobaseini Bílý, 2000

1285Tribe Mendizabaliini Cobos, 1968

1286Tribe Nascionini Holyński, 1988

1287Tribe Phrixiini Cobos, 1975

1288Tribe Pterobothrini Volkovitsh, 2001

1289Tribe Stigmoderini Lacordaire, 1857

1290Tribe Thomassetiini Bellamy, 1987

1291Subtribe Philanthaxiina Holyński, 1988

1292Subtribe Thomassetiina Bellamy, 1987

1293Tribe Trigonogeniini Cobos, 1956

1294Tribe Xenorhipidini Cobos, 1986

1295Subtribe Trichinorhipidina Bellamy, 2006

1296Subtribe Xenorhipidina Cobos, 1986

1297Subfamily Agrilinae Laporte, 1835

1298Tribe Agrilini Laporte, 1835

1299Subtribe Agrilina Laporte, 1835

1300Subtribe Amorphosternina Cobos, 1974

1301Subtribe Amyiina Holyński, 1993

1302Subtribe Rhaeboscelidina Cobos, 1976

1303Tribe Aphanisticini Jacquelin du Val, 1859

1304Subtribe Anthaxomorphina Holyński, 1993

1305Subtribe Aphanisticina Jacquelin du Val, 1859

1306Subtribe Cylindromorphina Portevin, 1931

1307Subtribe Cylindromorphoidina Cobos, 1979

1308Subtribe Germaricina Cobos, 1979

1309Tribe Coraebini Bedel, 1921

1310Subtribe Amorphosomatina Majer, 2000

1311Subtribe Cisseina Majer, 2000

1312Subtribe Clematina Majer, 2000

1313Subtribe Coraebina Bedel, 1921

1314Subtribe Dismorphina Cobos, 1990

1315Subtribe Ethoniina Majer, 2000

1316Subtribe Geraliina Cobos, 1988

1317Subtribe Meliboeina Majer, 2000

1318Subtribe Synechocerina Majer, 2000

1319Subtribe Toxoscelina Majer, 2000

1320Tribe Tracheini Laporte, 1835

1321Subtribe Brachina LeConte, 1861

1322Subtribe Leiopleurina Holyński, 1993

1323Subtribe Pachyschelina Böving and Craighead, 1931

1324Subtribe Tracheina Laporte, 1835

1325†Subfamily Parathyreinae Alexeev, 1994

1326Superfamily Byrrhoidea Latreille, 1804

1327Family Byrrhidae Latreille, 1804

1328Subfamily Byrrhinae Latreille, 1804

1329Tribe Byrrhini Latreille, 1804

1330Tribe Exomellini Casey, 1914

1331Tribe Morychini El Moursy, 1961

1332Tribe Pedilophorini Casey, 1912

1333Tribe Simplocariini Mulsant and Rey, 1869

1334Subfamily Syncalyptinae Mulsant and Rey, 1869

1335Tribe Microchaetini Paulus, 1973

1336Tribe Syncalyptini Mulsant and Rey, 1869

1337Subfamily Amphicyrtinae LeConte, 1861

1338Family Elmidae Curtis, 1830

1339Subfamily Larainae LeConte, 1861

1340Tribe Laraini LeConte, 1861

1341Tribe Potamophilini Mulsant and Rey, 1872

1342Subfamily Elminae Curtis, 1830

1343Tribe Ancyronychini Ganglbauer, 1904

1344Tribe Elmini Curtis, 1830

1345Subtribe Elmina Curtis, 1830

1346Subtribe Stenelmina Mulsant and Rey, 1872

1347Tribe Macronychini Gistel, 1848

1348Family Dryopidae Billberg, 1820 (1817)

1349Family Lutrochidae Kasap and Crowson, 1975

1350Family Limnichidae Erichson, 1846

1351Subfamily Hyphalinae Britton, 1971

1352Subfamily Limnichinae Erichson, 1846

1353Tribe Limnichini Erichson, 1846

1354Tribe Wooldridgeini Spangler, 1999

1355Subfamily Cephalobyrrhinae Champion, 1925

1356Subfamily Thaumastodinae Champion, 1924

1357Family Heteroceridae MacLeay, 1825

1358Subfamily Elythomerinae Pacheco, 1964

1359Subfamily Heterocerinae MacLeay, 1825

1360Tribe Augylini Pacheco, 1964

1361Tribe Heterocerini MacLeay, 1825

1362Tribe Micilini Pacheco, 1964

1363Tribe Tropicini Pacheco, 1964

1364Family Psephenidae Lacordaire, 1854

1365Subfamily Afroeubriinae Lee, Satô, Shepard and Jäch, 2007

1366Subfamily Eubriinae Lacordaire, 1857

1367Subfamily Eubrianacinae Jakobson, 1913

1368Subfamily Psephenoidinae Bollow, 1938

1369Subfamily Psepheninae Lacordaire, 1854

1370Family Cneoglossidae Champion, 1897

1371Family Ptilodactylidae Laporte, 1836

1372Subfamily Anchytarsinae Champion, 1897

1373Subfamily Cladotominae Pic, 1914

1374Subfamily Aploglossinae Champion, 1897

1375Subfamily Araeopidiinae Lawrence, 1991

1376Subfamily Ptilodactylinae Laporte, 1836

1377Family Podabrocephalidae Pic, 1930

1378Family Chelonariidae Blanchard, 1845

1379Family Eulichadidae Crowson, 1973

1380Family Callirhipidae Emden, 1924

1381Superfamily Elateroidea Leach, 1815

1382Family Rhinorhipidae Lawrence, 1988

1383Family Artematopodidae Lacordaire, 1857

1384Subfamily Electribiinae Crowson, 1975

1385Subfamily Allopogoniinae Crowson, 1973

1386Subfamily Artematopodinae Lacordaire, 1857

1387Tribe Artematopodini Lacordaire, 1857

1388Tribe Ctesibiini Crowson, 1973

1389Tribe Macropogonini LeConte, 1861

1390Family Brachypsectridae LeConte and Horn, 1883

1391Family Cerophytidae Latreille, 1834

1392Family Eucnemidae Eschscholtz, 1829

1393Subfamily Perothopinae Lacordaire, 1857

1394Subfamily Phyllocerinae Reitter, 1905

1395Tribe Anelastini Reitter, 1911

1396Tribe Phyllocerini Reitter, 1905

1397Subfamily Pseudomeninae Muona, 1993

1398Tribe Pseudomenini Muona, 1993

1399Tribe Schizophilini Muona, 1993

1400Subfamily Palaeoxeninae Muona, 1993

1401Subfamily Phlegoninae Muona, 1993

1402Subfamily Anischiinae Fleutiaux, 1936

1403Subfamily Melasinae Fleming, 1821

1404Tribe Calyptocerini Muona, 1993

1405Tribe Ceballosmelasini Muona, 1993

1406Tribe Dirhagini Reitter, 1911

1407Tribe Epiphanini Muona, 1993

1408Tribe Hylocharini Jacquelin du Val, 1859

1409Tribe Melasini Fleming, 1821

1410Subtribe Compsocnemina Muona, 1993

1411Subtribe Melasina Fleming, 1821

1412Tribe Neocharini Muona, 1993

1413Tribe Xylobiini Reitter, 1911

1414Subfamily Eucneminae Eschscholtz, 1829

1415Tribe Dendrocharini Fleutiaux, 1920

1416Tribe Dyscharachthini Muona, 1993

1417Tribe Entomosatopini Muona, 1993

1418Tribe Eucnemini Eschscholtz, 1829

1419Tribe Galbitini Muona, 1991

1420Tribe Mesogenini Muona, 1993

1421Tribe Muonajini Özdikmen, 2008

1422Tribe Perrotiini Muona, 1993

1423Tribe Phaenocerini Muona, 1993

1424Tribe Proutianini Muona, 1993

1425Subfamily Macraulacinae Fleutiaux, 1923

1426Tribe Anelastidini Muona, 1993

1427Tribe Echthrogasterini Cobos, 1965

1428Tribe Euryptychini Mamaev, 1976

1429Tribe Jenibuntorini Muona, 1993

1430Tribe Macraulacini Fleutiaux, 1923

1431Tribe Nematodini Leiler, 1976

1432Tribe Oisocerini Muona, 1993

1433Tribe Orodotini Muona, 1993

1434†Tribe Throscogeniini Iablokoff-Khnzorian, 1962

1435Family Throscidae Laporte, 1840 nomen protectum

1436†Family Praelateriidae Dolin, 1973

1437Family Elateridae Leach, 1815

1438Subfamily Cebrioninae Latreille, 1802

1439Subfamily Agrypninae Candèze, 1857 nomen protectum

1440Tribe Agrypnini Candèze, 1857 nomen protectum

1441Tribe Anaissini Golbach, 1984

1442†Tribe Cryptocardiini Dolin, 1980

1443Tribe Euplinthini Costa, 1975

1444Subtribe Cleidecostina Johnson, 2002

1445Subtribe Compsoplinthina Costa, 1975

1446Subtribe Euplinthina Costa, 1975

1447Tribe Hemirhipini Candèze, 1857

1448Tribe Oophorini Gistel, 1848

1449Tribe Platycrepidiini Costa and Casari-Chen, 1993

1450Tribe Pseudomelanactini Arnett, 1967

1451Tribe Pyrophorini Candèze, 1863

1452Subtribe Hapsodrilina Costa, 1975

1453Subtribe Nyctophyxina Costa, 1975

1454Subtribe Pyrophorina Candèze, 1863

1455Tribe Tetralobini Laporte, 1840

1456Subfamily Thylacosterninae Fleutiaux, 1920

1457Subfamily Lissominae Laporte, 1835

1458Subfamily Semiotinae Jakobson, 1913

1459Subfamily Campyloxeninae Costa, 1975

1460Subfamily Pityobiinae Hyslop, 1917

1461Subfamily Oxynopterinae Candèze, 1857

1462Subfamily Dendrometrinae Gistel, 1848

1463Tribe Crepidomenini Candèze, 1863

1464Tribe Dendrometrini Gistel, 1848

1465Subtribe Dendrometrina Gistel, 1848

1466Subtribe Denticollina Stein and Weise, 1877 (1848)

1467Subtribe Hemicrepidiina Champion, 1896

1468Tribe Dimini Candèze, 1863

1469Tribe Hypnoidini Schwarz, 1906 (1860)

1470Tribe Pleonomini Semenov and Pjatakova, 1936

1471Tribe Prosternini Gistel, 1856 nomen protectum

1472Tribe Senodoniini Schenkling, 1927

1473Subfamily Negastriinae Nakane and Kishii, 1956

1474Tribe Negastriini Nakane and Kishii, 1956

1475Tribe Quasimusini Schimmel and Tarnawski, 2009

1476Subtribe Loebliquasimusina Schimmel and Tarnawski, 2009

1477Subtribe Quasimusina Schimmel and Tarnawski, 2009

1478Subtribe Striatoquasimusina Schimmel and Tarnawski, 2009

1479Subtribe Wittmeroquasimusina Schimmel and Tarnawski, 2009

1480Subfamily Elaterinae Leach, 1815

1481Tribe Agriotini Laporte, 1840

1482Subtribe Agriotina Laporte, 1840

1483Subtribe Cardiorhinina Candèze, 1863

1484Tribe Ampedini Gistel, 1848

1485Tribe Dicrepidiini Thomson, 1858

1486Tribe Elaterini Leach, 1815

1487Tribe Megapenthini Gurjeva, 1973

1488Tribe Melanotini Candèze, 1859 (1848)

1489Tribe Odontonychini Girard, 1973

1490Tribe Physorhinini Candèze, 1859

1491Tribe Pomachiliini Candèze, 1859

1492Tribe Synaptini Gistel, 1856

1493Subfamily Cardiophorinae Candèze, 1859

1494Subfamily Hemiopinae Fleutiaux, 1941

1495Subfamily Physodactylinae Lacordaire, 1857

1496Subfamily Eudicronychinae Girard, 1971

1497Subfamily Subprotelaterinae Fleutiaux, 1920

1498Subfamily Morostomatinae Dolin, 2000

1499†Subfamily Protagrypninae Dolin, 1973

1500†Tribe Desmatini Dolin, 1975

1501†Tribe Hypnomorphini Dolin, 1975

1502†Tribe Protagrypnini Dolin, 1973

1503Family Plastoceridae Crowson, 1972

1504Family Drilidae Blanchard, 1845

1505Subfamily Drilinae Blanchard, 1845

1506Subfamily Thilmaninae Kazantsev, 2004

1507Tribe Euanomini Kazantsev, 2010

1508Tribe Thilmanini Kazantsev, 2004

1509Family Omalisidae Lacordaire, 1857

1510†Family Berendtimiridae Winkler, 1987

1511Family Lycidae Laporte, 1836

1512Subfamily Libnetinae Bocák and Bocáková, 1990

1513Subfamily Dictyopterinae Houlbert, 1922

1514Tribe Dictyopterini Houlbert, 1922

1515Tribe Lycoprogenthini Bocák and Bocáková, 2008

1516Tribe Taphini Bocák and Bocáková, 1990

1517Subfamily Lyropaeinae Bocák and Bocáková, 1989

1518Tribe Alyculini Bocák and Bocáková, 2008

1519Tribe Antennolycini Bocák and Bocáková, 2008

1520Tribe Lyropaeini Bocák and Bocáková, 1989

1521Tribe Miniduliticolini Kazantsev, 2003

1522Tribe Platerodrilini Kazantsev, 2004

1523Subfamily Ateliinae Kleine, 1929

1524Tribe Ateliini Kleine, 1929

1525Tribe Dilophotini Kleine, 1929

1526Subfamily Lycinae Laporte, 1836

1527Tribe Calochromini Lacordaire, 1857

1528Tribe Calopterini Green, 1949

1529Subtribe Acroleptina Bocáková, 2005

1530Subtribe Calopterina Green, 1949

1531Tribe Conderini Bocák and Bocáková, 1990

1532Tribe Dihammatini Bocák and Bocáková, 2008

1533Tribe Erotini LeConte, 1881

1534Tribe Eurrhacini Bocáková, 2005

1535Tribe Leptolycini Leng and Mutchler, 1922

1536Tribe Lycini Laporte, 1836

1537Tribe Lyponiini Bocák and Bocáková, 1990

1538Tribe Macrolycini Kleine, 1929

1539Tribe Melanerotini Kazantsev, 2010

1540Tribe Metriorrhynchini Kleine, 1926

1541Subtribe Hemiconderinina Bocák and Bocáková, 1990

1542Subtribe Metriorrhynchina Kleine, 1926

1543Subtribe Trichalina Kleine, 1929

1544Tribe Platerodini Kleine, 1929

1545Tribe Slipinskiini Bocák and Bocáková, 1992

1546Tribe Thonalmini Kleine, 1933

1547Subfamily Dexorinae Bocák and Bocáková, 1989

1548Family Telegeusidae Leng, 1920

1549Family Phengodidae LeConte, 1861

1550Subfamily Phengodinae LeConte, 1861

1551Subfamily Mastinocerinae LeConte, 1881

1552Subfamily Penicillophorinae Paulus, 1975

1553Family Rhagophthalmidae Olivier, 1907

1554Family Lampyridae Rafinesque, 1815

1555Subfamily Psilocladinae McDermott, 1964

1556Subfamily Amydetinae Olivier, 1907

1557Tribe Amydetini Olivier, 1907

1558Tribe Vestini McDermott, 1964

1559Subfamily Lampyrinae Rafinesque, 1815

1560Tribe Cratomorphini Green, 1948

1561Tribe Lamprocerini Olivier, 1907

1562Tribe Lamprohizini Kazantsev, 2010

1563Tribe Lampyrini Rafinesque, 1815

1564Tribe Lucidotini Lacordaire, 1857

1565Subtribe Dadophorina Olivier, 1907

1566Subtribe Lamprigerina McDermott, 1964

1567Subtribe Lucidotina Lacordaire, 1857

1568Subtribe Photinina LeConte, 1881

1569Tribe Pleotomini Summers, 1874

1570Subfamily Luciolinae Lacordaire, 1857

1571Tribe Curtosini McDermott, 1964

1572Tribe Luciolini Lacordaire, 1857

1573Subfamily Photurinae Lacordaire, 1857

1574Family Omethidae LeConte, 1861

1575Subfamily Omethinae LeConte, 1861

1576Subfamily Matheteinae LeConte, 1881

1577Subfamily Driloniinae Crowson, 1972

1578Family Cantharidae Imhoff, 1856 (1815)

1579Subfamily Cantharinae Imhoff, 1856 (1815)

1580Tribe Cantharini Imhoff, 1856 (1815)

1581Tribe Podabrini Gistel, 1856

1582Subfamily Silinae Mulsant, 1862

1583Tribe Silini Mulsant, 1862

1584Tribe Tytthonyxini Arnett, 1962

1585Subfamily Dysmorphocerinae Brancucci, 1980

1586Subfamily Malthininae Kiesenwetter, 1852

1587Tribe Malchinini Brancucci, 1980

1588Tribe Malthinini Kiesenwetter, 1852

1589Tribe Malthodini Böving and Craighead, 1931

1590Subfamily Chauliognathinae LeConte, 1861

1591Tribe Chauliognathini LeConte, 1861

1592Tribe Ichthyurini Champion, 1915

1593Subfamily Cydistinae Paulus, 1972

1594Subfamily Pterotinae LeConte, 1861

1595Subfamily Ototretinae McDermott, 1964

1596Subfamily Ototretadrilinae Crowson, 1972

1597†Subfamily Lasiosynidae Kirejtshuk, Chang, Ren and Kun, 2010

1598Series DERODONTIFORMIA

1599Superfamily Derodontoidea LeConte, 1861

1600Family Derodontidae LeConte, 1861

1601Subfamily Peltasticinae LeConte, 1861

1602Subfamily Derodontinae LeConte, 1861

1603Subfamily Laricobiinae Mulsant and Rey, 1864

1604Family Nosodendridae Erichson, 1846

1605Family Jacobsoniidae Heller, 1926

1606Series BOSTRICHIFORMIA

1607Superfamily Bostrichoidea Latreille, 1802

1608Family Dermestidae Latreille, 1804

1609Subfamily Dermestinae Latreille, 1804

1610Tribe Dermestini Latreille, 1804

1611Tribe Marioutini Jakobson, 1913

1612Subfamily Thorictinae Agassiz, 1846

1613Tribe Thaumaphrastini Anderson, 1949

1614Tribe Thorictini Agassiz, 1846

1615Subfamily Orphilinae LeConte, 1861

1616Subfamily Trinodinae Casey, 1900

1617†Tribe Cretonodini Kirejtshuk and Azar, 2009

1618Tribe Thylodriini Semenov, 1909

1619Tribe Trinodini Casey, 1900

1620Tribe Trinoparvini Háva, 2010

1621Subfamily Attageninae Laporte, 1840

1622Tribe Attagenini Laporte, 1840

1623Tribe Egidyellini Semenov, 1914

1624Subfamily Megatominae Leach, 1815

1625Tribe Anthrenini Gistel, 1848

1626Tribe Megatomini Leach, 1815

1627Family Endecatomidae LeConte, 1861

1628Family Bostrichidae Latreille, 1802

1629Subfamily Dysidinae Lesne, 1921

1630Subfamily Polycaoninae Lesne, 1896

1631Subfamily Bostrichinae Latreille, 1802

1632Tribe Apatini Billberg, 1820

1633Tribe Bostrichini Latreille, 1802

1634Tribe Dinapatini Lesne, 1910

1635Tribe Sinoxylini Marseul, 1857

1636Tribe Xyloperthini Lesne, 1921

1637Subfamily Psoinae Blanchard, 1851

1638Subfamily Dinoderinae Thomson, 1863

1639Subfamily Lyctinae Billberg, 1820

1640Tribe Lyctini Billberg, 1820

1641Tribe Trogoxylini Lesne, 1921

1642Subfamily Euderiinae Lesne, 1934

1643Family Ptinidae Latreille, 1802

1644Subfamily Eucradinae LeConte, 1861

1645Tribe Eucradini LeConte, 1861

1646Tribe Hedobiini Mulsant and Rey, 1868

1647Subfamily Ptininae Latreille, 1802

1648Tribe Gibbiini Jacquelin du Val, 1860

1649Tribe Meziini Bellés, 1985

1650Tribe Ptinini Latreille, 1802

1651Tribe Sphaericini Portevin, 1931

1652Subfamily Dryophilinae Gistel, 1848

1653Tribe Dryophilini Gistel, 1848

1654Tribe Ptilineurini Böving, 1927

1655Subfamily Ernobiinae Pic, 1912

1656Subfamily Anobiinae Fleming, 1821

1657Subfamily Ptilininae Shuckard, 1839

1658Subfamily Alvarenganiellinae Viana and Martínez, 1971

1659Subfamily Xyletininae Gistel, 1848

1660Tribe Lasiodermini Böving, 1927

1661Tribe Metholcini Zahradník, 2009

1662Tribe Xyletinini Gistel, 1848

1663Subfamily Dorcatominae Thomson, 1859

1664Subfamily Mesocoelopodinae Mulsant and Rey, 1864

1665Tribe Tricorynini White, 1971

1666Tribe Mesocoelopodini Mulsant and Rey, 1864

1667Series CUCUJIFORMIA

1668Superfamily Lymexyloidea Fleming, 1821

1669Family Lymexylidae Fleming, 1821

1670Subfamily Hylecoetinae Germar, 1818

1671Subfamily Lymexylinae Fleming, 1821

1672Subfamily Atractocerinae Laporte, 1840

1673Subfamily Melittommatinae Wheeler, 1986

1674Superfamily Cleroidea Latreille, 1802

1675Family Phloiophilidae Kiesenwetter, 1863

1676Family Trogossitidae Latreille, 1802

1677Subfamily Peltinae Latreille, 1806

1678Tribe Ancyronini Kolibáč, 2006

1679Tribe Colydiopeltini Kolibáč, 2006

1680Tribe Decamerini Crowson, 1964

1681Tribe Lophocaterini Crowson, 1964 nomen protectum

1682Tribe Peltini Latreille, 1806

1683Tribe Thymalini Léveillé, 1888

1684Subfamily Trogossitinae Latreille, 1802

1685Tribe Calityini Reitter, 1922

1686Tribe Egoliini Lacordaire, 1854

1687Tribe Gymnochilini Lacordaire, 1854

1688Tribe Larinotini Ślipiński, 1992

1689†Tribe Lithostomatini Kolibáč and Huang, 2008

1690Tribe Trogossitini Latreille, 1802

1691Family Chaetosomatidae Crowson, 1952

1692Family Metaxinidae Kolibáč, 2004

1693Family Thanerocleridae Chapin, 1924

1694Subfamily Zenodosinae Kolibáč, 1992

1695Subfamily Thaneroclerinae Chapin, 1924

1696Tribe Isoclerini Kolibáč, 1992

1697Tribe Thaneroclerini Chapin, 1924

1698Tribe Viticlerini Winkler, 1982

1699Family Cleridae Latreille, 1802

1700Subfamily Tillinae Fischer von Waldheim, 1813

1701Subfamily Hydnocerinae Spinola, 1844

1702Tribe Callimerini Kolibáč, 1998

1703Tribe Hydnocerini Spinola, 1844

1704Tribe Lemidiini Kolibáč, 1998

1705Subfamily Clerinae Latreille, 1802

1706Subfamily Korynetinae Laporte, 1836

1707Family Acanthocnemidae Crowson, 1964

1708Family Phycosecidae Crowson, 1952

1709Family Prionoceridae Lacordaire, 1857

1710Tribe Lobonychini Majer, 1987

1711Tribe Prionocerini Lacordaire, 1857

1712Family Mauroniscidae Majer, 1995

1713Family Melyridae Leach, 1815

1714Subfamily Rhadalinae LeConte, 1861

1715Subfamily Melyrinae Leach, 1815

1716Tribe Arthrobrachini Majer, 1987

1717Tribe Astylini Pic, 1929

1718Tribe Cerallini Pic, 1929

1719Tribe Melyrini Leach, 1815

1720Subfamily Dasytinae Laporte, 1840

1721Tribe Chaetomalachiini Majer, 1987

1722Tribe Danaceini Thomson, 1859

1723Tribe Dasytini Laporte, 1840

1724Tribe Gietellini Constantin and Menier, 1987

1725Tribe Listrini Majer, 1990

1726Subfamily Malachiinae Fleming, 1821

1727Tribe Amalthocini Majer, 2002

1728Tribe Attalomimini Majer, 1995

1729Tribe Carphurini Champion, 1923

1730Tribe Lemphini Wittmer, 1976

1731Tribe Malachiini Fleming, 1821

1732Tribe Pagurodactylini Constantin, 2001

1733Superfamily Cucujoidea Latreille, 1802

1734†Family Parandrexidae Kirejtshuk, 1994

1735†Family Sinisilvanidae Hong, 2002

1736Family Boganiidae Sen Gupta and Crowson, 1966

1737Subfamily Paracucujinae Endrödy-Younga and Crowson, 1986

1738Subfamily Boganiinae Sen Gupta and Crowson, 1966

1739Family Byturidae Gistel, 1848

1740Subfamily Platydascillinae Pic, 1914

1741Subfamily Byturinae Gistel, 1848

1742Family Helotidae Chapuis, 1876

1743Family Protocucujidae Crowson, 1954

1744Family Sphindidae Jacquelin du Val, 1860

1745Subfamily Protosphindinae Sen Gupta and Crowson, 1979

1746Subfamily Odontosphindinae Sen Gupta and Crowson, 1979

1747Subfamily Sphindiphorinae Sen Gupta and Crowson, 1979

1748Subfamily Sphindinae Jacquelin du Val, 1860

1749Family Biphyllidae LeConte, 1861

1750Family Erotylidae Latreille, 1802

1751Subfamily Xenoscelinae Ganglbauer, 1899

1752Subfamily Pharaxonothinae Crowson, 1952

1753Subfamily Loberinae Bruce, 1951

1754Subfamily Languriinae Hope, 1840

1755Tribe Hapalipini Leschen, 2003

1756Tribe Languriini Hope, 1840

1757Tribe Thallisellini Sen Gupta, 1968

1758Subfamily Cryptophilinae Casey, 1900

1759Tribe Cryptophilini Casey, 1900

1760Tribe Empocryptini Leschen, 2003

1761Tribe Toramini Sen Gupta, 1967

1762Subfamily Erotylinae Latreille, 1802

1763Tribe Dacnini Gistel, 1848

1764Tribe Encaustini Crotch, 1876

1765Tribe Erotylini Latreille, 1802

1766Tribe Megalodacnini Sen Gupta, 1970

1767Tribe Tritomini Curtis, 1834

1768Family Monotomidae Laporte, 1840

1769Subfamily Rhizophaginae Redtenbacher, 1845

1770Subfamily Monotominae Laporte, 1840

1771Tribe Europini Sen Gupta, 1988

1772Tribe Lenacini Crowson, 1952

1773Tribe Monotomini Laporte, 1840

1774†Tribe Rhizophtomini Kirejtshuk and Azar, 2009

1775Tribe Thionini Crowson, 1952

1776Family Hobartiidae Sen Gupta and Crowson, 1966

1777Family Cryptophagidae Kirby, 1826

1778Subfamily Cryptophaginae Kirby, 1826

1779Tribe Caenoscelini Casey, 1900

1780Tribe Cryptophagini Kirby, 1826

1781Tribe Picrotini Crowson, 1980

1782Subfamily Atomariinae LeConte, 1861

1783Tribe Atomariini LeConte, 1861

1784Tribe Cryptafricini Leschen, 1996

1785Tribe Hypocoprini Reitter, 1879

1786Family Agapythidae Sen Gupta and Crowson, 1969

1787Family Priasilphidae Crowson, 1973

1788Family Phloeostichidae Reitter, 1911

1789Family Silvanidae Kirby, 1837

1790Subfamily Brontinae Blanchard, 1845

1791Tribe Brontini Blanchard, 1845

1792Tribe Telephanini LeConte, 1861

1793Subfamily Silvaninae Kirby, 1837

1794Family Cucujidae Latreille, 1802

1795Family Myraboliidae Lawrence and Britton, 1991

1796Family Cavognathidae Sen Gupta and Crowson, 1966

1797Family Lamingtoniidae Sen Gupta and Crowson, 1969

1798Family Passandridae Blanchard, 1845

1799Family Phalacridae Leach, 1815

1800Subfamily Phaenocephalinae Matthews, 1899

1801Subfamily Phalacrinae Leach, 1815

1802Family Propalticidae Crowson, 1952

1803Family Laemophloeidae Ganglbauer, 1899

1804Family Tasmosalpingidae Lawrence and Britton, 1991

1805Family Cyclaxyridae Gimmel, Leschen and Ślipiński, 2009

1806Family Kateretida Kirby, 1837

1807Family Nitidulidae Latreille, 1802

1808Subfamily Calonecrinae Kirejtshuk, 1982

1809Subfamily Maynipeplinae Kirejtshuk, 1998

1810Subfamily Epuraeinae Kirejtshuk, 1986

1811Tribe Epuraeini Kirejtshuk, 1986

1812Tribe Taenioncini Kirejtshuk, 1998

1813Subfamily Carpophilinae Erichson, 1842

1814Subfamily Amphicrossinae Kirejtshuk, 1986

1815Subfamily Meligethinae Thomson, 1859

1816Subfamily Nitidulinae Latreille, 1802

1817Tribe Cychramini Gistel, 1848

1818Tribe Cychramptodini Kirejtshuk and Lawrence, 1992

1819Tribe Cyllodini Everts, 1898

1820Tribe Lawrencerosini Kirejtshuk, 1991

1821Tribe Mystropini Murray, 1864

1822Tribe Nitidulini Latreille, 1802

1823Subfamily Cillaeinae Kirejtshuk and Audisio, 1986

1824Subfamily Cryptarchinae Thomson, 1859

1825Tribe Arhinini Kirejtshuk, 1987

1826Tribe Cryptarchini Thomson, 1859

1827Tribe Eucalosphaerini Kirejtshuk, 1987

1828Tribe Platyarchini Kirejtshuk, 1998

1829Subfamily Cybocephalinae Jacquelin du Val, 1858

1830Family Smicripidae Horn, 1880

1831Family Bothrideridae Erichson, 1845

1832Subfamily Teredinae Seidlitz, 1888

1833Tribe Sosylopsini Dajoz, 1980

1834Tribe Sysolini Ślipiński and Pal, 1985

1835Tribe Teredini Seidlitz, 1888

1836Subfamily Xylariophilinae Pal and Lawrence, 1986

1837Subfamily Anommatinae Ganglbauer, 1899

1838Subfamily Bothriderinae Erichson, 1845

1839Family Cerylonidae Billberg, 1820

1840Subfamily Euxestinae Grouvelle, 1908

1841Subfamily Loeblioryloninae Ślipiński, 1990

1842Subfamily Ostomopsinae Sen Gupta and Crowson, 1973

1843Subfamily Murmidiinae Jacquelin du Val, 1858

1844Subfamily Ceryloninae Billberg, 1820

1845Family Alexiidae Imhoff, 1856

1846Family Discolomatidae Horn, 1878

1847Subfamily Notiophyginae Jakobson, 1915

1848Tribe Dystheamonini John, 1954

1849Tribe Notiophygini Jakobson, 1915

1850Tribe Pachyplacini John, 1954

1851Subfamily Discolomatinae Horn, 1878

1852Subfamily Aphanocephalinae Jakobson, 1904

1853Subfamily Cephalophaninae John, 1954

1854Subfamily Pondonatinae John, 1954

1855Family Endomychidae Leach, 1815

1856Subfamily Merophysiinae Seidlitz, 1872

1857Subfamily Pleganophorinae Jacquelin du Val, 1858

1858Subfamily Anamorphinae Strohecker, 1953

1859Subfamily Leiestinae Thomson, 1863

1860Subfamily Mycetaeinae Jacquelin du Val, 1857

1861Subfamily Eupsilobiinae Casey, 1895

1862Subfamily Xenomycetinae Strohecker, 1962

1863Subfamily Danascelinae Tomaszewska, 2000

1864Subfamily Endomychinae Leach, 1815

1865Subfamily Epipocinae Gorham, 1873

1866Subfamily Stenotarsinae Chapuis, 1876

1867Subfamily Lycoperdininae Bromhead, 1838

1868Family Coccinellidae Latreille, 1807

1869Subfamily Microweiseinae Leng, 1920

1870Tribe Microweiseini Leng, 1920

1871Tribe Serangiini Pope, 1962

1872Tribe Sukunahikonini Kamiya, 1960

1873Subfamily Coccinellinae Latreille, 1807

1874Tribe Argentipilosini Gordon and de Almeida, 1991

1875Tribe Aspidimerini Mulsant, 1850

1876Tribe Azyini Mulsant, 1850

1877Tribe Brachiacanthini Mulsant, 1850

1878Tribe Carinodulini Gordon, Pakaluk and Ślipiński, 1989

1879Tribe Cephaloscymnini Gordon, 1985

1880Tribe Chilocorini Mulsant, 1846

1881Tribe Chnoodini Mulsant, 1850

1882Tribe Coccidulini Mulsant, 1846

1883Tribe Coccinellini Latreille, 1807

1884Tribe Cranophorini Mulsant, 1850

1885Tribe Cryptognathini Mulsant, 1850

1886Tribe Cynegetini Thomson, 1866

1887Tribe Diomini Gordon, 1999

1888Tribe Epilachnini Mulsant, 1846

1889Tribe Epivertini Pang and Mao, 1979

1890Tribe Eremochilini Gordon and Vanderberg, 1987

1891Tribe Hyperaspidini Mulsant, 1846

1892Tribe Limnichopharini Miyatake, 1994

1893Tribe Monocorynini Miyatake, 1988

1894Tribe Noviini Mulsant, 1846

1895Tribe Ortaliini Mulsant, 1850

1896Tribe Oryssomini Gordon, 1974

1897Tribe Platynaspini Mulsant, 1846

1898Tribe Plotinini Miyatake, 1994

1899Tribe Poriini Mulsant, 1850

1900Tribe Scymnillini Casey, 1899

1901Tribe Scymnini Mulsant, 1846

1902Tribe Selvadiini Gordon, 1985

1903Tribe Shirozuellini Sasaji, 1967

1904Tribe Stethorini Dobzhansky, 1924

1905Tribe Sticholotidini Weise, 1901

1906Tribe Telsimiini Casey, 1899

1907Tribe Tetrabrachini Kapur, 1948

1908Family Corylophidae LeConte, 1852

1909Subfamily Periptyctinae Ślipiński, Lawrence and Tomaszewska, 2001

1910Subfamily Corylophinae LeConte, 1852

1911Tribe Aenigmaticini Casey, 1900

1912Tribe Cleidostethini Bowestead, Booth, Ślipiński and Lawrence, 2001

1913Tribe Corylophini LeConte, 1852

1914Tribe Foadiini Ślipiński, Tomaszewska and Lawrence, 2009

1915Tribe Orthoperini Jacquelin du Val, 1857

1916Tribe Parmulini Poey, 1854

1917Tribe Peltinodini Paulian, 1950

1918Tribe Rypobiini Paulian, 1950

1919Tribe Sericoderini Matthews, 1888

1920Tribe Teplinini Pakaluk, Ślipiński and Lawrence, 1994

1921Family Akalyptoischiidae Lord, Hartley, Lawrence, McHugh and Miller, 2010

1922Family Latridiidae Erichson, 1842

1923Subfamily Latridiinae Erichson, 1842

1924Subfamily Corticariinae Curtis, 1829

1925†Subfamily Tetrameropseinae Kirejtshuk and Azar, 2008

1926Superfamily Tenebrionoidea Latreille, 1802

1927Family Mycetophagidae Leach, 1815

1928Subfamily Esarcinae Reitter, 1882

1929Subfamily Mycetophaginae Leach, 1815

1930Tribe Mycetophagini Leach, 1815

1931Tribe Typhaeini Thomson, 1863

1932Subfamily Bergininae Leng, 1920

1933Family Archeocrypticidae Kaszab, 1964

1934Family Pterogeniidae Crowson, 1953

1935Family Ciidae Leach, 1819

1936Subfamily Sphindociinae Lawrence, 1974

1937Subfamily Ciinae Leach, 1819

1938Tribe Ciini Leach, 1819

1939Tribe Orophiini Thomson, 1863

1940Tribe Xylographellini Kawanabe and Miyatake, 1996

1941Subtribe Syncosmetina Lopes-Andrade, 2008

1942Subtribe Xylographellina Kawanabe and Miyatake, 1996

1943Family Tetratomidae Billberg, 1820

1944Subfamily Tetratominae Billberg, 1820

1945Subfamily Piseninae Miyatake, 1960

1946Subfamily Penthinae Lacordaire, 1859

1947Subfamily Hallomeninae Gistel, 1848

1948Subfamily Eustrophinae Gistel, 1848

1949Tribe Eustrophini Gistel, 1848

1950Tribe Holostrophini Nikitsky, 1998

1951Family Melandryidae Leach, 1815

1952Subfamily Melandryinae Leach, 1815

1953Tribe Anisoxiellini Nikitsky, 2007

1954Tribe Dircaeini Kirby, 1837

1955Tribe Hypulini Gistel, 1848

1956Tribe Melandryini Leach, 1815

1957Tribe Orchesiini Mulsant, 1856

1958Tribe Serropalpini Latreille, 1829

1959Tribe Xylitini Thomson, 1864

1960Tribe Zilorini Desbrochers des Loges, 1900

1961Subfamily Osphyinae Mulsant, 1856 (1839)

1962Family Mordellidae Latreille, 1802

1963Subfamily Ctenidiinae Franciscolo, 1951

1964Subfamily Mordellinae Latreille, 1802

1965Tribe Conaliini Ermisch, 1956

1966Tribe Mordellini Latreille, 1802

1967Tribe Mordellistenini Ermisch, 1941

1968Tribe Reynoldsiellini Franciscolo, 1957

1969Tribe Stenaliini Franciscolo, 1955

1970Family Ripiphoridae Gemminger, 1870 (1855)

1971Subfamily Ptilophorinae Gerstaecker, 1855

1972Subfamily Pelecotominae Seidlitz, 1875

1973Subfamily Hemirhipidiinae Heller, 1921

1974Subfamily Ripidiinae Gerstaecker, 1855

1975Tribe Eorhipidiini Iablokoff-Khnzorian, 1986

1976Tribe Ripidiini Gerstaecker, 1855

1977Subfamily Ripiphorinae Gemminger, 1870 (1855)

1978Tribe Macrosiagonini Heyden, 1908

1979Tribe Ripiphorini Gemminger, 1870 (1855)

1980Family Zopheridae Solier, 1834

1981Subfamily Colydiinae Billberg, 1820

1982Tribe Acropini Sharp, 1894

1983Tribe Adimerini Sharp, 1894

1984Tribe Colydiini Billberg, 1820

1985Tribe Gempylodini Sharp, 1893

1986Tribe Nematidiini Horn, 1878

1987Tribe Orthocerini Blanchard, 1845

1988Tribe Rhagoderini Horn, 1878

1989Tribe Rhopalocerini Reitter, 1911

1990Tribe Synchitini Erichson, 1845

1991Subfamily Zopherinae Solier, 1834

1992Tribe Latometini Ślipiński and Lawrence, 1999

1993Tribe Monommatini Blanchard, 1845

1994Tribe Phellopsini Ślipiński and Lawrence, 1999

1995Tribe Pycnomerini Erichson, 1845

1996Tribe Usechini Horn, 1867

1997Tribe Zopherini Solier, 1834

1998Family Ulodidae Pascoe, 1869

1999Family Promecheilidae Lacordaire, 1859

2000Family Chalcodryidae Watt, 1974

2001Family Trachelostenidae Lacordaire, 1859

2002Family Tenebrionidae Latreille, 1802

2003Subfamily Zolodininae Watt, 1975

2004Subfamily Lagriinae Latreille, 1825 (1820)

2005Tribe Adeliini Kirby, 1828

2006Tribe Belopini Reitter, 1917

2007Tribe Chaerodini Doyen, Matthews and Lawrence, 1990

2008Tribe Cossyphini Latreille, 1802

2009Tribe Goniaderini Lacordaire, 1859

2010Tribe Laenini Seidlitz, 1895

2011Tribe Lagriini Latreille, 1825 (1820)

2012Subtribe Lagriina Latreille, 1825 (1820)

2013Subtribe Statirina Blanchard, 1845

2014Tribe Lupropini Ardoin, 1958

2015Tribe Pycnocerini Lacordaire, 1859 nomen protectum

2016Subfamily Nilioninae Oken, 1843

2017Subfamily Phrenapatinae Solier, 1834

2018Tribe Archaeoglenini Watt, 1975

2019Tribe Penetini Lacordaire, 1859

2020Tribe Phrenapatini Solier, 1834

2021Subfamily Pimeliinae Latreille, 1802

2022Tribe Adelostomini Solier, 1834

2023Tribe Adesmiini Lacordaire, 1859 nomen protectum

2024Tribe Akidini Billberg, 1820

2025Tribe Anepsiini LeConte, 1862

2026Tribe Asidini Fleming, 1821

2027Tribe Boromorphini Skopin, 1978

2028Tribe Branchini LeConte, 1862

2029Tribe Caenocrypticini Koch, 1958

2030Tribe Ceratanisini Gebien, 1937

2031Tribe Cnemeplatiini Jacquelin du Val, 1861

2032Subtribe Actizetina Watt, 1992

2033Subtribe Cnemeplatiina Jacquelin du Val, 1861

2034Subtribe Rondoniellina Ferrer and Moragues, 2000

2035Subtribe Thorictosomatina Watt, 1992

2036Tribe Cnemodinini Gebien, 1910

2037Tribe Coniontini Waterhouse, 1858

2038Tribe Cossyphodini Wasmann, 1899

2039Subtribe Cossyphodina Wasmann, 1899

2040Subtribe Cossyphoditina Basilewsky, 1950

2041Subtribe Esemephina Steiner, 1980

2042Subtribe Paramellonina Andreae, 1961

2043Tribe Cryptochilini Solier, 1841

2044Subtribe Calognathina Lacordaire, 1859

2045Subtribe Cryptochilina Solier, 1841

2046Subtribe Homebiina Endrödy-Younga, 1989

2047Subtribe Horatomina Koch, 1955

2048Subtribe Vansoniina Koch, 1955

2049Tribe Cryptoglossini LeConte, 1862 nomen protectum

2050Tribe Edrotini Lacordaire, 1859

2051Tribe Elenophorini Solier, 1837

2052Tribe Epitragini Blanchard, 1845 nomen protectum

2053Tribe Erodiini Billberg, 1820 nomen protectum

2054Tribe Evaniosomini Lacordaire, 1859

2055Tribe Falsomycterini Gebien, 1910

2056Tribe Idisiini Medvedev, 1973

2057Tribe Klewariini Gebien, 1910

2058Tribe Kuhitangiini Medvedev, 1962

2059Tribe Lachnogyini Seidlitz, 1894

2060Subtribe Lachnodactylina Reitter, 1904

2061Subtribe Lachnogyina Seidlitz, 1894

2062Subtribe Netuschiliina Ferrer and Yvinec, 2004

2063Tribe Leptodini Lacordaire, 1859

2064Tribe Nycteliini Solier, 1834

2065Tribe Nyctoporini Lacordaire, 1859

2066Tribe Phrynocarenini Gebien, 1928

2067Tribe Physogasterini Lacordaire, 1859

2068Tribe Pimeliini Latreille, 1802

2069Tribe Praociini Eschscholtz, 1829

2070Tribe Sepidiini Eschscholtz, 1829

2071Subtribe Hypomelina Koch, 1955

2072Subtribe Molurina Solier, 1834

2073Subtribe Oxurina Koch, 1955

2074Subtribe Phanerotomeina Koch, 1958

2075Subtribe Sepidiina Eschscholtz, 1829

2076Subtribe Trachynotina Koch, 1955

2077Tribe Stenosini Schaum, 1859 (1834)

2078Tribe Tentyriini Eschscholtz, 1831

2079Tribe Thinobatini Lacordaire, 1859

2080Tribe Trilobocarini Lacordaire, 1859

2081Tribe Vacronini Gebien, 1910

2082Tribe Zophosini Solier, 1834

2083Subfamily Tenebrioninae Latreille, 1802

2084Tribe Acropteronini Doyen, 1989

2085Tribe Alphitobiini Reitter, 1917

2086Tribe Amarygmini Gistel, 1848

2087Tribe Amphidorini LeConte, 1862

2088Tribe Apocryphini Lacordaire, 1859

2089Tribe Blaptini Leach, 1815

2090Subtribe Blaptina Leach, 1815

2091Subtribe Gnaptorina Medvedev, 2001

2092Subtribe Gnaptorinina Medvedev, 2001

2093Subtribe Prosodina Skopin, 1960

2094Subtribe Remipedellina Semenov, 1907

2095Tribe Bolitophagini Kirby, 1837 nomen protectum

2096Tribe Centronopini Doyen, 1989

2097Tribe Cerenopini Horn, 1870

2098Tribe Dissonomini Medvedev, 1968

2099Tribe Eulabini Horn, 1870

2100Tribe Falsocossyphini Ferrer, 2006

2101Tribe Heleini Fleming, 1821

2102Subtribe Asphalina Matthews and Lawrence, 2005

2103Subtribe Cyphaleina Lacordaire, 1859

2104Subtribe Heleina Fleming, 1821

2105Tribe Helopini Latreille, 1802

2106Subtribe Helopina Latreille, 1802

2107Subtribe Cylindrinotina Español, 1956

2108Tribe Helopinini Lacordaire, 1859

2109Subtribe Aptilina Koch, 1958

2110Subtribe Helopinina Lacordaire, 1859

2111Subtribe Micrantereina Reitter, 1917

2112Subtribe Oncosomina Koch, 1958

2113Tribe Melanimonini Seidlitz, 1894 (1854)

2114Tribe Opatrini Brullé, 1832

2115Subtribe Heterocheirina Koch, 1956

2116Subtribe Heterotarsina Blanchard, 1845

2117Subtribe Opatrina Brullé, 1832

2118Subtribe Neopachypterina Bouchard, Löbl and Merkl, 2007

2119Tribe Palorini Matthews, 2003

2120Tribe Pedinini Eschscholtz, 1829

2121Subtribe Dendarina Mulsant and Rey, 1854

2122Subtribe Eurynotina Mulsant and Rey, 1854

2123Subtribe Leichenina Mulsant, 1854

2124Subtribe Loensina Koch, 1956

2125Subtribe Melambiina Mulsant and Rey, 1854

2126Subtribe Pedinina Eschscholtz, 1829

2127Subtribe Platynotina Mulsant and Rey, 1853

2128Subtribe Pythiopina Koch, 1953

2129Tribe Platyscelidini Lacordaire, 1859

2130Tribe Praeugenini De Moor, 1970

2131Tribe Rhysopaussini Wasmann, 1896

2132Tribe Scaurini Billberg, 1820

2133Tribe Scotobiini Solier, 1838

2134Tribe Tenebrionini Latreille, 1802

2135Tribe Titaenini Fauvel, 1905

2136Tribe Toxicini Oken, 1843

2137Subtribe Eudysantina Bouchard, Lawrence, Davies and Newton, 2005

2138Subtribe Nycteropina Lacordaire, 1859

2139Subtribe Toxicina Oken, 1843

2140Tribe Triboliini Gistel, 1848

2141Tribe Ulomini Blanchard, 1845

2142Subfamily Alleculinae Laporte, 1840

2143Tribe Alleculini Laporte, 1840

2144Subtribe Alleculina Laporte, 1840

2145Subtribe Gonoderina Seidlitz, 1896

2146Subtribe Mycetocharina Gistel, 1848

2147Subtribe Xystropodina Solier, 1835

2148Tribe Cteniopodini Solier, 1835

2149Subfamily Diaperinae Latreille, 1802

2150Tribe Crypticini Brullé, 1832

2151Tribe Diaperini Latreille, 1802

2152Subtribe Adelinina LeConte, 1862

2153Subtribe Diaperina Latreille, 1802

2154Tribe Ectychini Doyen, Matthews and Lawrence, 1990

2155Tribe Gnathidiini Gebien, 1921

2156Subtribe Anopidiina Jeannel and Paulian, 1945

2157Subtribe Gnathidiina Gebien, 1921

2158Tribe Hyociini Medvedev and Lawrence, 1982

2159Subtribe Brittonina Medvedev and Lawrence, 1986

2160Subtribe Hyociina Medvedev and Lawrence, 1982

2161Subtribe Uptonina Medvedev and Lawrence, 1986

2162Tribe Hypophlaeini Billberg, 1820

2163Tribe Leiochrinini Lewis, 1894

2164Tribe Myrmechixenini Jacquelin du Val, 1858

2165Tribe Phaleriini Blanchard, 1845

2166Tribe Scaphidemini Reitter, 1922

2167Tribe Trachyscelini Blanchard, 1845

2168Subfamily Stenochiinae Kirby, 1837

2169Tribe Cnodalonini Oken, 1843

2170Tribe Stenochiini Kirby, 1837

2171Tribe Talanini Champion, 1887 (1883)

2172Family Prostomidae Thomson, 1859

2173Family Synchroidae Lacordaire, 1859

2174Family Stenotrachelidae Thomson, 1859

2175Subfamily Stenotrachelinae Thomson, 1859

2176Subfamily Cephaloinae LeConte, 1862

2177Subfamily Nematoplinae LeConte, 1862

2178Subfamily Stoliinae Nikitsky, 1985

2179Family Oedemeridae Latreille, 1810

2180Subfamily Polypriinae Lawrence, 2005

2181Subfamily Calopodinae Costa, 1852 nomen protectum

2182Subfamily Oedemerinae Latreille, 1810

2183Tribe Asclerini Gistel, 1848

2184Tribe Ditylini Mulsant, 1858

2185Tribe Nacerdini Mulsant, 1858

2186Tribe Oedemerini Latreille, 1810

2187Tribe Stenostomatini Mulsant, 1858

2188Family Meloidae Gyllenhal, 1810

2189Subfamily Eleticinae Wellman, 1910

2190Tribe Derideini Wellman, 1910

2191Tribe Eleticini Wellman, 1910

2192Subtribe Eleticina Wellman, 1910

2193Subtribe Eospastina Selander, 1966

2194Tribe Ertlianini Selander, 1966

2195Tribe Spasticini Kaszab, 1959

2196Subtribe Anthicoxenina Selander, 1966

2197Subtribe Protomeloina Abdullah, 1965

2198Subtribe Spasticina Kaszab, 1959

2199Subtribe Xenospastina Selander, 1966

2200Subfamily Meloinae Gyllenhal, 1810

2201Tribe Cerocomini Leach, 1815

2202Tribe Epicautini Parker and Böving, 1924

2203Tribe Eupomphini LeConte, 1862

2204Tribe Lyttini Solier, 1851

2205Tribe Meloini Gyllenhal, 1810

2206Tribe Mylabrini Rafinesque, 1815

2207Tribe Pyrotini MacSwain, 1956

2208Subfamily Tetraonycinae Böving and Craighead, 1931

2209Subfamily Nemognathinae Laporte, 1840

2210Tribe Horiini Latreille, 1802

2211Tribe Nemognathini Laporte, 1840

2212Subtribe Nemognathina Laporte, 1840

2213Subtribe Zonitidina Mulsant, 1857

2214Subtribe Sitarina Mulsant, 1857

2215Tribe Stenoderini Selander, 1991

2216Family Mycteridae Oken, 1843

2217Subfamily Mycterinae Oken, 1843

2218Subfamily Eurypinae Thomson, 1860

2219Subfamily Hemipeplinae Lacordaire, 1854

2220Family Boridae Thomson, 1859

2221Subfamily Borinae Thomson, 1859

2222Subfamily Synercticinae Lawrence and Pollock, 1994

2223Family Trictenotomidae Blanchard, 1845

2224Family Pythidae Solier, 1834

2225Family Pyrochroidae Latreille, 1806

2226Subfamily Tydessinae Nikitsky, 1986

2227Subfamily Pilipalpinae Abdullah, 1964

2228Subfamily Pedilinae Lacordaire, 1859

2229Subfamily Pyrochroinae Latreille, 1806

2230Subfamily Agnathinae Lacordaire, 1859

2231Family Salpingidae Leach, 1815

2232Subfamily Othniinae LeConte, 1861

2233Subfamily Prostominiinae Grouvelle, 1914

2234Subfamily Agleninae Horn, 1878

2235Subfamily Inopeplinae Grouvelle, 1908

2236Subfamily Dacoderinae LeConte, 1862

2237Subfamily Aegialitinae LeConte, 1862

2238Subfamily Salpinginae Leach, 1815

2239Family Anthicidae Latreille, 1819

2240Subfamily Eurygeniinae LeConte, 1862

2241Tribe Eurygeniini LeConte, 1862

2242Tribe Ictistygnini Borchmann, 1936

2243Tribe Mitraelabrini Abdullah, 1969

2244Subfamily Macratriinae LeConte, 1862

2245Tribe Macratriini LeConte, 1862

2246†Tribe Camelomorphini Kirejtshuk and Azar, 2008

2247Subfamily Steropinae Jacquelin du Val, 1863

2248Subfamily Copobaeninae Abdullah, 1969

2249Subfamily Lemodinae Lawrence and Britton, 1991

2250Subfamily Tomoderinae Bonadona, 1961

2251Subfamily Anthicinae Latreille, 1819

2252Tribe Anthicini Latreille, 1819

2253Tribe Endomiini Kaszab, 1956

2254Tribe Formicomini Bonadona, 1974

2255Tribe Microhoriini Bonadona, 1974

2256Subfamily Notoxinae Stephens, 1829

2257Family Aderidae Csiki, 1909

2258Tribe Aderini Csiki, 1909

2259Subtribe Aderina Csiki, 1909

2260Subtribe Cnopina Báguena Corella, 1948

2261Subtribe Gompeliina Bouchard, 2011

2262Subtribe Syzetoninina Báguena Corella, 1948

2263Tribe Emelinini Báguena Corella, 1948

2264Tribe Euglenesini Seidlitz, 1875

2265Subtribe Euglenesina Seidlitz, 1875

2266Subtribe Pseudolotelina Báguena Corella, 1948

2267Tribe Phytobaenini Báguena Corella, 1948

2268Family Scraptiidae Gistel, 1848

2269Subfamily Scraptiinae Gistel, 1848

2270Tribe Allopodini Franciscolo, 1964

2271Tribe Scraptiini Gistel, 1848

2272Subfamily Anaspidinae Mulsant, 1856

2273Tribe Anaspidini Mulsant, 1856

2274Tribe Anaspimordini Franciscolo, 1954

2275Tribe Menuthianaspidini Franciscolo, 1972

2276Tribe Pentariini Franciscolo, 1954

2277Subfamily Lagrioidinae Abdullah and Abdullah, 1968

2278Subfamily Afreminae Levey, 1985

2279Subfamily Ischaliinae Blair, 1920

2280Superfamily Chrysomeloidea Latreille, 1802

2281Family Oxypeltidae Lacordaire, 1868

2282Family Vesperidae Mulsant, 1839

2283Subfamily Philinae Thomson, 1861

2284Subfamily Vesperinae Mulsant, 1839

2285Subfamily Anoplodermatinae Guérin-Méneville, 1840

2286Tribe Anoplodermatini Guérin-Méneville, 1840

2287Tribe Hypocephalini Blanchard, 1845

2288Tribe Mysteriini Prosen, 1960

2289Family Disteniidae Thomson, 1861

2290Tribe Cyrtonopini Gressitt, 1940

2291Tribe Disteniini Thomson, 1861

2292Tribe Dynamostini Lacordaire, 1868

2293Tribe Heteropalpini Villiers, 1961

2294Family Cerambycidae Latreille, 1802

2295Subfamily Parandrinae Blanchard, 1845

2296Tribe Erichsoniini Thomson, 1861

2297Tribe Parandrini Blanchard, 1845

2298Subfamily Prioninae Latreille, 1802

2299Tribe Acanthophorini Thomson, 1864

2300Tribe Aegosomatini Thomson, 1861

2301Tribe Anacolini Thomson, 1857

2302Tribe Cacoscelini Thomson, 1861

2303Tribe Callipogonini Thomson, 1861

2304Tribe Calocomini Galileo and Martins, 1993

2305Tribe Cantharocnemini Thomson, 1861

2306Tribe Ergatini Fairmaire, 1864

2307Tribe Eurypodini Gahan, 1906 (1868)

2308Tribe Hopliderini Thomson, 1864

2309Tribe Macrodontiini Thomson, 1861

2310Tribe Macrotomini Thomson, 1861

2311Subtribe Archetypina Lameere, 1912

2312Subtribe Basitoxina Lameere, 1912

2313Subtribe Macrotomina Thomson, 1861

2314Subtribe Platygnathina Gilmour, 1954

2315Subtribe Xixuthrina Lameere, 1912

2316Tribe Mallaspini Thomson, 1861

2317Tribe Mallodonini Thomson, 1861

2318Tribe Meroscelisini Thomson, 1861

2319Tribe Prionini Latreille, 1802

2320Tribe Remphanini Lacordaire, 1868

2321Tribe Solenopterini Lacordaire, 1868

2322Tribe Tereticini Lameere, 1913

2323Tribe Vesperoctenini Vives, 2005

2324Subfamily Lepturinae Latreille, 1802

2325Tribe Desmocerini Blanchard, 1845

2326Tribe Encyclopini LeConte, 1873

2327Tribe Lepturini Latreille, 1802

2328Tribe Oxymirini Danilevsky, 1997

2329Tribe Rhagiini Kirby, 1837

2330Tribe Rhamnusiini Sama, 2009

2331Tribe Teledapini Pascoe, 1871

2332Tribe Sachalinobiini Danilevsky, 2010

2333Tribe Xylosteini Reitter, 1913

2334Subfamily Spondylidinae Audinet-Serville, 1832

2335Tribe Anisarthrini Mamaev and Danilevsky, 1973

2336Tribe Asemini Thomson, 1861

2337Tribe Atimiini LeConte, 1873

2338Tribe Saphanini Gistel, 1848

2339Tribe Spondylidini Audinet-Serville, 1832

2340Subfamily Necydalinae Latreille, 1825

2341Subfamily Dorcasominae Lacordaire, 1868

2342Subfamily Apatophyseinae Lacordaire, 1869

2343Subfamily Cerambycinae Latreille, 1802

2344Tribe Acangassuini Galileo and Martins, 2001

2345Tribe Achrysonini Lacordaire, 1868

2346Tribe Agallissini LeConte, 1873

2347Tribe Alanizini Di Iorio, 2003

2348Tribe Anaglyptini Lacordaire, 1868 nomen protectum

2349Tribe Aphanasiini Lacordaire, 1868

2350Tribe Aphneopini Lacordaire, 1868

2351Tribe Auxesini Lepesme and Breuning, 1952

2352Tribe Basipterini Fragoso, Monné and Campos Seabra, 1987

2353Tribe Bimiini Lacordaire, 1868

2354Tribe Bothriospilini Lane, 1950

2355Tribe Brachypteromatini Sama, 2008

2356Tribe Callichromatini Swainson, 1840

2357Tribe Callidiini Kirby, 1837

2358Tribe Callidiopini Lacordaire, 1868

2359Tribe Cerambycini Latreille, 1802

2360Subtribe Cerambycina Latreille, 1802

2361Subtribe Sphallotrichina Martins and Monné, 2005

2362Tribe Certallini Fairmaire, 1864

2363Tribe Chlidonini Waterhouse, 1879

2364Tribe Cleomenini Lacordaire, 1868

2365Tribe Clytini Mulsant, 1839

2366Tribe Compsocerini Thomson, 1864

2367Tribe Coptommatini Lacordaire, 1869

2368Tribe Curiini LeConte, 1873

2369Tribe Deilini Fairmaire, 1864

2370Tribe Dejanirini Lacordaire, 1868

2371Tribe Diorini Lane, 1950

2372Tribe Distichocerini Pascoe, 1867

2373Tribe Dodecosini Aurivillius, 1912

2374Tribe Dryobiini Arnett, 1962 nomen protectum

2375Tribe Eburiini Blanchard, 1845

2376Tribe Ectenessini Martins, 1998

2377Tribe Elaphidiini Thomson, 1864

2378Tribe Eligmodermini Lacordaire, 1868

2379Tribe Erlandiini Aurivillius, 1912

2380Tribe Eroschemini Lacordaire, 1868

2381Tribe Eumichthini Linsley, 1940

2382Tribe Gahaniini Quentin and Villiers, 1969

2383Tribe Glaucytini Lacordaire, 1868

2384Tribe Graciliini Mulsant, 1839

2385Tribe Hesperophanini Mulsant, 1839

2386Subtribe Daramina Sama, 2008

2387Subtribe Hesperophanina Mulsant, 1839

2388Tribe Hesthesini Pascoe, 1867

2389Tribe Heteropsini Lacordaire, 1868 nomen protectum

2390Tribe Hexoplini Martins, 2006

2391Tribe Holopleurini Chemsak and Linsley, 1974

2392Tribe Holopterini Lacordaire, 1868

2393Tribe Hyboderini Linsley, 1940

2394Tribe Hylotrupini Zagajkevich, 1991

2395Tribe Ibidionini Thomson, 1861

2396Subtribe Compsina Martins and Galileo, 2007

2397Subtribe Ibidionina Thomson, 1861

2398Subtribe Tropidina Martins and Galileo, 2007

2399Tribe Ideratini Martins and Napp, 2009

2400Tribe Lissonotini Swainson, 1840

2401Tribe Luscosmodicini Martins, 2003

2402Tribe Lygrini Sama, 2008

2403Tribe Macronini Lacordaire, 1868

2404Tribe Megacoelini Quentin and Villiers, 1969

2405Tribe Methiini Thomson, 1860

2406Tribe Molorchini Gistel, 1848

2407Tribe Mythodini Lacordaire, 1868

2408Tribe Necydalopsini Lacordaire, 1868

2409Tribe Neocorini Martins, 2005

2410Tribe Neostenini Lacordaire, 1868

2411Tribe Obriini Mulsant, 1839

2412Tribe Ochyrini Pascoe, 1871

2413Tribe Oedenoderini Aurivillius, 1912

2414Tribe Oemini Lacordaire, 1868

2415Subtribe Methioidina Martins, 1997

2416Subtribe Oemina Lacordaire, 1868

2417Tribe Opsimini LeConte, 1873

2418Tribe Oxycoleini Martins and Galileo, 2003

2419Tribe Paraholopterini Martins, 1997

2420Tribe Phalotini Lacordaire, 1868

2421Tribe Phlyctaenodini Lacordaire, 1868

2422Tribe Phoracanthini Newman, 1840

2423Tribe Phyllarthriini Lepesme and Breuning, 1956

2424Tribe Piesarthriini McKeown, 1947

2425Tribe Piezocerini Lacordaire, 1868

2426Subtribe Haruspicina Martins, 1976

2427Subtribe Piezocerina Lacordaire, 1868

2428Tribe Platyarthrini Bates, 1870

2429Tribe Plectogasterini Quentin and Villiers, 1969

2430Tribe Plectromerini Nearns and Braham, 2008

2431Tribe Pleiarthrocerini Lane, 1950

2432Tribe Protaxini Gahan, 1906

2433Tribe Prothemini Lacordaire, 1868

2434Tribe Psebiini Lacordaire, 1868

2435Tribe Pseudocephalini Aurivillius, 1912 (1861)

2436Tribe Pseudolepturini Thomson, 1861

2437Tribe Psilomorphini Lacordaire, 1868

2438Tribe Pteroplatini Thomson, 1861

2439Tribe Pyrestini Lacordaire, 1868

2440Tribe Rhagiomorphini Newman, 1841

2441Tribe Rhinotragini Thomson, 1861

2442Tribe Rhopalophorini Blanchard, 1845

2443Tribe Rosaliini Fairmaire, 1864

2444Tribe Sestyrini Lacordaire, 1868

2445Tribe Smodicini Lacordaire, 1868

2446Tribe Spintheriini Lacordaire, 1869

2447Tribe Stenhomalini Miroshnikov, 1989

2448Tribe Stenoderini Pascoe, 1867

2449Tribe Stenopterini Gistel, 1848

2450Tribe Strongylurini Lacordaire, 1868

2451Tribe Tessarommatini Lacordaire, 1868

2452Tribe Thraniini Gahan, 1906

2453Tribe Thyrsiini Marinoni and Napp, 1984

2454Tribe Tillomorphini Lacordaire, 1868

2455Tribe Torneutini Thomson, 1861

2456Tribe Trachyderini Dupont, 1836

2457Subtribe Ancylocerina Thomson, 1864

2458Subtribe Trachyderina Dupont, 1836

2459Tribe Tragocerini Pascoe, 1867

2460Tribe Trichomesiini Aurivillius, 1912

2461Tribe Tropocalymmatini Lacordaire, 1868

2462Tribe Typhocesini Lacordaire, 1868

2463Tribe Unxiini Napp, 2007

2464Tribe Uracanthini Blanchard, 1853

2465Tribe Vesperellini Sama, 2008

2466Tribe Xystrocerini Blanchard, 1845

2467Subfamily Lamiinae Latreille, 1825

2468Tribe Acanthocinini Blanchard, 1845

2469Tribe Acanthoderini Thomson, 1860

2470Tribe Acmocerini Thomson, 1864

2471Tribe Acridocephalini Dillon and Dillon, 1959

2472Tribe Acrocinini Swainson, 1840

2473Tribe Aderpasini Breuning and Teocchi, 1978

2474Tribe Aerenicini Lacordaire, 1872

2475Tribe Agapanthiini Mulsant, 1839

2476Tribe Amphoecini Breuning, 1951

2477Tribe Ancitini Aurivillius, 1917

2478Tribe Ancylonotini Lacordaire, 1869

2479Tribe Anisocerini Thomson, 1860

2480Tribe Apomecynini Thomson, 1860

2481Tribe Astathini Thomson, 1864

2482Tribe Batocerini Thomson, 1864

2483Tribe Calliini Thomson, 1864

2484Tribe Ceroplesini Thomson, 1860

2485Subtribe Ceroplesina Thomson, 1860

2486Subtribe Crossotina Thomson, 1864

2487Tribe Cloniocerini Lacordaire, 1872

2488Tribe Colobotheini Thomson, 1860

2489Tribe Compsosomatini Thomson, 1857

2490Tribe Cyrtinini Thomson, 1864

2491Tribe Desmiphorini Thomson, 1860

2492Tribe Dorcadionini Swainson, 1840

2493Tribe Dorcaschematini Thomson, 1860

2494Tribe Elytracanthinini Bousquet, 2009

2495Tribe Enicodini Thomson, 1864

2496Tribe Eupromerini Galileo and Martins, 1995

2497Tribe Forsteriini Tippmann, 1960

2498Tribe Gnomini Thomson, 1860

2499Tribe Gyaritini Breuning, 1950

2500Tribe Heliolini Breuning, 1951

2501Tribe Hemilophini Thomson, 1868 nomen protectum

2502Tribe Homonoeini Thomson, 1864

2503Tribe Hyborhabdini Aurivillius, 1911

2504Tribe Lamiini Latreille, 1825

2505Tribe Laticraniini Lane, 1959

2506Tribe Mauesiini Lane, 1956

2507Tribe Megabasini Thomson, 1860

2508Tribe Mesosini Mulsant, 1839

2509Tribe Microcymaturini Breuning and Teocchi, 1985

2510Tribe Moneilemini Thomson, 1864

2511Tribe Monochamini Gistel, 1848

2512Tribe Morimonellini Lobanov, Danilevsky and Murzin, 1981

2513Tribe Morimopsini Lacordaire, 1869

2514Tribe Nyctimeniini Gressitt, 1951

2515Tribe Obereini Thomson, 1864

2516Tribe Oculariini Breuning, 1950

2517Tribe Onciderini Thomson, 1860

2518Tribe Oncideropsidini Aurivillius, 1922

2519Tribe Onocephalini Thomson, 1860

2520Tribe Onychogleneini Aurivillius, 1923

2521Tribe Parmenini Mulsant, 1839

2522Tribe Petrognathini Blanchard, 1845

2523Tribe Phacellini Lacordaire, 1872

2524Tribe Phantasini Kolbe, 1897

2525Tribe Phrynetini Thomson, 1864

2526Tribe Phymasternini Teocchi, 1989

2527Tribe Phytoeciini Mulsant, 1839

2528Tribe Pogonocherini Mulsant, 1839

2529Tribe Polyrhaphidini Thomson, 1860

2530Tribe Pretiliini Martins and Galileo, 1990

2531Tribe Proctocerini Aurivillius, 1922

2532Tribe Prosopocerini Thomson, 1864

2533Tribe Pteropliini Thomson, 1860

2534Tribe Rhodopinini Gressitt, 1951

2535Tribe Saperdini Mulsant, 1839

2536Tribe Stenobiini Breuning, 1950

2537Tribe Sternotomini Thomson, 1860

2538Tribe Tapeinini Thomson, 1857

2539Tribe Tetraopini Thomson, 1860

2540Tribe Tetraulaxini Breuning and Teocchi, 1977

2541Tribe Tetropini Portevin, 1927

2542Tribe Theocrini Lacordaire, 1872

2543Tribe Tmesisternini Blanchard, 1853

2544Tribe Tragocephalini Thomson, 1857

2545Tribe Xenicotelini Matsushita, 1933

2546Tribe Xenofreini Aurivillius, 1923

2547Tribe Xenoleini Lacordaire, 1872

2548Tribe Xylorhizini Lacordaire, 1872

2549Tribe Zygocerini Thomson, 1864

2550Family Megalopodidae Latreille, 1802

2551Subfamily Megalopodinae Latreille, 1802

2552Subfamily Palophaginae Kuschel and May, 1990

2553Subfamily Zeugophorinae Böving and Craighead, 1931

2554Family Orsodacnidae Thomson, 1859

2555Subfamily Orsodacninae Thomson, 1859

2556Subfamily Aulacoscelidinae Chapuis, 1874

2557Family Chrysomelidae Latreille, 1802

2558Subfamily Sagrinae Leach, 1815

2559Tribe Carpophagini Chapuis, 1874

2560Tribe Diaphanopsidini Monrós, 1958

2561Tribe Megamerini Chapuis, 1874

2562Tribe Sagrini Leach, 1815

2563Subfamily Bruchinae Latreille, 1802

2564Tribe Amblycerini Bridwell, 1932

2565Subtribe Amblycerina Bridwell, 1932

2566Subtribe Spermophagina Borowiec, 1987

2567Tribe Bruchini Latreille, 1802

2568Subtribe Acanthoscelidina Bridwell, 1946

2569Subtribe Bruchina Latreille, 1802

2570Subtribe Megacerina Bridwell, 1946

2571Tribe Eubaptini Bridwell, 1932

2572Tribe Kytorhinini Bridwell, 1932

2573Tribe Pachymerini Bridwell, 1929

2574Subtribe Caryedontina Bridwell, 1929

2575Subtribe Caryopemina Bridwell, 1929

2576Subtribe Pachymerina Bridwell, 1929

2577Tribe Rhaebini Blanchard, 1845

2578Subfamily Donaciinae Kirby, 1837

2579Tribe Donaciini Kirby, 1837

2580Tribe Haemoniini Chen, 1941

2581Tribe Plateumarini Böving, 1922

2582Subfamily Criocerinae Latreille, 1804

2583Tribe Criocerini Latreille, 1804

2584Tribe Lemini Gyllenhal, 1813

2585Tribe Pseudocriocerini Heinze, 1962

2586Subfamily Cassidinae Gyllenhal, 1813

2587Tribe Alurnini Chapuis, 1875

2588Tribe Anisoderini Chapuis, 1875

2589Tribe Aproidini Weise, 1911

2590Tribe Arescini Chapuis, 1875

2591Tribe Aspidimorphini Chapuis, 1875

2592Tribe Basiprionotini Gressitt, 1952 (1929)

2593Tribe Botryonopini Chapuis, 1875

2594Tribe Callispini Chapuis, 1875

2595Tribe Callohispini Uhmann, 1960

2596Tribe Cassidini Gyllenhal, 1813

2597Tribe Cephaloleiini Chapuis, 1875

2598Tribe Chalepini Weise, 1910

2599Tribe Coelaenomenoderini Weise, 1911

2600Tribe Cryptonychini Chapuis, 1875

2601Tribe Delocraniini Spaeth, 1929

2602Tribe Dorynotini Monrós and Viana, 1949 (1923)

2603Tribe Eugenysini Hincks, 1952

2604Tribe Eurispini Chapuis, 1875

2605Tribe Exothispini Weise, 1911

2606Tribe Goniocheniini Spaeth, 1942

2607Tribe Gonophorini Chapuis, 1875

2608Tribe Hemisphaerotini Monrós and Viana, 1951 (1929)

2609Tribe Hispini Gyllenhal, 1813

2610Tribe Hispoleptini Chapuis, 1875

2611Tribe Hybosispini Weise, 1910

2612Tribe Imatidiini Hope, 1840

2613Tribe Ischyrosonychini Chapuis, 1875

2614Tribe Leptispini Fairmaire, 1868

2615Tribe Mesomphaliini Hope, 1840

2616Tribe Nothosacanthini Gressitt, 1952 (1929)

2617Tribe Oediopalpini Monrós and Viana, 1947 (1910)

2618Tribe Omocerini Hincks, 1952 (1923)

2619Tribe Oncocephalini Chapuis, 1875

2620Tribe Promecothecini Chapuis, 1875

2621Tribe Prosopodontini Weise, 1910

2622Tribe Sceloenoplini Uhmann, 1930

2623Tribe Spilophorini Chapuis, 1875

2624Tribe Uroplatini Weise, 1910

2625Subfamily Chrysomelinae Latreille, 1802

2626Tribe Chrysomelini Latreille, 1802

2627Tribe Timarchini Motschulsky, 1860

2628Subfamily Galerucinae Latreille, 1802

2629Tribe Alticini Newman, 1834

2630Tribe Decarthrocerini Laboissière, 1937

2631Tribe Galerucini Latreille, 1802

2632Tribe Hylaspini Chapuis, 1875

2633Tribe Luperini Gistel, 1848

2634Tribe Metacyclini Chapuis, 1875

2635Tribe Oidini Laboissière, 1921 (1875)

2636Subfamily Lamprosomatinae Lacordaire, 1848

2637Tribe Lamprosomatini Lacordaire, 1848

2638Tribe Neochlamysini Monrós, 1959

2639Tribe Sphaerocharini Chapuis, 1874

2640Subfamily Cryptocephalinae Gyllenhal, 1813

2641Tribe Clytrini Kirby, 1837

2642Tribe Cryptocephalini Gyllenhal, 1813

2643Subtribe Achaenopina Chapuis, 1874

2644Subtribe Cryptocephalina Gyllenhal, 1813

2645Subtribe Monachulina Leng, 1920

2646Subtribe Pachybrachina Chapuis, 1874

2647Subtribe Stylosomina Chapuis, 1874

2648Tribe Fulcidacini Jakobson, 1924

2649Subfamily Eumolpinae Hope, 1840

2650Tribe Bromiini Baly, 1865 (1863)

2651Tribe Caryonodini Bechyné, 1951

2652Tribe Cubispini Monrós, 1954

2653Tribe Eumolpini Hope, 1840

2654Tribe Euryopini Chapuis, 1874

2655Tribe Habrophorini Bechyné and Špringlová de Bechyné, 1969

2656Tribe Hemydacnini Bechyné, 1951

2657Tribe Megascelidini Chapuis, 1874

2658Tribe Merodini Chapuis, 1874

2659Tribe Pygomolpini Bechyné, 1949

2660Tribe Rosiroiini Bechyné, 1950

2661Tribe Typophorini Baly, 1865

2662Subfamily Spilopyrinae Chapuis, 1874

2663Subfamily Synetinae LeConte and Horn, 1883

2664†Subfamily Protoscelidinae Medvedev, 1968

2665Superfamily Curculionoidea Latreille, 1802

2666Family Nemonychidae Bedel, 1882

2667Subfamily Nemonychinae Bedel, 1882

2668Subfamily Cimberidinae Gozis, 1882

2669Tribe Cimberidini Gozis, 1882

2670Tribe Doydirhynchini Pierce, 1916

2671†Tribe Kuschelomacrini Riedel, 2010

2672Subfamily Rhinorhynchinae Voss, 1922

2673Tribe Mecomacerini Kuschel, 1994

2674Subtribe Brarina Legalov, 2009

2675Subtribe Mecomacerina Kuschel, 1994

2676Tribe Rhinorhynchini Voss, 1922

2677†Subfamily Slonikinae Zherikhin, 1977

2678†Tribe Slonikini Zherikhin, 1977

2679†Tribe Ulyaniscini Legalov, 2009

2680†Subfamily Eccoptarthrinae Arnoldi, 1977

2681†Subfamily Brenthorrhininae Arnoldi, 1977

2682†Tribe Brenthorrhinini Arnoldi, 1977

2683†Tribe Brenthorrhinoidini Legalov, 2003

2684†Subfamily Distenorrhininae Arnoldi, 1977

2685†Subfamily Eobelinae Arnoldi, 1977

2686†Tribe Eobelini Arnoldi, 1977

2687†Subtribe Eobelina Arnoldi, 1977

2688†Subtribe Procurculionina Arnoldi, 1977

2689†Tribe Karataucarini Legalov, 2009

2690†Tribe Nanophydini Arnoldi, 1977

2691†Tribe Oxycorynoidini Arnoldi, 1977

2692†Tribe Probelini Legalov, 2009

2693†Subfamily Paleocartinae Legalov, 2003

2694†Tribe Nebrenthorrhinini Legalov, 2007

2695†Tribe Paleocartini Legalov, 2003

2696†Subfamily Metrioxenoidinae Legalov, 2009

2697†Subfamily Cretonemonychinae Gratshev and Legalov, 2009

2698†Subfamily Selengarhynchinae Gratshev and Legalov, 2009

2699Family Anthribidae Billberg, 1820

2700Subfamily Anthribinae Billberg, 1820

2701Tribe Anthribini Billberg, 1820

2702Tribe Basitropini Lacordaire, 1865

2703Tribe Corrhecerini Lacordaire, 1865

2704Tribe Cratoparini LeConte, 1876

2705†Tribe Cretanthribini Legalov, 2009

2706Tribe Decataphanini Lacordaire, 1865

2707Tribe Discotenini Lacordaire, 1865

2708Tribe Ecelonerini Lacordaire, 1865

2709Tribe Ischnocerini Lacordaire, 1865

2710Tribe Gymnognathini Valentine, 1960

2711Tribe Jordanthribini Morimoto, 1980

2712Tribe Mauiini Valentine, 1990

2713Tribe Mecocerini Lacordaire, 1865

2714Tribe Mycteini Morimoto, 1972

2715Tribe Ozotomerini Morimoto, 1972

2716Tribe Piesocorynini Valentine, 1960

2717Tribe Platyrhinini Bedel, 1882

2718Tribe Platystomini Pierce, 1916

2719Tribe Proscoporhinini Lacordaire, 1865

2720Tribe Ptychoderini Jekel, 1855

2721Tribe Sintorini Lacordaire, 1865

2722Tribe Stenocerini Kolbe, 1895

2723Tribe Tophoderini Lacordaire, 1865

2724Tribe Trigonorhinini Valentine, 1999

2725Tribe Tropiderini Lacordaire, 1865

2726Tribe Xenocerini Lacordaire, 1865

2727Tribe Xylinadini Lacordaire, 1865

2728Tribe Zygaenodini Lacordaire, 1865

2729Subfamily Choraginae Kirby, 1819

2730Tribe Apolectini Lacordaire, 1865

2731Tribe Araecerini Lacordaire, 1865

2732Tribe Cisanthribini Zimmerman, 1994

2733Tribe Choragini Kirby, 1819

2734Tribe Valenfriesiini Alonso-Zarazaga and Lyal, 1999

2735Tribe Xenorchestini Lacordaire, 1865

2736Subfamily Urodontinae Thomson, 1859

2737†Family Ulyanidae Zherikhin, 1993

2738Family Belidae Schönherr, 1826

2739Subfamily Belinae Schönherr, 1826

2740Tribe Agnesiotidini Zimmerman, 1994

2741Tribe Belini Schönherr, 1826

2742Subtribe Belina Schönherr, 1826

2743Subtribe Homalocerina Legalov, 2009

2744Tribe Pachyurini Kuschel, 1959

2745Subfamily Oxycoryninae Schönherr, 1840

2746Tribe Aglycyderini Wollaston, 1864

2747Tribe Alloxycorynini Legalov, 2009

2748Tribe Distenorrhinoidini Legalov, 2009

2749Tribe Metrioxenini Voss, 1953

2750Subtribe Afrocorynina Voss, 1957

2751Subtribe Metrioxenina Voss, 1953

2752Subtribe Zherichinixenina Legalov, 2009

2753Tribe Oxycorynini Schönherr, 1840

2754Subtribe Allocorynina Sharp, 1890

2755Subtribe Oxycorynina Schönherr, 1840

2756Subtribe Oxycraspedina Marvaldi and Oberprieler, 2006

2757Family Caridae Thompson, 1992

2758Subfamily Carinae Thompson, 1992

2759Subfamily Chilecarinae Legalov, 2009

2760Tribe Carodini Legalov, 2009

2761Tribe Chilecarini Legalov, 2009

2762†Subfamily Baissorhynchinae Zherikhin, 1993

2763Family Attelabidae Billberg, 1820

2764Subfamily Attelabinae Billberg, 1820

2765Tribe Attelabini Billberg, 1820

2766Subtribe Attelabina Billberg, 1820

2767Subtribe Euscelina Voss, 1925

2768Subtribe Euscelophilina Voss, 1925

2769Subtribe Henicolabina Legalov, 2007

2770Subtribe Himatolabina Legalov, 2003

2771Subtribe Hybolabina Voss, 1925

2772Subtribe Isolabina Legalov, 2007

2773Subtribe Lagenoderina Voss, 1925

2774Subtribe Lamprolabina Voss, 1925

2775Subtribe Metocalolabina Legalov, 2003

2776Subtribe Omolabina Legalov, 2003

2777Subtribe Paramecolabina Legalov, 2003

2778Subtribe Phialodina Legalov, 2003

2779Subtribe Phymatolabina Voss, 1925

2780Subtribe Phymatopsinina Legalov, 2003

2781Subtribe Pleurolabina Legalov, 2003

2782Tribe Euopini Voss, 1925

2783Tribe Pilolabini Voss, 1925

2784Subfamily Apoderinae Jekel, 1860

2785Tribe Apoderini Jekel, 1860

2786Tribe Clitostylini Voss, 1929

2787Subtribe Allapoderina Legalov, 2003

2788Subtribe Clitostylina Voss, 1929

2789Subtribe Pseudophrysina Legalov, 2003

2790Tribe Hoplapoderini Voss, 1926

2791Subtribe Afroapoderina Legalov, 2003

2792Subtribe Hoplapoderina Voss, 1926

2793Subtribe Paratomapoderina Legalov, 2003

2794Tribe Trachelophorini Voss, 1926

2795Subfamily Rhynchitinae Gistel, 1848

2796Tribe Auletini Desbrochers des Loges, 1908

2797Subtribe Auletina Desbrochers des Loges, 1908

2798Subtribe Auletobiina Legalov, 2001

2799Subtribe Guineauletina Legalov, 2003

2800Subtribe Mandelschtamiina Legalov, 2003

2801Subtribe Pseudauletina Voss, 1933

2802Subtribe Pseudomesauletina Legalov, 2003

2803Tribe Auletorhinini Voss, 1935

2804Tribe Byctiscini Voss, 1923

2805Subtribe Byctiscina Voss, 1923

2806Subtribe Listrobyctiscina Legalov, 2003

2807Subtribe Svetlanaebyctiscina Legalov, 2003

2808Tribe Cesauletini Legalov, 2003

2809Tribe Deporaini Voss, 1929

2810Subtribe Chonostropheina Morimoto, 1962

2811Subtribe Deporaina Voss, 1929

2812Tribe Minurini Legalov, 2003

2813Tribe Rhinocartini Voss, 1931

2814Tribe Rhynchitini Gistel, 1848

2815Subtribe Acritorrhynchitina Legalov, 2007

2816Subtribe Anisomerinina Legalov, 2003

2817Subtribe Eugnamptina Voss, 1930

2818Subtribe Lasiorhynchitina Legalov, 2003

2819Subtribe Perrhynchitina Legalov, 2003

2820Subtribe Rhynchitallina Legalov, 2003

2821Subtribe Rhynchitina Gistel, 1848

2822Subtribe Temnocerina Legalov, 2003

2823Subfamily Isotheinae Scudder, 1893

2824Tribe Isotheini Scudder, 1893

2825Subtribe Depasophilina Legalov, 2003

2826†Subtribe Isotheina Scudder, 1893

2827†Tribe Toxorhynchini Scudder, 1893

2828Subfamily Pterocolinae Lacordaire, 1865

2829Family Brentidae Billberg, 1820

2830Subfamily Brentinae Billberg, 1820

2831Tribe Brentini Billberg, 1820

2832Subtribe Arrhenodina Lacordaire, 1865

2833Subtribe Brentina Billberg, 1820

2834Subtribe Eremoxenina Semenov, 1892

2835Tribe Cyladini Schönherr, 1823

2836Tribe Cyphagogini Kolbe, 1892

2837Subtribe Atopobrentina Damoiseau, 1965

2838Subtribe Cyphagogina Kolbe, 1892

2839†Subtribe Dominibrentina Poinar, 2009

2840Subtribe Hoplopisthiina Senna and Calabresi, 1919

2841Subtribe Stereodermina Sharp, 1895

2842Tribe Pholidochlamydini Damoiseau, 1962

2843Tribe Taphroderini Lacordaire, 1865

2844Tribe Trachelizini Lacordaire, 1865

2845Subtribe Acratina Alonso-Zarazaga, Lyal, Bartolozzi and Sforzi, 1999

2846Subtribe Ithystenina Lacordaire, 1865

2847Subtribe Microtrachelizina Zimmerman, 1994

2848Subtribe Pseudoceocephalina Kleine, 1922

2849Subtribe Rhyticephalina Kleine, 1922

2850Subtribe Trachelizina Lacordaire, 1865

2851Subtribe Tychaeina Schoenfeldt, 1908

2852Tribe Ulocerini Schönherr, 1823

2853Subfamily Eurhynchinae Lacordaire, 1863

2854†Tribe Axelrodiellini Legalov, 2009

2855Tribe Eurhynchini Lacordaire, 1863

2856Subfamily Apioninae Schönherr, 1823

2857Supertribe Apionitae Schönherr, 1823

2858Tribe Apionini Schönherr, 1823

2859Subtribe Apionina Schönherr, 1823

2860Subtribe Aplemonina Kissinger, 1968

2861Subtribe Aspidapiina Alonso-Zarazaga, 1990

2862Subtribe Catapiina Alonso-Zarazaga, 1990

2863Subtribe Ceratapiina Alonso-Zarazaga, 1990

2864Subtribe Exapiina Alonso-Zarazaga, 1990

2865Subtribe Ixapiina Alonso-Zarazaga, 1990

2866Subtribe Kalcapiina Alonso-Zarazaga, 1990

2867Subtribe Malvapiina Alonso-Zarazaga, 1990

2868Subtribe Metapiina Alonso-Zarazaga, 1990

2869Subtribe Oxystomatina Alonso-Zarazaga, 1990

2870Subtribe Piezotrachelina Voss, 1959

2871Subtribe Prototrichapiina Wanat, 1995

2872Subtribe Synapiina Alonso-Zarazaga, 1990

2873Subtribe Trichapiina Alonso-Zarazaga, 1990

2874Tribe Chilapiini Wanat, 2001

2875Tribe Noterapiini Kissinger, 2004

2876Tribe Podapiini Wanat, 2001

2877Tribe Rhinorhynchidiini Zimmerman, 1994

2878Supertribe Antliarhinitae Schönherr, 1823

2879Supertribe Cybebitae Lacordaire, 1863

2880Supertribe Mecolenitae Wanat, 2001

2881Supertribe Myrmacicelitae Zimmerman, 1994

2882Tribe Lispotheriini Wanat, 2001

2883Tribe Myrmacicelini Zimmerman, 1994

2884Supertribe Rhadinocybitae Alonso-Zarazaga, 1992

2885Tribe Notapionini Zimmerman, 1994

2886Tribe Rhadinocybini Alonso-Zarazaga, 1992

2887Supertribe Tanaitae Schönherr, 1839

2888Subfamily Ithycerinae Schönherr, 1823

2889Subfamily Microcerinae Lacordaire, 1863

2890Subfamily Nanophyinae Gistel, 1848

2891Tribe Corimaliini Alonso-Zarazaga, 1989

2892Tribe Nanophyini Gistel, 1848

2893Family Dryophthoridae Schönherr, 1825

2894Subfamily Dryophthorinae Schönherr, 1825

2895Subfamily Cryptodermatinae Bovie, 1908

2896Subfamily Orthognathinae Lacordaire, 1865

2897Tribe Orthognathini Lacordaire, 1865

2898Tribe Rhinostomini LeConte, 1874

2899Subfamily Rhynchophorinae Schönherr, 1833

2900Tribe Diocalandrini Zimmerman, 1993

2901Tribe Litosomini Lacordaire, 1865

2902Tribe Ommatolampini Lacordaire, 1865

2903Tribe Polytini Zimmerman, 1993

2904Tribe Rhynchophorini Schönherr, 1833

2905Tribe Sphenophorini Lacordaire, 1865

2906Subfamily Stromboscerinae Lacordaire, 1865

2907Family Brachyceridae Billberg, 1820

2908Subfamily Brachycerinae Billberg, 1820

2909Tribe Brachycerini Billberg, 1820

2910Tribe Byrsopini Germar, 1829

2911Subfamily Cryptolarynginae Schalkwyk, 1966

2912Subfamily Erirhininae Schönherr, 1825

2913Tribe Aonychini Zimmerman, 1993

2914Tribe Arthrostenini Reitter, 1913

2915†Tribe Cretuliini Legalov, 2009

2916Tribe Erirhinini Schönherr, 1825

2917Tribe Himasthlophallini Zherikhin, 1991

2918Tribe Stenopelmini LeConte, 1876

2919Tribe Tadiini Zimmerman, 1993

2920Tribe Tanysphyrini Gistel, 1848

2921Subfamily Ocladiinae Lacordaire, 1865

2922Tribe Desmidophorini Morimoto, 1962

2923Tribe Ocladiini Lacordaire, 1865

2924Subfamily Raymondionyminae Reitter, 1913

2925Tribe Myrtonymini Kuschel, 1990

2926Tribe Raymondionymini Reitter, 1913

2927Family Curculionidae Latreille, 1802

2928Subfamily Curculioninae Latreille, 1802

2929Tribe Acalyptini Thomson, 1859

2930Subtribe Acalyptina Thomson, 1859

2931Subtribe Derelomina Lacordaire, 1865

2932Subtribe Notolomina Franz, 2006

2933Subtribe Phyllotrogina Franz, 2006

2934Subtribe Staminodeina Franz, 2006

2935Tribe Acentrusini Alonso-Zarazaga, 2005

2936Tribe Ancylocnemidini Voss, 1962

2937Tribe Anthonomini Thomson, 1859

2938Tribe Camarotini Schönherr, 1833

2939Subtribe Camarotina Schönherr, 1833

2940Subtribe Prionomerina Lacordaire, 1863

2941Tribe Ceratopodini Lacordaire, 1863

2942Tribe Cionini Schönherr, 1825

2943Tribe Cranopoeini Kuschel, 2009

2944Tribe Cryptoplini Lacordaire, 1863

2945Tribe Curculionini Latreille, 1802

2946Subtribe Curculionina Latreille, 1802

2947Subtribe Pseudobalaninina Heller, 1925

2948Subtribe Timolina Heller, 1925

2949Tribe Diabathrariini Lacordaire, 1863

2950Tribe Ellescini Thomson, 1859

2951Subtribe Dorytomina Bedel, 1886

2952Subtribe Ellescina Thomson, 1859

2953Tribe Erodiscini Lacordaire, 1863

2954Tribe Eugnomini Lacordaire, 1863

2955Subtribe Eugnomina Lacordaire, 1863

2956Subtribe Meriphina Marshall, 1937

2957Tribe Gonipterini Lacordaire, 1863

2958Tribe Mecinini Gistel, 1848

2959Tribe Nerthopini Lacordaire, 1865

2960Tribe Otidocephalini Lacordaire, 1863

2961Tribe Piazorhinini Lacordaire, 1863

2962Tribe Prionobrachiini Hustache, 1938

2963Tribe Pyropini Lacordaire, 1865

2964Tribe Rhamphini Rafinesque, 1815

2965Subtribe Dinorhopalina Voss, 1936

2966Subtribe Ixalmina Voss, 1936

2967Subtribe Rhamphina Rafinesque, 1815

2968Subtribe Tachygonina Lacordaire, 1865

2969Tribe Smicronychini Seidlitz, 1891 nomen protectum

2970Tribe Sphaeriopoeini Kuschel, 2003

2971Tribe Storeini Lacordaire, 1863

2972Tribe Styphlini Jekel, 1861

2973Tribe Tychiini Gistel, 1848

2974Subtribe Demimaeina Voss, 1937

2975Subtribe Lignyodina Bedel, 1883

2976Subtribe Ochyromerina Voss, 1935

2977Subtribe Tychiina Gistel, 1848

2978Tribe Ulomascini Lacordaire, 1865

2979Tribe Viticiini Morimoto, 1983

2980Subfamily Bagoinae Thomson, 1859 nomen protectum

2981Subfamily Baridinae Schönherr, 1836

2982Tribe Ambatini Lacordaire, 1863

2983Tribe Anopsilini Bondar, 1942

2984Tribe Apostasimerini Schönherr, 1844

2985Subtribe Apostasimerina Schönherr, 1844

2986Subtribe Madopterina Lacordaire, 1865

2987Subtribe Thaliabaridina Bondar, 1943

2988Subtribe Torcina Bondar, 1943

2989Subtribe Zygobaridina Pierce, 1907

2990Tribe Baridini Schönherr, 1836

2991Subtribe Baridina Schönherr, 1836

2992Subtribe Coelonertina Casey, 1922

2993Subtribe Coleomerina Casey, 1922

2994Subtribe Diorymerina Jekel, 1865

2995Subtribe Eurhinina Lacordaire, 1865

2996Tribe Madarini Jekel, 1865

2997Subtribe Barymerina Lacordaire, 1865

2998Subtribe Eutoxina Champion, 1908

2999Subtribe Leptoschoinina Lacordaire, 1865

3000Subtribe Madarina Jekel, 1865

3001Subtribe Tonesiina Alonso-Zarazaga and Lyal, 1999

3002Tribe Neosharpiini Hoffmann, 1956

3003Tribe Nertinini Voss, 1954

3004Tribe Optatini Champion, 1907

3005Tribe Pantotelini Lacordaire, 1865

3006Subtribe Cyrionychina Casey, 1922

3007Subtribe Pantotelina Lacordaire, 1865

3008Tribe Peridinetini Lacordaire, 1865

3009Subfamily Ceutorhynchinae Gistel, 1848

3010Tribe Ceutorhynchini Gistel, 1848

3011Tribe Cnemogonini Colonnelli, 1979

3012Tribe Egriini Pajni and Kohli, 1982

3013Tribe Hypohypurini Colonnelli, 2004

3014Tribe Hypurini Schultze, 1902

3015Tribe Lioxyonychini Colonnelli, 1984

3016Tribe Mecysmoderini Wagner, 1938

3017Tribe Mononychini LeConte, 1876

3018Tribe Phytobiini Gistel, 1848

3019Tribe Scleropterini Schultze, 1902

3020Subfamily Conoderinae Schönherr, 1833

3021Tribe Arachnopodini Lacordaire, 1865

3022Tribe Campyloscelini Schönherr, 1845

3023Subtribe Campyloscelina Schönherr, 1845

3024Subtribe Corynemerina Hustache, 1929

3025Subtribe Phaenomerina Faust, 1898

3026Tribe Conoderini Schönherr, 1833

3027Tribe Coryssomerini Thomson, 1859

3028Tribe Coryssopodini Lacordaire, 1865

3029Tribe Lechriopini Lacordaire, 1865

3030Tribe Lobotrachelini Lacordaire, 1865

3031Tribe Mecopini Lacordaire, 1865

3032Tribe Menemachini Lacordaire, 1865

3033Tribe Othippiini Morimoto, 1962

3034Tribe Peloropodini Hustache, 1932

3035Tribe Piazurini Lacordaire, 1865

3036Tribe Sphadasmini Lacordaire, 1865

3037Tribe Trichodocerini Champion, 1906

3038Tribe Zygopini Lacordaire, 1865

3039Subfamily Cossoninae Schönherr, 1825

3040Tribe Acamptini LeConte, 1876

3041Tribe Acanthinomerini Voss, 1972

3042Tribe Allomorphini Folwaczny, 1973

3043Tribe Aphyllurini Voss, 1955

3044Tribe Araucariini Kuschel, 1966

3045Tribe Choerorhinini Folwaczny, 1973

3046Tribe Cossonini Schönherr, 1825

3047Tribe Cryptommatini Voss, 1972

3048Tribe Dryotribini LeConte, 1876

3049Tribe Microxylobiini Voss, 1972

3050Tribe Nesiobiini Alonso-Zarazaga and Lyal, 1999

3051Tribe Neumatorini Folwaczny, 1973

3052Tribe Onychiini Chapuis, 1869

3053Tribe Onycholipini Wollaston, 1873

3054Tribe Pentarthrini Lacordaire, 1865

3055Tribe Proecini Voss, 1956

3056Tribe Pseudapotrepini Champion, 1909

3057Tribe Rhyncolini Gistel, 1848

3058Subtribe Phloeophagina Voss, 1955

3059Subtribe Pseudomimina Voss, 1939

3060Subtribe Rhyncolina Gistel, 1848

3061Tribe Tapiromimini Voss, 1972

3062Subfamily Cryptorhynchinae Schönherr, 1825

3063Tribe Aedemonini Faust, 1898

3064Tribe Camptorhinini Lacordaire, 1865

3065Tribe Cryptorhynchini Schönherr, 1825

3066Subtribe Cryptorhynchina Schönherr, 1825

3067Subtribe Mecistostylina Lacordaire, 1865

3068Subtribe Tylodina Lacordaire, 1865

3069Tribe Gasterocercini Zherikhin, 1991

3070Tribe Psepholacini Lacordaire, 1865

3071Tribe Sophrorhinini Lacordaire, 1865

3072Tribe Torneumatini Bedel, 1884

3073Subfamily Cyclominae Schönherr, 1826

3074Tribe Amycterini Waterhouse, 1854

3075Tribe Aterpini Lacordaire, 1863 nomen protectum

3076Subtribe Aterpina Lacordaire, 1863 nomen protectum

3077Subtribe Rhadinosomina Lacordaire, 1863

3078Tribe Cyclomini Schönherr, 1826

3079Tribe Dichotrachelini Hoffmann, 1957

3080Tribe Hipporhinini Lacordaire, 1863

3081Tribe Listroderini LeConte, 1876

3082Tribe Notiomimetini Wollaston, 1873

3083Tribe Rhythirrinini Lacordaire, 1863

3084Subfamily Entiminae Schönherr, 1823

3085Tribe Agraphini Horn, 1876

3086Tribe Alophini LeConte, 1874

3087Tribe Anomophthalmini Morrone, 1998

3088Tribe Anypotactini Champion, 1911

3089Tribe Blosyrini Lacordaire, 1863

3090Tribe Brachyderini Schönherr, 1826

3091Tribe Celeuthetini Lacordaire, 1863

3092Subtribe Celeuthetina Lacordaire, 1863

3093Subtribe Isopterina Morimoto and Kojima, 2001

3094Tribe Cneorhinini Lacordaire, 1863

3095Tribe Cratopodini Hustache, 1919

3096Tribe Cylydrorhinini Lacordaire, 1863

3097Tribe Cyphicerini Lacordaire, 1863

3098Subtribe Acanthotrachelina Marshall, 1944

3099Subtribe Cyphicerina Lacordaire, 1863

3100Subtribe Mylacorrhinina Reitter, 1913

3101Subtribe Myllocerina Pierce, 1913

3102Subtribe Phytoscaphina Lacordaire, 1863

3103Tribe Ectemnorhinini Lacordaire, 1863

3104Tribe Elytrurini Marshall, 1956

3105Tribe Embrithini Marshall, 1942

3106Tribe Entimini Schönherr, 1823

3107Tribe Episomini Lacordaire, 1863

3108Tribe Eudiagogini LeConte, 1874

3109Tribe Eupholini Günther, 1943

3110Tribe Eustylini Lacordaire, 1863

3111Tribe Geonemini Gistel, 1848

3112Tribe Holcorhinini Desbrochers des Loges, 1898

3113Tribe Hormorini Horn, 1876

3114Tribe Laparocerini Lacordaire, 1863

3115Tribe Leptostethini Lacordaire, 1863

3116Tribe Lordopini Schönherr, 1823

3117Tribe Mesostylini Reitter, 1913

3118Tribe Myorhinini Marseul, 1863

3119Tribe Nastini Reitter, 1913

3120Tribe Naupactini Gistel, 1848 nomen protectum

3121Tribe Nothognathini Marshall, 1916

3122Tribe Omiini Shuckard, 1839

3123Tribe Oosomini Lacordaire, 1863

3124Tribe Ophryastini Lacordaire, 1863

3125Tribe Ophtalmorrhynchini Hoffmann, 1965

3126Tribe Otiorhynchini Schönherr, 1826

3127Tribe Ottistirini Heller, 1925

3128Tribe Pachyrhynchini Schönherr, 1826

3129Tribe Peritelini Lacordaire, 1863

3130Tribe Phyllobiini Schönherr, 1826

3131Tribe Polycatini Marshall, 1956

3132Tribe Polydrusini Schönherr, 1823

3133Tribe Premnotrypini Kuschel, 1956

3134†Tribe Pristorhynchini Heer, 1847

3135Tribe Prypnini Lacordaire, 1863

3136Tribe Psallidiini Lacordaire, 1863

3137Tribe Rhyncogonini Sharp, 1919

3138Tribe Sciaphilini Sharp, 1891

3139Tribe Sitonini Gistel, 1848

3140Tribe Tanymecini Lacordaire, 1863

3141Subtribe Piazomiina Reitter, 1913

3142Subtribe Tainophthalmina Desbrochers des Loges, 1873

3143Subtribe Tanymecina Lacordaire, 1863

3144Tribe Tanyrhynchini Schönherr, 1826

3145Tribe Thecesternini Lacordaire, 1863

3146Tribe Trachyphloeini Gistel, 1848

3147Subtribe Trachyphilina Voss, 1948

3148Subtribe Trachyphloeina Gistel, 1848

3149Tribe Tropiphorini Marseul, 1863

3150Tribe Typhlorhinini Kuschel, 1954

3151Subfamily Hyperinae Marseul, 1863 (1848)

3152Tribe Cepurini Capiomont, 1867

3153Tribe Hyperini Marseul, 1863 (1848)

3154Subfamily Lixinae Schönherr, 1823

3155Tribe Cleonini Schönherr, 1826 nomen protectum

3156Tribe Lixini Schönherr, 1823

3157Tribe Rhinocyllini Lacordaire, 1863

3158Subfamily Mesoptiliinae Lacordaire, 1863

3159Tribe Carciliini Pierce, 1916

3160Tribe Laemosaccini Lacordaire, 1865

3161Tribe Magdalidini Pascoe, 1870 nomen protectum

3162Tribe Mesoptiliini Lacordaire, 1863

3163Subfamily Molytinae Schönherr, 1823

3164Tribe Anoplini Bedel, 1883

3165Tribe Amalactini Lacordaire, 1863

3166Tribe Aminyopini Voss, 1956

3167Tribe Amorphocerini Voss, 1939

3168Tribe Anchonini Imhoff, 1856

3169Tribe Brachyceropseini Aurivillius, 1926

3170Tribe Cholini Schönherr, 1825

3171Subtribe Cholina Schönherr, 1825

3172Subtribe Cholomina Vaurie, 1974

3173Subtribe Rhinastina Vaurie, 1973

3174Tribe Cleogonini Gistel, 1848

3175Tribe Conotrachelini Jekel, 1865

3176Tribe Cycloterini Lacordaire, 1863

3177Subtribe Cycloterina Lacordaire, 1863

3178Subtribe Thrombosternina Voss, 1965

3179Tribe Dinomorphini Lacordaire, 1863

3180Tribe Emphyastini Lacordaire, 1863

3181Tribe Euderini Lacordaire, 1865

3182Tribe Galloisiini Morimoto, 1962

3183Tribe Guioperini Lacordaire, 1865

3184Tribe Hylobiini Kirby, 1837

3185Subtribe Epistrophina Marshall, 1932

3186Subtribe Hylobiina Kirby, 1837

3187Tribe Ithyporini Lacordaire, 1865

3188Subtribe Colobodina Voss, 1958

3189Subtribe Ithyporina Lacordaire, 1865

3190Subtribe Sclerocardiina Lacordaire, 1865

3191Tribe Itini Reitter, 1913

3192Tribe Juanorhinini Aurivillius, 1931

3193Tribe Lepyrini Kirby, 1837

3194Tribe Lithinini Lacordaire, 1863

3195Subtribe Lithinina Lacordaire, 1863

3196Subtribe Rhytidophloeina Voss, 1963

3197Tribe Lymantini Lacordaire, 1865

3198Tribe Mecysolobini Reitter, 1913

3199Tribe Metatygini Pascoe, 1888

3200Tribe Molytini Schönherr, 1823

3201Subtribe Leiosomatina Reitter, 1913

3202Subtribe Molytina Schönherr, 1823

3203Subtribe Plinthina Lacordaire, 1863

3204Subtribe Typoderina Voss, 1965

3205Tribe Nettarhinini Lacordaire, 1865

3206Tribe Pacholenini Lacordaire, 1863

3207Tribe Paipalesomini Marshall, 1932

3208Tribe Petalochilini Lacordaire, 1863

3209Tribe Phoenicobatini Champion, 1914

3210Tribe Phrynixini Kuschel, 1964

3211Tribe Pissodini Gistel, 1848

3212Subtribe Cotasteromimina Morimoto, 1962

3213Subtribe Orthorhinina Jekel, 1865

3214Subtribe Pissodina Gistel, 1848

3215Tribe Sternechini Lacordaire, 1863

3216Tribe Styanacini Chûjô and Voss, 1960

3217Tribe Trachodini Gistel, 1848

3218Tribe Trigonocolini Lacordaire, 1863

3219Tribe Trypetidini Lacordaire, 1865

3220Subfamily Orobitidinae Thomson, 1859

3221Subfamily Xiphaspidinae Marshall, 1920

3222Subfamily Scolytinae Latreille, 1804

3223Tribe Amphiscolytini Mandelshtam and Beaver, 2003

3224Tribe Bothrosternini Blandford, 1896

3225Tribe Cactopinini Chamberlin, 1939

3226Tribe Carphodicticini Wood, 1971

3227Tribe Coptonotini Chapuis, 1869

3228Tribe Corthylini LeConte, 1876

3229Subtribe Corthylina LeConte, 1876

3230Subtribe Pityophthorina Eichhoff, 1878

3231Tribe Cryphalini Lindemann, 1877

3232Tribe Crypturgini LeConte, 1876

3233†Tribe Cylindrobrotini Kirejtshuk, Azar, Beaver, Mandelshtam and Nel, 2009

3234Tribe Diamerini Hagedorn, 1909

3235Tribe Dryocoetini Lindemann, 1877

3236Tribe Hexacolini Eichhoff, 1878

3237Tribe Hylastini LeConte, 1876

3238Tribe Hylesinini Erichson, 1836

3239Tribe Hylurgini Gistel, 1848

3240Tribe Hyorrhynchini Hopkins, 1915

3241Tribe Hypoborini Nüsslin, 1911

3242Tribe Ipini Bedel, 1888

3243Tribe Micracidini LeConte, 1876

3244Tribe Phloeosinini Nüsslin, 1912

3245Tribe Phloeotribini Chapuis, 1869

3246Tribe Phrixosomatini Wood, 1978

3247Tribe Polygraphini Chapuis, 1869

3248Tribe Premnobiini Browne, 1962

3249Tribe Scolytini Latreille, 1804

3250Tribe Scolytoplatypodini Blandford, 1893

3251Tribe Xyleborini LeConte, 1876

3252Tribe Xyloctonini Eichhoff, 1878

3253Tribe Xyloterini LeConte, 1876

3254Subfamily Platypodinae Shuckard, 1839

3255Tribe Mecopelmini Thompson, 1992

3256Tribe Platypodini Shuckard, 1839

3257Tribe Schedlariini Wood and Bright, 1992

3258Tribe Tesserocerini Strohmeyer, 1914

3259Subtribe Diapodina Strohmeyer, 1914

3260Subtribe Tesserocerina Strohmeyer, 1914

3261Rohdendorf, 1944

3262Tshekardocoleidae Rohdendorf, 1944: 252 [stem: Tshekardocole-]. Type genus: Tshekardocoleus Rohdendorf, 1944.

3263Rohdendorf, 1944

3264Tshekardocoleidae Rohdendorf, 1944: 252 [stem: Tshekardocole-]. Type genus: Tshekardocoleus Rohdendorf, 1944.

3265Uralocoleidae Zalesskiy, 1947: 857 [stem: Uralocole-]. Type genus: Uralocoleus Zalessky, 1947.

3266Ponomarenko, 1969

3267Labradorocoleidae Ponomarenko, 1969b: 307 [stem: Labradorocole-]. Type genus: Labradorocoleus Ponomarenko, 1969.

3268Kukalová, 1969

3269Oborocoleidae Kukalová, 1969: 155 [stem: Oborocole-]. Type genus: Oborocoleus Kukalová, 1969.

3270Martynov, 1933

3271Permocupidae Martynov, 1933: 85 [stem: Permocuped-]. Type genus: Permocupes Martynov, 1933.

3272Martynov, 1933

3273Permocupidae Martynov, 1933: 85 [stem: Permocuped-]. Type genus: Permocupes Martynov, 1933. Comment: incorrect original stem formation, not in prevailing usage.

3274Kaltanocoleidae Rohdendorf, 1961: 397 [stem: Kaltanocole-]. Type genus: Kaltanocoleus Rohdendorf, 1961.

3275Rohdendorf, 1961

3276Taldycupidae Rohdendorf, 1961: 412 [stem: Taldycuped-]. Type genus: Taldycupes Rohdendorf, 1961. Comment: usage of Trachycupidae by Fujiyama (1973: 375) was in error for Taldycupidae (Ponomarenko pers. comm. August 2009); incorrect original stem formation, not in prevailing usage.

3277Tillyard, 1924

3278Permosynidae Tillyard, 1924: 431 [stem: Permosyn-]. Type genus: Permosyne Tillyard, 1924.

3279Ponomarenko, 1968

3280Ademosynidae Ponomarenko, 1968: 128 [stem: Ademosyn-]. Type genus: Ademosyne Handlirsch, 1906.

3281Tillyard, 1924

3282Permosynidae Tillyard, 1924: 431 [stem: Permosyn-]. Type genus: Permosyne Tillyard, 1924.

3283Iablokoff-Khnzorian, 1983

3284Crowsoniellidae Iablokoff-Khnzorian, 1983: 65 [stem: Crowsoniell-]. Type genus: Crowsoniella Pace, 1975.

3285Laporte, 1836

3286Cupesidae Laporte, 1836: 56 [stem: Cuped-]. Type genus: Cupes Fabricius, 1801.

3287Crowson, 1962

3288Priacmini Crowson, 1962: 152 [stem: Priacm-]. Type genus: Priacma J. L. LeConte, 1874.

3289Ponomarenko, 1969

3290Mesocupedini Ponomarenko, 1969a: 105 [stem: Mesocuped-]. Type genus: Mesocupes Martynov, 1926.

3291Laporte, 1836

3292Cupesidae Laporte, 1836: 56 [stem: Cuped-]. Type genus: Cupes Fabricius, 1802. Comment: incorrect original stem formation, not in prevailing usage.

3293Barber, 1913

3294Micromalthidae Barber, 1913: 185 [stem: Micromalth-]. Type genus: Micromalthus J. L. LeConte, 1878.

3295Sharp and Muir, 1912

3296Ommadidae Sharp and Muir, 1912: 521 [stem: Ommat-]. Type genus: Omma Newman, 1839.

3297Hong, 1982

3298Brochocoleidae Hong, 1982: 100 [stem: Brochocole-]. Type genus: Brochocoleus Hong, 1982.

3299Crowson, 1962

3300Tetraphalerini Crowson, 1962: 152 [stem: Tetraphaler-]. Type genus: Tetraphalerus C. O. Waterhouse, 1901.

3301Sharp and Muir, 1912

3302Ommadidae Sharp and Muir, 1912: 521 [stem: Ommat-]. Type genus: Omma Newman, 1839.

3303Ponomarenko, 1969

3304Lithocupedini Ponomarenko, 1969a: 82 [stem: Lithocuped-]. Type genus: Lithocupes Ponomarenko, 1966.

3305Ponomarenko, 1966

3306Notocupedini Ponomarenko, 1966: 60 [stem: Notocup-]. Type genus: Notocupes Ponomarenko, 1966.

3307Sharp and Muir, 1912

3308Ommadidae Sharp and Muir, 1912: 521 [stem: Ommat-]. Type genus: Omma Newman, 1839. Comment: incorrect original stem formation, not in prevailing usage.

3309Ponomarenko, 1985

3310Jurodidae Ponomarenko, 1985: 53 [stem: Jurod-]. Type genus: Jurodes Ponomarenko, 1985.

3311Sikhotealiniidae Lafer, 1996: 390 [stem: Sikhotealini-]. Type genus: Sikhotealinia Lafer, 1996.

3312Ponomarenko, 1966

3313Triadocupedinae Ponomarenko, 1966: 48 [stem: Triadocuped-]. Type genus: Triadocupes Ponomarenko, 1966.

3314Hong, 1998

3315Magnocoleidae Hong, 1998: 41 [stem: Magnocole-]. Type genus: Magnocoleus Hong, 1998.

3316Zherikhin and Gratshev, 1994

3317Obrieniidae Zherikhin and Gratshev, 1994: 51 [stem: Obrieni-]. Type genus: Obrienia Zherikhin and Gratshev, 1994. Comment: precedence (Obrieniidae Zherikhin and Gratshev, 1994 vs Kararhynchidae Zherikhin and Gratshev, 1994) given to taxon originally proposed at the higher rank (Art. 24.1).

3318Zherikhin and Gratshev, 1994

3319Kararhynchinae Zherikhin and Gratshev, 1994: 58 [stem: Kararhynch-]. Type genus: Kararhynchus Zherikhin and Gratshev, 1994.

3320Zherikhin and Gratshev, 1994

3321Kararhynchinae Zherikhin and Gratshev, 1994: 58 [stem: Kararhynch-]. Type genus: Kararhynchus Zherikhin and Gratshev, 1994.

3322Legalov, 2009

3323Kenderlykanini Legalov, 2009c: 285 [stem: Kenderlyka-]. Type genus: Kenderlyka Zherikhin and Gratshev, 1994. Comment: incorrect original stem formation, not in prevailing usage.

3324Zherikhin and Gratshev, 1994

3325Obrieniidae Zherikhin and Gratshev, 1994: 51 [stem: Obrieni-]. Type genus: Obrienia Zherikhin and Gratshev, 1994.

3326Rohdendorf, 1961

3327Asiocoleidae Rohdendorf, 1961: 396 [stem: Asiocole-]. Type genus: Asiocoleus Rohdendorf, 1961.

3328Rohdendorf, 1961

3329Asiocoleidae Rohdendorf, 1961: 396 [stem: Asiocole-]. Type genus: Asiocoleus Rohdendorf, 1961.

3330Ponomarenko, 1969

3331Tricoleidae Ponomarenko, 1969a: 138 [stem: Tricole-]. Type genus: Tricoleus Ponomarenko, 1969.

3332Rohdendorf, 1961

3333Rhombocoleidae Rohdendorf, 1961: 432 [stem: Rhombocole-]. Type genus: Rhombocoleus Rohdendorf, 1961.

3334Rohdendorf, 1961

3335Rhombocoleidae Rohdendorf, 1961: 432 [stem: Rhombocole-]. Type genus: Rhombocoleus Rohdendorf, 1961.

3336Ponomarenko, 1968

3337Schizophoridae Ponomarenko, 1968: 130 [stem: Schizophor-]. Type genus: Schizophorus Ponomarenko, 1968. Comment: usage of this name conserved over Schizocoleoidea Rohdendorf, 1961 (Art. 35.5).

3338Ponomarenko, 1968

3339Schizophoridae Ponomarenko, 1968: 130 [stem: Schizophor-]. Type genus: Schizophorus Ponomarenko, 1968.

3340Ponomarenko, 1968

3341Catiniidae Ponomarenko, 1968: 137 [stem: Catini-]. Type genus: Catinius Ponomarenko, 1968.

3342Rohdendorf, 1961

3343Curculiopsidae Martynov, 1937: 39 [stem: Curculiopse-]. Type genus: Curculiopsis Martynov, 1937 [preoccupied genus name, not Curculiopsis Handlirsch, 1907 [fossil Curculionoidea]; syn. of Aenigmocoleus Rohdendorf, 1961 or Rossocoleus Rohdendorf, 1961]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.

3344Schizocoleidae Rohdendorf, 1961: 438 [stem: Schizocole-]. Type genus: Schizocoleus Rohdendorf, 1961.

3345Hinton, 1936 (1882)

3346Lepiceridae Hinton, 1936: 473 [stem: Lepicer-]. Type genus: Lepicerus Motschulsky, 1855. Comment: use of family-group name conserved over Cyathoceroidea Sharp, 1882 (Art. 40.2) (see Lawrence and Newton 1995: 804).

3347Hinton, 1936 (1882)

3348Cyathoceridae Sharp, 1882: 141 [stem: Cyathocer-]. Type genus: Cyathocerus Sharp, 1882 [syn. of Lepicerus Motschulsky, 1855]. Comment: use of younger name Lepiceridae Hinton, 1936 conserved over this name (Art. 40.2) (see Lawrence and Newton 1995: 804).

3349Lepiceridae Hinton, 1936: 473 [stem: Lepicer-]. Type genus: Lepicerus Motschulsky, 1855. Comment: use of family-group name conserved over Cyathoceridae Sharp, 1882 (Art. 40.2) (see Lawrence and Newton 1995: 804).

3350†Haplochelidae Kirejtshuk and Poinar, 2006: 156 [stem: Haplochel-]. Type genus: Haplochelus Kirejtshuk and Poinar, 2006. Comment: synonymy with Lepiceridae by Ge et al. (2010: 336).

3351Erichson, 1845

3352Sphaerina Erichson, 1845: 38 [stem: Sphaerius-]. Type genus: Sphaerius Waltl, 1838 [originally placed on the Official Index of Rejected and Invalid Generic Names in Zoology by the Commission (ICZN 1985d) but later placed on the Official List of Generic Names in Zoology (ICZN 2000)]. Comment: correct stem determined to be Sphaerius- and Sphaeriusidae Erichson, 1845 placed on the Official List of Family-Group Names in Zoology (ICZN 2000).

3353Steffan, 1964

3354Torridincolidae Steffan, 1964: 199 [stem: Torridincol-]. Type genus: Torridincola Steffan, 1964.

3355Steffan, 1964

3356Torridincolidae Steffan, 1964: 199 [stem: Torridincol-]. Type genus: Torridincola Steffan, 1964.

3357Ptyopterinae M. Abdullah, 1974: 961 [stem: Ptyopteryg-]. Type genus: Ptyopteryx H. Reichardt and C. Costa, 1967 [preoccupied genus name, not Ptyopteryx Kolenati, 1848 [Trichoptera]; syn. of Iapir Py-Daniel et al., 1993]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.

3358Endrödy-Younga, 1997

3359Deleveinae Endrödy-Younga, 1997: 317 [stem: Deleve-]. Type genus: Delevea H. Reichardt, 1976.

3360LeConte, 1874

3361Hydroscaphidae J. L. LeConte, 1874a: 45 [stem: Hydroscaph-]. Type genus: Hydroscapha J. L. LeConte, 1874.

3362Erichson, 1845

3363Sphaerina Erichson, 1845: 38 [stem: Sphaerius-]. Type genus: Sphaerius Waltl, 1838 [originally placed on the Official Index of Rejected and Invalid Generic Names in Zoology by the Commission (ICZN 1985d) but later placed on the Official List of Generic Names in Zoology (ICZN 2000)]. Comment: family-group name and its type genus were rejected by the Commission (ICZN 1985d) but rescinded afterwards (ICZN 2000); family-group name placed on the Official List of Family-Group Names in Zoology and correct stem ruled to be Sphaerius- (ICZN 2000).

3364Microsporidae Crotch, 1873a: 78 [stem: Microspor-]. Type genus: Microsporus Kolenati, 1846 [placed on the Official List of Generic Names in Zoology (ICZN 1985d)]. Comment: name originally placed on the Official List of Family-Group Names in Zoology as “Microsporidae H. Reichardt, 1976” (ICZN 1985d) but author and year subsequently corrected to Crotch, 1873 (ICZN 2000).

3365Hong, 2002

3366Tritarsusidae Hong, 2002: 102 [stem: Tritars-]. Type genus: Tritarsus Hong, 2002. Comment: incorrect original stem formation, not in prevailing usage.

3367Latreille, 1810

3368Gyrinites Latreille, 1810: 141 [stem: Gyrin-]. Type genus: Gyrinus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].

3369Folkerts, 1979

3370Spanglerogyrinae Folkerts, 1979: 7 [stem: Spanglerogyr-]. Type genus: Spanglerogyrus Folkerts, 1979.

3371Latreille, 1810

3372Gyrinites Latreille, 1810: 141 [stem: Gyrin-]. Type genus: Gyrinus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].

3373Régimbart, 1882

3374Enhydrini Régimbart, 1882: 392 [stem: Enhydr-]. Type genus: Enhydrus Laporte, 1834 [placed on the Official List of Generic Names in Zoology (ICZN 1964)]. Comment: usage of this name conserved over Dineutini Desmarest, 1851 (Art. 35.5).

3375Desmarest, 1851

3376Dineutides Desmarest, 1851: 223 [stem: Dineut-]. Type genus: Dineutes W. S. MacLeay, 1825.

3377Régimbart, 1882

3378Enhydrini Régimbart, 1882: 392 [stem: Enhydr-]. Type genus: Enhydrus Laporte, 1834 [placed on the Official List of Generic Names in Zoology (ICZN 1964)]. Comment: junior homonym of Enhydrini Gray, 1825 (type genus Enhydra Fleming, 1822) in Mammalia; an application was recently submitted to the Commission by Özdikmen and Darilmaz (2010; see Appendix 6) to remove the homonymy (Art. 55.3.1).

3379Latreille, 1810

3380Gyrinites Latreille, 1810: 141 [stem: Gyrin-]. Type genus: Gyrinus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].

3381Latreille, 1810

3382Gyrinites Latreille, 1810: 141 [stem: Gyrin-]. Type genus: Gyrinus Geoffroy, 1762 [placed on the Official List of Generic Names in Zoology (ICZN 1994a)].

3383Brinck, 1956

3384Heterogyrini P. Brinck, 1956: 37, in key [stem: Heterogyr-]. Type genus: Heterogyrus Legros, 1953.

3385Régimbart, 1882

3386Orectochilini Régimbart, 1882: 391 [stem: Orectochil-]. Type genus: Orectochilus Dejean, 1833.

3387Thomson, 1857

3388Trachypachini C. G. Thomson, 1857: 5 [stem: Trachypach-]. Type genus: Trachypachus Motschulsky, 1844.

3389Ponomarenko, 1977

3390Eodromeinae Ponomarenko, 1977: 46 [stem: Eodrome-]. Type genus: Eodromeus Ponomarenko, 1977. Comment: the type genus originally included four species: antiquus, stenalis, dissectus and major; Ponomarenko (1977: 66) designated “fasciatus” as the type species of the genus which was a lapsuscalami for dissectus (see Carpenter 1992: 291); this genus is considered here as being available in Ponomarenko (1977: 66) and not Ponomarenko (in Carpenter 1992: 291); this action enables us to treat the family-group name Eodromeinae as available from the original description in Ponomarenko (1977: 46).

3391Leptopodocoleidae Hong, 1982: 118 [stem: Leptopodocole-]. Type genus: Leptopodocoleus Hong, 1982.

3392Thomson, 1857

3393Trachypachini C. G. Thomson, 1857: 5 [stem: Trachypach-]. Type genus: Trachypachus Motschulsky, 1844.

3394*Systolosomini Erwin, 1985: 467 [stem: Systolosomat-]. Type genus: Systolosoma Solier, 1849. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1); incorrect original stem formation, not in prevailing usage.

3395Systolosomini Erwin, 1991: 4, in key [stem: Systolosomat-]. Type genus: Systolosoma Solier, 1849. Comment: incorrect original stem formation, not in prevailing usage.

3396Laporte, 1840

3397Rhysodites Laporte, 1840a: 291 [stem: Rhysod-]. Type genus: Rhysodes Germar, 1822.

3398Bell and Bell, 1978

3399Leoglymmiina R. T. Bell and J. R. Bell, 1978: 53 [stem: Leoglymmi-]. Type genus: Leoglymmius Bell and Bell, 1978.

3400Bell and Bell, 1978

3401Dhysorina R. T. Bell and J. R. Bell, 1978: 53 [stem: Dhysor-]. Type genus: Dhysores Grouvelle, 1903.

3402Bell and Bell, 1987

3403Medisorina R. T. Bell and J. R. Bell, 1987: 287 [stem: Medisor-]. Type genus: Medisores Bell and Bell, 1987.

3404Laporte, 1840

3405Rhysodites Laporte, 1840a: 291 [stem: Rhysod-]. Type genus: Rhysodes Germar, 1822. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Agassiz (1846b: 326, as Rhyssodeoidae [incorrect stem formation]), generally accepted as in R. T. Bell (2003: 78, as Rhysodidae).

3406Bell and Bell, 1978

3407Clinidiina R. T. Bell and J. R. Bell, 1978: 59 [stem: Clinidi-]. Type genus: Clinidium Kirby, 1830.

3408Bell and Bell, 1978

3409Omoglymmiina R. T. Bell and J. R. Bell, 1978: 66 [stem: Omoglymmi-]. Type genus: Omoglymmius Ganglbauer, 1891.

3410Bell and Bell, 1991

3411Sloanoglymmiina R. T. Bell and J. R. Bell, 1991: 183 [stem: Sloanoglymmi-]. Type genus: Sloanoglymmius Bell and Bell, 1991.

3412Latreille, 1802

3413Carabici Latreille, 1802: 80 [stem: Carab-]. Type genus: Carabus Linnaeus, 1758 [placed on the Official List of Generic Names in Zoology (ICZN 1950)]. Comment: First Reviser (Carabidae Latreille, 1802 vs Cicindelidae Latreille, 1802 vs Elaphridae Latreille, 1802) not determined, current usage maintained.

3414Ponomarenko, 1977

3415Protorabinae Ponomarenko, 1977: 71 [stem: Protorab-]. Type genus: Protorabus Ponomarenko, 1977.

3416Ponomarenko, 1977

3417Conjunctiini Ponomarenko, 1977: 85 [stem: Conjuncti-]. Type genus: Conjunctia Ponomarenko, 1977.

3418Laporte, 1834

3419Nebriidae Laporte, 1834b: 90 [stem: Nebri-]. Type genus: Nebria Latreille, 1802.

3420Laporte, 1834

3421Nebriidae Laporte, 1834b: 90 [stem: Nebri-]. Type genus: Nebria Latreille, 1802.

3422Kavanaugh and Nègre, 1983

3423*Notiokasiini Erwin, 1979: 578 [stem: Notiokasi-]. Type genus: Notiokasis Kavanaugh and Nègre, 1983. Comment: family-group name unavailable (Art. 11.7.1.1): not based on an available genus name at the time.

3424Notiokasiini Kavanaugh and Nègre, 1983: 551 [stem: Notiokasi-]. Type genus: Notiokasis Kavanaugh and Nègre, 1983.

3425Motschulsky, 1850

3426*Notiophiles Motschulsky, 1849: 54 [stem: Notiophil-]. Type genus: Notiophilus Duméril, 1805. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Motschulsky (1849).

3427Notiophili Motschulsky, 1850: 16 [stem: Notiophil-]. Type genus: Notiophilus Duméril, 1805.

3428Dupuis, 1912

3429Opisthiinae Dupuis, 1912: 1 [stem: Opisthi-]. Type genus: Opisthius Kirby, 1837.

3430Kavanaugh, 1996

3431Pelophilini Kavanaugh, 1996: 35 [stem: Pelophil-]. Type genus: Pelophila Dejean, 1821.

3432Csiki, 1927

3433Cicindini Csiki, 1927: 425 [stem: Cicind-]. Type genus: Cicindis Bruch, 1908.

3434Latreille, 1802

3435Cicindeletae Latreille, 1802: 77 [stem: Cicindel-]. Type genus: Cicindela Linné, 1758.

3436Csiki, 1903

3437Amblychilinae Csiki, 1903: 124 [stem: Amblycheil-]. Type genus: Amblycheila Say, 1830 [as Amblychila, unjustified emendation of type genus name by Agassiz (1846b), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.

3438Omites W. Horn, 1907: 466 [stem: Om-]. Type genus: Omus Eschscholtz, 1829.

3439Latreille, 1802

3440Cicindeletae Latreille, 1802: 77 [stem: Cicindel-]. Type genus: Cicindela Linnaeus, 1758.

3441Rivalier, 1971

3442Apteroessina Rivalier, 1971: 143 [stem: Apteroess-]. Type genus: Apteroessa Hope, 1838.

3443Latreille, 1802

3444Cicindeletae Latreille, 1802: 77 [stem: Cicindel-]. Type genus: Cicindela Linnaeus, 1758.

3445Thomson, 1859

3446Dromicitae J. Thomson, 1859: 89 [stem: Dromic-]. Type genus: Dromica Dejean, 1826.

3447Caledonicini W. Horn, 1893: 324 [stem: Caledonic-]. Type genus: Caledonica Chaudoir, 1861.

3448Odontochilini W. Horn, 1899: 41 [stem: Odontocheil-]. Type genus: Odontocheila Laporte, 1834 [as Odontochila, unjustified emendation of type genus name by Agassiz (1846b), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.

3449Prepusini W. Horn, 1899: 44 [stem: Prepus-]. Type genus: Prepusa Chaudoir, 1850 [syn. of Eulampra Chaudoir, 1848].

3450Euryodini W. Horn, 1899: 37 [stem: Euryod-]. Type genus: Euryoda Lacordaire, 1842 [syn. of Prothyma Hope, 1838].

3451Prothymini W. Horn, 1906: 86 [stem: Prothym-]. Type genus: Prothyma Hope, 1838.

3452Rivalier, 1971

3453Euprosopini W. Horn, 1893: 324 [stem: Euprosop-]. Type genus: Euprosopus Dejean, 1825.

3454Eucalliini W. Horn, 1893: 324 [stem: Eucalli-]. Type genus: Eucallia Guérin-Méneville, 1844 [syn. of Callidema Guérin-Méneville, 1843].

3455Distypsiderini W. Horn, 1893: 324 [stem: Distipsider-]. Type genus: Distipsidera Westwood, 1837 [as Distypsidera, unjustified emendation of type genus name by Gemminger and Harold (1868a: 32), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.

3456Iresina Rivalier, 1971: 138 [stem: Iresi-]. Type genus: Iresia Dejean, 1829. Comment: although this is not the oldest name for the subtribe, we recommend that an application be sent to the Commission in order to conserve usage of Iresiina Rivalier, 1971 over the three older names proposed by W. Horn in 1893 which have not been used as valid recently to our knowledge; incorrect original stem formation, not in prevailing usage.

3457Horn, 1893

3458Theratidae W. Horn, 1893: 324 [stem: Therat-]. Type genus: Therates Latreille, 1816.

3459Brullé, 1834

3460Collyriens Brullé, 1834: 96 [stem: Collyrid-]. Type genus: Collyris Fabricius, 1801.

3461Brullé, 1834

3462Collyriens Brullé, 1834: 96 [stem: Collyrid-]. Type genus: Collyris Fabricius, 1801. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Hope (1838a: 30, as Collyridae [incorrect stem formation]), generally accepted as in Puchkov and Matalin (2003: 116, as Collyridina); incorrect original stem formation, not in prevailing usage.

3463Tracheloniadae Gistel, 1850: 75 [stem: Tracheloni-]. Type genus: Trachelonia Gistel, 1850 [syn. of Collyris Fabricius, 1801].

3464*Colliurides Motschulsky, 1855: 34 [stem: Colliur-]. Type genus: Colliuris sensu Latreille, 1802 [not Colliuris DeGeer, 1774; syn. of Collyris Fabricius, 1801]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized.

3465Naviaux, 1991

3466Tricondylina Naviaux, 1991: 219 [stem: Tricondyl-]. Type genus: Tricondyla Latreille, 1822.

3467Laporte, 1834

3468Ctenostomidae Laporte, 1834b: 38 [stem: Ctenostomat-]. Type genus: Ctenostoma Klug, 1821. Comment: incorrect original stem formation, not in prevailing usage.

3469Laporte, 1834

3470Manticoridae Laporte, 1834b: 33 [stem: Manticor-]. Type genus: Manticora Fabricius, 1781.

3471Laporte, 1834

3472Megacephalidae Laporte, 1834b: 33 [stem: Megacephal-]. Type genus: Megacephala Latreille, 1802.

3473Oxycheilites J. Thomson, 1857a: 17 [stem: Oxycheil-]. Type genus: Oxycheila Dejean, 1825. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Chaudoir (1861a: 326, as Oxychilini [incorrect stem formation]), generally accepted as in Fleutiaux (1892a: 13, as Oxychilini [incorrect stem formation]); the stem Oxychil- should not be used for this taxon in order to avoid homonymy with Oxychilini Hesse, 1927 (type genus Oxychilus Fitzinger 1833), currently being used as valid in Mollusca.

3474Platychilidae W. Horn, 1893: 325 [stem: Platychil-]. Type genus: Platychile W. S. MacLeay, 1825.

3475Tetrachae Leng and Mutchler, 1916: 683 [stem: Tetrach-]. Type genus: Tetracha Hope, 1838.

3476Latreille, 1802

3477Carabici Latreille, 1802: 80 [stem: Carab-]. Type genus: Carabus Linnaeus, 1758.

3478Latreille, 1802

3479Carabici Latreille, 1802: 80 [stem: Carab-]. Type genus: Carabus Linnaeus, 1758.

3480Calosomii Bonelli, 1810: Tabula Synoptica [stem: Calosomat-]. Type genus: Calosoma Weber, 1801. Comment: incorrect original stem formation, not in prevailing usage.

3481Proceridae Laporte, 1834b: 87 [stem: Procer-]. Type genus: Procerus Dejean, 1821.

3482Callisthenisidae Gistel, 1848: [2] [stem: Callisthen-]. Type genus: Callisthenes Fischer von Waldheim, 1820. Comment: incorrect original stem formation, not in prevailing usage.

3483*Cechenogenici Morawitz, 1889: 40 [stem: Cechen-]. Type genus: Cechenus Fischer von Waldheim, 1822. Comment: family-group name unavailable (Art. 11.7.1.1): original name not proposed as a noun.

3484*Tribacogenici Morawitz, 1889: 40 [stem: Tribac-]. Type genus: Tribax Fischer von Waldheim, 1817. Comment: family-group name unavailable (Art. 11.7.1.1): original name not proposed as a noun.

3485*Procrustogenici Morawitz, 1889: 40 [stem: Procrust-]. Type genus: Procrustes Bonelli, 1810. Comment: family-group name unavailable (Art. 11.7.1.1): original name not proposed as a noun.

3486Procrustogenici Roeschke, 1898: 285 [stem: Procrust-]. Type genus: Procrustes Bonelli, 1810.

3487Aplothoracina Lapouge, 1927: 45 [stem: Aplothorac-]. Type genus: Aplothorax G. R. Waterhouse, 1842.

3488*Callistheniens Lapouge, 1927: 47 [stem: Callisthen-]. Type genus: Callisthenes Fischer von Waldheim, 1820. Comment: original vernacular name unavailable (Art. 11.7.2): proposed after 1899; family-group name proposed as new without reference to Callisthenisidae Gistel, 1848.

3489Cechenogenici Csiki, 1927: 110 [stem: Cechen-]. Type genus: Cechenus Fischer von Waldheim, 1822.

3490Tribacogenici Csiki, 1927: 121 [stem: Tribac-]. Type genus: Tribax Fischer von Waldheim, 1817.

3491Megadontici Csiki, 1927: 60 [stem: Megodont-]. Type genus: Megodontus Solier, 1848 [as Megadontus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.

3492Lapouge, 1927

3493Ceroglossina Lapouge, 1927: 45 [stem: Cerogloss-]. Type genus: Ceroglossus Solier, 1848.

3494Perty, 1830

3495Cychrii Perty, 1830: 6 [stem: Cychr-]. Type genus: Cychrus Fabricius, 1794. Comment: name previously attributed to Laporte (1834b: 86).

3496Hope, 1838

3497Pamboridae Hope, 1838a: 47 [stem: Pambor-]. Type genus: Pamborus Latreille, 1812.

3498Bonelli, 1810

3499Loricerides Bonelli, 1810: Tabula Synoptica [stem: Loricer-]. Type genus: Loricera Latreille, 1802.

3500Bonelli, 1810

3501Omophronii Bonelli, 1810: Tabula Synoptica [stem: Omophron-]. Type genus: Omophron Latreille, 1802.

3502Scolyti Motschulsky, 1850: 91 [stem: Scolyt-]. Type genus: Scolytus Fabricius, 1790 [preoccupied genus name, not Scolytus Geoffroy, 1762 [Coleoptera: Curculionidae]; syn. of Omophron Latreille, 1802]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; Scolytinae/-ini Latreille, 1804 are currently used as valid in Curculionidae.

3503Epactiini Fauvel, 1888: 1 [stem: Epacti-]. Type genus: Epactius Schneider, 1791 [syn. of Omophron Latreille, 1802].

3504Latreille, 1802

3505Elaphrii Latreille, 1802: 81 [stem: Elaphr-]. Type genus: Elaphrus Fabricius, 1775.

3506Chaudoir, 1861

3507Migadopidae Chaudoir, 1861b: 510 [stem: Migadop-]. Type genus: Migadops G. R. Waterhouse, 1842.

3508Erwin, 1985

3509Amarotypini Erwin, 1985: 468 [stem: Amarotyp-]. Type genus: Amarotypus H. W. Bates, 1872.

3510Chaudoir, 1861

3511Migadopidae Chaudoir, 1861b: 510 [stem: Migadop-]. Type genus: Migadops G. R. Waterhouse, 1842.

3512Moret, 2005

3513Aquilicina Moret, 2005: 30 [stem: Aquilic-]. Type genus: Aquilex Moret, 1989.

3514Chaudoir, 1861

3515Migadopidae Chaudoir, 1861b: 510 [stem: Migadop-]. Type genus: Migadops G. R. Waterhouse, 1842.

3516Monolobinae Jeannel, 1938: 10, in key [stem: Monolob-]. Type genus: Monolobus Solier, 1849.

3517Loxomeriformes Erwin, 1985: 446 [stem: Loxomer-]. Type genus: Loxomerus Chaudoir, 1842.

3518Schiødte, 1848

3519Hiletini Schiødte, 1848: 69 [stem: Hilet-]. Type genus: Hiletus Schiødte, 1848.

3520Camaragnathini Csiki, 1927: 341 [stem: Camaragnath-]. Type genus: Camaragnathus Bertrand-Bocandé, 1849 [syn. of Hiletus Schiødte, 1847].

3521Bonelli, 1810

3522Scaritides Bonelli, 1810: Tabula Synoptica [stem: Scarit-]. Type genus: Scarites Fabricius, 1775.

3523MacLeay, 1887

3524Carenides W. J. MacLeay, 1887: 117 [stem: Caren-]. Type genus: Carenum Bonelli, 1813.

3525Rafinesque, 1815

3526Clivinidia Rafinesque, 1815: 109 [stem: Clivin-]. Type genus: Clivina Latreille, 1802.

3527Putzeys, 1867

3528Ardistomides Putzeys, 1867: 200 [stem: Ardistom-]. Type genus: Ardistomis Putzeys, 1846. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Csiki (1927: 547, as Ardistomina), generally accepted as in Balkenohl (2003: 219, as Ardistomina); current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Ardistomid-).

3529Rafinesque, 1815

3530Clivinidia Rafinesque, 1815: 109 [stem: Clivin-]. Type genus: Clivina Latreille, 1802.

3531Reicheina Jeannel, 1957: 141, in key [stem: Reichei-]. Type genus: Reicheia Saulcy, 1862. Comment: incorrect original stem formation, not in prevailing usage.

3532Italodytina Jeannel, 1957: 141, in key [stem: Italodyt-]. Type genus: Italodytes Müller, 1938.

3533Reicheiina Basilewsky, 1980: 293 [stem: Reichei-]. Type genus: Reicheia Saulcy, 1862. Comment: family-group name proposed as new without reference to Reicheiina Jeannel, 1957.

3534Bänninger, 1938

3535*Oxystomides Putzeys, 1867: 12 [stem: Oxystom-]. Type genus: Oxystomus Dejean, 1825 [preoccupied genus name, not Oxystomus Fischer von Waldheim, 1803 [Mammalia]; syn. of Forcipator Maindron, 1904]. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Putzeys (1867).

3536Oxystomina Csiki, 1927: 491. Type genus: Oxystomus Dejean, 1825 [preoccupied genus name, not Oxystomus Fischer von Waldheim, 1803 [Mammalia]; syn. of Forcipator Maindron, 1904]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.

3537Forcipatorina Bänninger, 1938: 83 [stem: Forcipator-]. Type genus: Forcipator Maindron, 1904.

3538Mateu, 2002

3539Dalyatinae Mateu, 2002: 67 [stem: Dalyat-]. Type genus: Dalyat Mateu, 2002.

3540Kolbe, 1880

3541Dyschiriini Kolbe, 1880: 266 [stem: Dyschiri-]. Type genus: Dyschirius Bonelli, 1810.

3542McKay, 1991

3543Palaeoaxinidini McKay, 1991: 10 [stem: Palaeoaxinidi-]. Type genus: Palaeoaxinidium McKay, 1991. Comment: incorrect original stem formation, not in prevailing usage.

3544Putzeys, 1867

3545Pasimachides Putzeys, 1867: 3 [stem: Pasimach-]. Type genus: Pasimachus Bonelli, 1813. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form and generally accepted as in Csiki (1927: 444, as Pasimachina).

3546LeConte, 1853

3547Promecognathi J. L. LeConte, 1853b: 394 [stem: Promecognath-]. Type genus: Promecognathus Chaudoir, 1846.

3548Axinidiini Basilewsky, 1963: 307 [stem: Axinidi-]. Type genus: Axinidium Sturm, 1843.

3549Alluaud, 1930

3550Salcediini Alluaud, 1930a: 21 [stem: Salcedi-]. Type genus: Salcedia Fairmaire, 1899.

3551Bell, 1998

3552Androzelmina R. T. Bell, 1998: 271 [stem: Androzelm-]. Type genus: Androzelma Dostal, 1993.

3553Alluaud, 1930 (1929)

3554Zelmides Andrewes, 1929: 209, 416 [stem: Zelm-]. Type genus: Zelma Andrewes, 1920 [syn. of Salcedia Fairmaire, 1899]. Comment: use of younger name Salcediina Alluaud, 1930 conserved over this name (Art. 40.2).

3555Salcediini Alluaud, 1930a: 21 [stem: Salcedi-]. Type genus: Salcedia Fairmaire, 1899. Comment: name proposed to replace Zelmini Andrewes, 1929 because of the synonymy of the type genus; use of Salcediina conserved over Zelmina Andrewes, 1929 (Art. 40.2).

3556Bell, 1998

3557Solenogenyina R. T. Bell, 1998: 271 [stem: Solenogeny-]. Type genus: Solenogenys Westwood, 1859.

3558Bonelli, 1810

3559Scaritides Bonelli, 1810: Tabula Synoptica [stem: Scarit-]. Type genus: Scarites Fabricius, 1775.

3560Csiki, 1927

3561Acanthoscelina Csiki, 1927: 489 [stem: Acanthoscel-]. Type genus: Acanthoscelis Dejean, 1825. Comment: the incorrect stem formation is in prevailing usage (should be Acanthoscelid-) and is maintained (Art. 29.3.1.1); usage of the stem Acanthoscel- avoids homonymy with Acanthoscelidina Bridwell, 1946 (type genus Acanthoscelides Schilsky, 1905) in Chrysomelidae.

3562Basilewsky, 1973

3563Corintascarini Basilewsky, 1973a: 10 [stem: Corintascar-]. Type genus: Corintascaris Basilewsky, 1952. Comment: current spelling maintained (Art. 29.3.1.1): incorrect stem formation in prevailing usage (should be Corintascarid-).

3564Basilewsky, 1973

3565Dyscherina Basilewsky, 1973a: 101 [stem: Dyscher-]. Type genus: Dyscherus Chaudoir, 1855.

3566Basilewsky, 1973

3567Ochyropini Basilewsky, 1973a: 9 [stem: Ochyrop-]. Type genus: Ochyropus Schiødte, 1847. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Ochyropod-).

3568Andrewes, 1929

3569Oxylobides Andrewes, 1929: 209, 292 [stem: Oxylob-]. Type genus: Oxylobus Chaudoir, 1855.

3570Putzeys, 1867

3571Scaptérides Putzeys, 1867: 7 [stem: Scapter-]. Type genus: Scapterus Dejean, 1826. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Csiki (1927: 490, as Scapterina), generally accepted as in Balkenohl (2003: 231, as Scapterina).

3572Passalidiina Csiki, 1927: 489 [stem: Passalidi-]. Type genus: Passalidius Chaudoir, 1863.

3573Bonelli, 1810

3574Scaritides Bonelli, 1810: Tabula Synoptica [stem: Scarit-]. Type genus: Scarites Fabricius, 1775.

3575Jeannel, 1946

3576Storthodontini Jeannel, 1946: 237, in key [stem: Storthodont-]. Type genus: Storthodontus Chaudoir, 1855.

3577Hope, 1838

3578Broschidae Hope, 1838a: 80 [stem: Brosc-]. Type genus: Broscus Panzer, 1813.

3579Hope, 1838

3580Broschidae Hope, 1838a: 80 [stem: Brosc-]. Type genus: Broscus Panzer, 1813.

3581Roig-Juñent, 2000

3582Axonyina Roig-Juñent, 2000: 18 [stem: Axony-]. Type genus: Axonya Andrewes, 1923.

3583Jeannel, 1941

3584Barypitae Jeannel, 1941: 287, in key [stem: Baripod-]. Type genus: Baripus Dejean, 1828 [as Barypus, unjustified emendation of type genus name by Agassiz (1846b: 43), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.

3585Hope, 1838

3586Broschidae Hope, 1838a: 80 [stem: Brosc-]. Type genus: Broscus Panzer, 1813 [as Broschus, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.

3587Cephalotida Heer, 1838: 12 [stem: Cephalot-]. Type genus: Cephalotes Bonelli, 1810 [preoccupied genus name, not Cephalotes Latreille, 1802 [Hymenoptera]; syn. of Broscus Panzer, 1813]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.

3588Zacotini G. H. Horn, 1881: 169 [stem: Zacot-]. Type genus: Zacotus J. L. LeConte, 1869.

3589Jeannel, 1941

3590Creobitae Jeannel, 1941: 287, in key [stem: Creobi-]. Type genus: Creobius Guérin-Méneville, 1838. Comment: incorrect original stem formation, not in prevailing usage.

3591Roig-Juñent, 2000

3592Nothobroscina Roig-Juñent, 2000: 32 [stem: Nothobrosc-]. Type genus: Nothobroscus Roig-Juñent and Ball, 1995.

3593LeConte, 1853

3594Apotomi J. L. LeConte, 1853b: 370 [stem: Apotom-]. Type genus: Apotomus Illiger, 1807.

3595Bonelli, 1813

3596Siagones Bonelli, 1813: 24 [stem: Siagon-]. Type genus: Siagona Latreille, 1804.

3597Horn, 1881

3598Enceladini G. H. Horn, 1881: 118 [stem: Encelad-]. Type genus: Enceladus Bonelli, 1813.

3599Lecordier, 1977

3600Lupercini Lecordier, 1977: 628 [stem: Luperc-]. Type genus: Luperca Laporte, 1840.

3601Bonelli, 1813

3602Siagones Bonelli, 1813: 24 [stem: Siagon-]. Type genus: Siagona Latreille, 1804.

3603Csiki, 1933

3604Coscinides Chaudoir, 1876b: 115 [stem: Coscini-]. Type genus: Coscinia Dejean, 1831 [preoccupied genus name, not Coscinia Hübner, 1819 [Lepidoptera]; syn. of Cymbionotum Baudi, 1864]. Comment: permanently invalid (Art. 39): based on preoccupied type genus; incorrect original stem formation, not in prevailing usage.

3605Granigerini Bedel, 1900: 24 [stem: Graniger-]. Type genus: Graniger sensu Chaudoir, 1876 [not Graniger Motschulsky, 1864; syn. of Cymbionotum Baudi, 1864]. Comment: based on a misidentified type genus; an application will need to be submitted to the Commission to suppress this name for the Principles of Priority and Homonymy (Art. 65.2.1) if Granigerini Antoine, 1959 in Harpalinae: Harpalini: Harpalina is to be used as valid in the future.

3606Melaeninae Csiki, 1933: 1650 [stem: Melaen-]. Type genus: Melaenus Dejean, 1831. Comment: published 26 May 1933; name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Lorenz (1998b: 149) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).

3607Cymbionotini Andrewes, 1933: 3 [stem: Cymbionot-]. Type genus: Cymbionotum Baudi di Selve, 1864. Comment: published 30 June 1933.

3608Darlington, 1933

3609Gehringiini Darlington, 1933: 110 [stem: Gehringi-]. Type genus: Gehringia Darlington, 1933.

3610Darlington, 1933

3611Gehringiini Darlington, 1933: 110 [stem: Gehringi-]. Type genus: Gehringia Darlington, 1933.

3612Darlington, 1933

3613Gehringiini Darlington, 1933: 110 [stem: Gehringi-]. Type genus: Gehringia Darlington, 1933.

3614Deuve, 2007

3615Helenaeina Deuve, 2007: 217 [stem: Helenae-]. Type genus: Helenaea Schatzmayr and Koch, 1934.

3616Bonelli, 1810

3617Trechii Bonelli, 1810: Tabula Synoptica [stem: Trech-]. Type genus: Trechus Clairville, 1806.

3618Stephens, 1827

3619Bembidiidae Stephens, 1827: 5 [stem: Bembidi-]. Type genus: Bembidion Latreille, 1802 [as Bembidium, unjustified emendation of type genus name by Gyllenhal (1810: 12), not in prevailing usage].

3620Jeannel, 1937

3621Anillini Jeannel, 1937: 244, in key [stem: Anill-]. Type genus: Anillus Jacquelin du Val, 1851. Comment: precedence (Anillina Jeannel, 1937 vs Scotodipnina Jeannel, 1937) given to taxon originally proposed at the higher rank (Art. 24.1).

3622Scotodipnina Jeannel, 1937: 265, in key [stem: Scotodipn-]. Type genus: Scotodipnus Schaum, 1860.

3623Typhlocharina Jeanne, 1973: 95, in key [stem: Typhlocharit-]. Type genus: Typhlocharis Dieck, 1869. Comment: incorrect original stem formation, not in prevailing usage.

3624Stephens, 1827

3625Bembidiidae Stephens, 1827: 5 [stem: Bembidi-]. Type genus: Bembidion Latreille, 1802 [as Bembidium, unjustified emendation of type genus name by Gyllenhal (1810), not in prevailing usage].

3626Peryphidae Kirby, 1837: 52 [stem: Peryph-]. Type genus: Peryphus Dejean, 1821.

3627Motschulsky, 1862

3628Tachyaires Motschulsky, 1862: 24 [stem: Tachy-]. Type genus: Tachys Dejean, 1821 [placed on the Official List of Generic Names in Zoology (ICZN 1990c)]. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Jeannel (1941: 401, 422, as Tachyini), generally accepted as in J. K. Park et al. (2006: 91, as Tachyini); current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Tache-, see Alonso-Zarazaga 2007).

3629Micratopini Casey, 1914a: 42 [stem: Micratop-]. Type genus: Micratopus Casey, 1914.

3630Limnastini Jeannel, 1937: 245, in key [stem: Lymnast-]. Type genus: Lymnastis Motschulsky, 1862 [as Limnastis, unjustified emendation of type genus name by Ganglbauer (1891a: 181), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.

3631Erwin, 1994

3632Xystosomina Erwin, 1994: 560 [stem: Xystosom-]. Type genus: Xystosomus Schaum, 1863.

3633Jeannel, 1949

3634Horologionidae Jeannel, 1949b: 91 [stem: Horologion-]. Type genus: Horologion J. M. Valentine, 1932. Comment: current spelling maintained (Art. 29.5): incorrect stem formation in prevailing usage (should be Horologi-).

3635Laporte, 1834

3636Pogonidae Laporte, 1834b: 70 [stem: Pogon-]. Type genus: Pogonus Dejean, 1821.

3637Pogonopsini Bedel, 1900: 20 [stem: Pogonopse-]. Type genus: Pogonopsis Bedel, 1898. Comment: incorrect original stem formation, not in prevailing usage.

3638Bonelli, 1810

3639Trechii Bonelli, 1810: Tabula Synoptica [stem: Trech-]. Type genus: Trechus Clairville, 1806.

3640Fowler, 1887

3641Aëpyes Fowler, 1887: 123 [stem: Aep-]. Type genus: Aepus Samouelle, 1819.

3642Temnostegini Enderlein, 1909: 376 [stem: Temnosteg-]. Type genus: Temnostega Enderlein, 1905.

3643Jeannel, 1958

3644Cnidina Jeannel, 1958b: 733 [stem: Cnid-]. Type genus: Cnides Motschulsky, 1862. Comment: name proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1), however available because it was used as valid before 2000 as in Casale and Laneyrie (1982: 9, as Cnidini) and was not rejected by an author who, between 1961 and 1999, applied Article 13 of the then current edition of the Code (see Art. 13.2.1).

3645Sloane, 1903

3646Perileptides Sloane, 1903: 583 [stem: Perilept-]. Type genus: Perileptus Schaum, 1860. Comment: Perileptides Sloane, 1903 was treated as Latin and available in Madge (1989: 466) but the valid name for this taxon was listed as Perilepti Jeannel, 1922 by Csiki (1928) and Perileptina Jeannel, 1922 by Lorenz (2005); we follow the usage of Madge (1989).

3647Ochthephilini Jeannel, 1922: 165 [stem: Ochthephil-]. Type genus: Ochthephilus Nietner, 1857 [preoccupied genus name, not Ochthephilus Mulsant and Rey, 1856 [Coleoptera: Staphylinidae]; syn. of Perileptus Schaum, 1860]. Comment: permanently invalid (Art. 39): based on preoccupied type genus.

3648Jeannel, 1960

3649*Plocamotrechina Jeannel, 1958b: 733 [stem: Plocamotrech-]. Type genus: Plocamotrechus Jeannel, 1926 [syn. of Pachydesus Motschulsky, 1864]. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).

3650Plocamotrechina Jeannel, 1960: 53, in key [stem: Plocamotrech-]. Type genus: Plocamotrechus Jeannel, 1926 [syn. of Pachydesus Motschulsky, 1864].

3651Bonelli, 1810

3652Trechii Bonelli, 1810: Tabula Synoptica [stem: Trech-]. Type genus: Trechus Clairville, 1806.

3653*Aphaenopses Fauvel, 1888: 7 [stem: Aphoenop-]. Type genus: Aphoenops Bonvouloir, 1861 [as Aphaenops, incorrect subsequent spelling of type genus name, not in prevailing usage]. Comment: original vernacular name unavailable (Art. 11.7.2): not subsequently latinized; incorrect original stem formation, not in prevailing usage.

3654Homaloderini Jeannel, 1926: 397, in key [stem: Omaloder-]. Type genus: Omalodera Blanchard, 1842 [as Homalodera, unjustified emendation of type genus name by Gemminger and Harold (1868a: 389), not in prevailing usage]. Comment: incorrect original stem formation, not in prevailing usage.

3655Jeannel, 1926

3656Trechodini Jeannel, 1926: 469 [stem: Trechod-]. Type genus: Trechodes Blackburn, 1901.

3657Thalassophili Csiki, 1928: 233 [stem: Thalassophil-]. Type genus: Thalassophilus Wollaston, 1854.

3658Sharp, 1886

3659Zolini Sharp, 1886a: 371 [stem: Zol-]. Type genus: Zolus Sharp, 1886.

3660Roig-Juñent and Cicchino, 2001

3661Chalteniina Roig-Juñent and Cicchino, 2001: 658 [stem: Chalteni-]. Type genus: Chaltenia Roig-Juñent and Cicchino, 2001.

3662Deuve, 1997

3663Sinozolini Deuve, 1997: 35 [stem: Sinozol-]. Type genus: Sinozolus Bedel, 1898.

3664Sharp, 1886

3665Zolini Sharp, 1886a: 371 [stem: Zol-]. Type genus: Zolus Sharp, 1886.

3666Merizodini Sloane, 1920: 139 [stem: Merizodont-]. Type genus: Merizodus Solier, 1849. Comment: incorrect original stem formation, not in prevailing usage.

3667*Oopterini Jeannel, 1938: 45 [stem: Oopter-]. Type genus: Oopterus Guérin-Méneville, 1841. Comment: unavailable family-group name, proposed after 1930 without description or bibliographic reference to such a description (Art. 13.1).

3668Oopterini Jeannel, 1940: 93 [stem: Oopter-]. Type genus: Oopterus Guérin-Méneville, 1841.

3669Kirby, 1837

3670Patrobidae Kirby, 1837: 50 [stem: Patrob-]. Type genus: Patrobus Dejean, 1821.

3671Zamotajlov, 2000

3672Lissopogonini Zamotajlov, 2000: 266 [stem: Lissopogon-]. Type genus: Lissopogonus Andrewes, 1923.

3673Zolinopatrobina Deuve and Tian, 2001: 421 [stem: Zolinopatrob-]. Type genus: Zolinopatrobus Deuve and Tian, 2001 [syn. of Lissopogonus Andrewes, 1923].

3674Kirby, 1837

3675Patrobidae Kirby, 1837: 50 [stem: Patrob-]. Type genus: Patrobus Dejean, 1821.

3676Chaudoir, 1871

3677Deltomeridae Chaudoir, 1871a: 51 [stem: Deltomer-]. Type genus: Deltomerus Motschulsky, 1850.

3678Zamotajlov, 2002

3679Deltomerodina Zamotajlov, 2002: 83 [stem: Deltomerod-]. Type genus: Deltomerodes Deuve, 1992.

3680Kirby, 1837

3681Patrobidae Kirby, 1837: 50 [stem: Patrob-]. Type genus: Patrobus Dejean, 1821.

3682Zamotajlov and Lafer, 2001

3683Platidiolini Zamotajlov and Lafer, 2001: 411 [stem: Platidiol-]. Type genus: Platidiolus Chaudoir, 1878.