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World checklist of hornworts and liverworts

Lars Söderström, Anders Hagborg, Matt von Konrat and 36 more

PhytoKeys · 2016 · CC BY

doi:10.3897/phytokeys.59.6261

Abstract

A working checklist of accepted taxa worldwide is vital in achieving the goal of developing an online flora of all known plants by 2020 as part of the Global Strategy for Plant Conservation. We here present the first-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 7486 species in 398 genera representing 92 families from the two phyla. The checklist has far reaching implications and applications, including providing a valuable tool for taxonomists and systematists, analyzing phytogeographic and diversity patterns, aiding in the assessment of floristic and taxonomic knowledge, and identifying geographical gaps in our understanding of the global liverwort and hornwort flora. The checklist is derived from a working data set centralizing nomenclature, taxonomy and geography on a global scale. Prior to this effort a lack of centralization has been a major impediment for the study and analysis of species richness, conservation and systematic research at both regional and global scales. The success of this checklist, initiated in 2008, has been underpinned by its community approach involving taxonomic specialists working towards a consensus on taxonomy, nomenclature and distribution.

Full text

From PubMed Central, U.S. National Library of Medicine, as current on Oct 25, 2026. Paragraph numbers are ours, so an answer can point at one.

This checklist is dedicated to Elizabeth Brown and Ray Stotler, who sadly left us during our work with it.

“……Charles Darwin wrote of his desire to provide financial support ‘for the formation of a perfect M.S. catalogue of all known plants’ (Darwin 1881, letter 13570). It is a personal embarrassment to me, and should be chastening to us all, that more than 120 years later we still have not delivered on that commitment.”.

Introduction

The natural world is changing fast. The Global Strategy for Plant Conservation (GSPC) was adopted by the Conference of Parties (COP) of the Convention on Biological Diversity (CBD) in April 2002 (http://www.cbd.int/decision/cop/?id=7183; accessed 2014.06.02). The Strategy set out 16 outcome-oriented targets that were to be achieved by 2010. The GSPC was designed as a framework for action to halt the loss of plant diversity. Target 1 of the Strategy was to complete “a widely accessible working list of all known plant species, as a step towards a complete world Flora”. As early as 1881 Charles Darwin expressed a wish to have a catalogue of all known plants. However, over 125 years later, this wish is not yet fulfilled. Without a working checklist, many of the other objectives in the GSPC cannot be met and botanists around the world cannot communicate about plants on a global basis. A working list of known plant species is critical for (I) its underpinning role in the effective implementation of the other targets through provision of baseline information; (II) increasing the accessibility and use of accurate botanical name information for research, conservation and sustainable use; and (III) real-world politics and how taxonomists respond to the decisions of policy makers.

Global Strategy for Plant Conservation

Conference of Parties

Convention on Biological Diversity

Version 1.0 of The Plant List (http://www.theplantlist.org/) was released in December 2010 aimed to be comprehensive for species of vascular plants (flowering plants, conifers, ferns and their allies) and of bryophytes (mosses, liverworts and hornworts), as a response to the 2010 Target 1 of the GSPC and to a clear global need for such data. The Plant List is a broad collaboration, coordinated by the Royal Botanic Gardens, Kew, and the Missouri Botanical Garden (MO), involving diverse partnerships. noted The Plant List represents a work in progress with future versions planned to improve the quality of names and decrease the number of unresolved names. Target 1 has since been revised and agreed at the 10th COP of the CBD in Nagoya, Japan, with the goal of developing ‘‘an online flora of all known plants’’ by 2020 (http://www.cbd.int/decision/cop/default.shtml?id=12283; accessed 2014.06.02). identified several factors that might assist in the implementation of the revised target, stressing that achieving a working list for plant taxa worldwide was a vital step to an online flora.

Earlier, assessed the progress made at that time and discussed prospects for the completion of Target 1. Paton noted that good progress had been made in bryophytes (mosses, liverworts and hornworts), ferns and gymnosperms with widely accessible working lists either complete or almost so for those groups. For bryophytes alone, he tabulated 13,370 accepted species noting that the data was largely derived from Tropicos. However, although Tropicos is an indispensable reference for anyone dealing with bryophyte names and the database is very strong for mosses, the nomenclatural and auxiliary data for liverworts is less complete, especially for larger genera. The lack of a central source providing a synthesis of nomenclatural data and global distributional data was the impetus toward developing the current checklist of liverworts and hornworts. Checklists are powerful and important tools that can integrate the almost overwhelmingly scattered information concerning taxonomy, systematics, nomenclature, distribution, and even frequency.

Ecological and biological significance of liverworts and hornworts

Liverworts and hornworts (Figure 1) are of critical biological, ecological, and phylogenetic significance (e.g.,,,,, ). Liverworts are found on soil, rocks, and trees throughout the world, from coastal Antarctica to the tundra of the Northern Hemisphere, and from semi-arid areas of Australia to the Amazon rainforests. Although there are xero-tolerant taxa, the majority of liverworts are found in relatively humid and shaded terrestrial ecosystems. Liverworts and hornworts are an important component of the vegetation in many regions of the world, constituting a major part of the biodiversity in moist forest, wetlands, mountain, and tundra ecosystems.

Liverworts, hornworts and mosses offer microhabitats that are critical to the survival of a tremendous diversity of organisms such as single-celled eukaryotes, protozoa, and numerous groups of invertebrates. Their structural contribution to levels of diversity might be as significant as that of vascular plants, albeit at a smaller scale. These plants are also important environmental and ecological indicators. Liverworts, in concert with mosses and hornworts, play a significant role in the global carbon budget and CO2 exchange, plant succession, net production and phytomass, nutrient cycling, and water retention. These groups of land plants also have been used as indicators of past climate change, to validate climate models, and as potential indicators of global warming.

Species numbers

There are estimated to exist only 215 hornwort species. By comparison estimates of liverwort species richness have varied considerably, by as much as 50%, with estimates ranging from 4,500 to 9,000 (e.g., ). In the last decade or so estimates in the range of 5,000 to 6,000 liverwort species have been widely accepted (e.g.,, ). provided a mean estimate of 7,500 for the number of liverwort species based on estimating rates of synonymy in a sample of recently monographed and revised taxa. A standardized global worldwide liverwort checklist with strong community participation, as presented here, coupled with the critical need for ongoing monographs and revisions, will aid in arriving at clearer estimates of liverwort and hornwort diversity. Significantly, a list of standardized names reviewed by taxonomic experts will enable more meaningful geographical and biogeographical species comparative studies. The global checklist of accepted liverwort and hornwort taxa has vast potential, not only in aiding our understanding of liverwort diversity, patterns, and processes, but also to the broader biological community.

Below we provide context to the worldwide checklist, including a brief historical account and current informational resources, structure and layout, systematic and classification concepts, and methodology, including a brief overview of the underlying data set and nomenclatural elements.

Major historical works

has provided a detailed account on the history of hepaticology, up to the first half of the 20th century. Earlier, also outlined a valuable historical review of important figures in hepaticology. In the early 19th century, possibly the two most influential persons who provided an early framework for taxonomic concepts were W.J. Hooker and G. Raddi. During the middle of the 19th century C.G. Nees von Esenbeck, C.M. Gottsche and J.B.W. Lindenberg provided vastly significant contributions towards developing a classification framework. provided a brief biography of these workers. This was later followed by R. Spruce and the Austrian-born H. Leitgeb in the second half 19th century. Leitgeb in particular was instrumental in liverwort plant morphology and anatomy including a six part treatise, “Untersuchungen ueber die Lebermoose”, from 1874 to 1881. Towards the end of 19th century and early 20th century V. Schiffner, F. Stephani, and A. Evans were prominent hepaticologists. In the first half of the 20th century A. Evans, H. Buch, F. Verdoorn, and K. Müller (Frib.) were leading influential hepaticologists. In the second half of the 20th century, R. Grolle and R.M. Schuster were of extraordinary influence in working towards contemporary classification schemes. Many of these groundbreaking concepts still serve the foundation from which our current classification scheme is derived. R.M. Schuster’s major contribution to botany and hepaticology lies in the astounding new diversity of liverworts he added to our knowledge. He described a staggering 463 species, 83 genera and 15 families new to science. It is difficult to name another contemporary botanist who discovered this much new diversity of a major clade of land plants. R.M. Schuster’s contribution to botany went beyond hepatics. He was one of the first botanists who recognized the importance of Wallace’s Line in plant biogeography, separating Australia of Gondwanaland from Southeast Asia of Laurasia.

Figure 2 depicts the number of novel liverwort species, excluding new combinations that have been described over the last 250 years. The first major peak corresponds to the works of several early 19th century botanists, including Synopsis Hepaticarum by Gottsche, Lindenberg, and Nees (1844–1847). The three decades leading into, but prior to the highest peak, between 1860 and 1890, correspond to publications by a number of prominent bryologists including W. Mitten, J.D. Hooker, T. Taylor, and V. Schiffner. The second and highest peak of almost 1,200 names, in the early 1900s corresponds largely to the plethora of taxa described by F. Stephani (1898–1924) in his monumental work Species Hepaticarum. The periods of highest rates of new species described in the 1830s and around 1900 are the same for seed plants. The third peak over the four decades between 1950 and 1980 can be attributed mainly to the works of R.M. Schuster, H. Inoue, and S. Hattori. H. Inoue and S. Hattori were influential on large species-rich genera such as Plagiochila, Frullania, and Porella. The decline in newly described species since 1970 does not necessarily translate to the conclusion that taxonomists are closer to discovering all known species. The almost 200 novel liverwort species that have been described in the past six years alone still represent a significant number, considering the relatively few liverwort taxonomists and monographers. Moreover, bryological exploration has been very uneven in many parts of the world. For example, many areas of the Neotropics still remain without a single bryophyte record. Paradoxically, scores of new species are still being discovered and described in relatively well-studied areas such as New Zealand, e.g. over 50 new taxa since 2000 alone (e.g.,,, ). Recent attention to cryptic speciation in bryophytes is also revealing novel liverwort species (e.g. ). The combination of collecting in yet-to-be explored areas, the continued discovery of species in well-studied regions, and an increased understanding of the biology of liverworts (including cryptic speciation), will lead to a significant number of newly discovered species in the foreseeable future. The corollary of this, coupled with increased monographic and revisional work, will be the increased discovery and the unravelling of synonymy.

Contemporary resources

Data availability and information needs associated with liverworts and hornworts have been reviewed extensively by, ). A fundamental problem common to all nomenclatural indexing projects is the dispersed nature of the biological literature, some of which may date back over 250 years. Our own data records, from 1990 to the present alone, indicate there are over 190 periodical titles and non-serials in which liverwort nomenclatural novelties were published. The most useful and successful web-accessible bryophyte nomenclatural database is Tropicos (http://www.tropicos.org), which offers name data with references and type information as well as links to specimen data of its holdings. Tropicos has therefore become an indispensable reference for anyone dealing with bryophyte names. However, as stated above, the database is particularly strong for mosses and far less comprehensive for liverworts. Moreover, Tropicos does not prescribe taxonomic disposition of names, i.e., there is no attempt to adopt a single consistent view on the status of any particular name.

A major nomenclatural work is the Index Hepaticarum (http://www.ville-ge.ch/musinfo/bd/cjb/hepatic/index.php), which includes all effectively published liverwort epithets spanning 12 volumes with the closing date of 1973. The indices were prepared as a purely nomenclatural resource and did not claim to express any particular taxonomic concept. Recently, provided an equally valuable nomenclatural resource and catalog of names at all ranks for liverworts and hornworts published during 1974 to 2000. Subsequently, indices of the citations for names published for bryophytes have been compiled for the years 2001-2004 and 2005. The Early Land Plants Today initiative has continued this series.

The application of molecular phylogenetics continues to generate new insights into the evolutionary history of liverworts and hornworts. In recent years, inferences made from these phylogenies have especially revolutionized liverwort classification. Yet, the reconstruction of the phylogenetic history of the Marchantiophyta and Anthocerotophyta remains an ongoing effort and the classification is fluid. The current checklist largely follows the comprehensive phylogenetic liverwort classification scheme provided by and the hornwort classification scheme of. Thus the checklist largely adheres to their higher classification namely from class to suborder, and the majority of family and genera arrangements provided therein. However, there are some notable exceptions, especially in the systematic treatment of liverworts, which largely either reflect publications subsequent to 2009, or are a slight departure in taxonomic opinion based on earlier works, e.g. the concepts of Saccogynaceae and Stephaniellaceae depart from. Since 2010 there have been a number of realignments and novel hypotheses proposed that were generated from conclusions based on molecular evidence which have influenced the systematic treatment presented here. These include, for example, Adelanthaceae (e.g.,,,, ), Anastrophyllaceae (e.g. ), Gymnomitriaceae (e.g.,,, ), Lepidoziaceae, Lophocoleaceae, and Scapaniaceae (e.g., ). In all cases, any such departures, new families, or realignments are noted and very briefly discussed throughout the checklist. As noted by many small families still remain to be investigated using molecular data, and many of the large families and genera that have been sampled appear to be either polyphyletic or paraphyletic.

The systematic treatment as outlined here also includes the addition of subfamily and infrageneric classification; ranks not presented by, ). Although there are arguably a number of ambiguous placements, especially at the infrageneric rank, and there remain many areas of contention, particularly at the familial and generic level, the systematic treatment itself serves as an excellent synopsis of contemporary taxonomic opinion, including infrageneric ranks. The infrageneric treatments will also aid future efforts in providing a taxonomic framework that will be particularly useful for extremely large genera. In large part, the infrageneric treatments only deal with those taxa that represent phylogenetic clades with strong support. Care has been taken to reduce ambiguity and those taxa where infrageneric placement is weak or without evidence are simply listed under the genus and not placed systematically.

Structure and format of the checklist

The checklist is presented in a taxonomic framework as outlined above with four main sections. The first is arranged systematically with taxa ordered alphabetically within the nearest higher rank. A brief discussion for selected families is provided in the main body of the text under the respective families. This does not appear uniformly for all families, but typically for larger more complicated families. Rather, a brief description is provided mainly where concepts depart from or if new information has come to light since that publication. Subfamilies and infrageneric ranks have been incorporated where appropriate. However, if the subgeneric placement is unclear the species are listed under “Incertae sedis”. From the rank of genus and below, all entries include a three star confidence ranking (described below), accepted taxon name, authority, abbreviated publication title, page number and date, which is cross- referenced to the full citation in the reference list, as well as the basionym if appropriate. A typical entry appears as follows:

*** Oleolophozia L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010.

*** Oleolophozia perssonii (H.Buch et S.W.Arnell) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 51, 2010. Bas.: Lophozia perssonii H.Buch et S.W.Arnell, Bot. Not. 97: 382, 1944.

There are comprehensive footnotes throughout the systematic section of the checklist. These are applied to selected names representing the rank of genus and below only. Footnotes without any literature citation represent observations from the authors of the respective family. Footnotes are wide ranging in their content. Generally, footnotes can be categorized into the following: i) general statements relating to taxonomic opinion, ii) comments on the type specimen, iii) comments about possible conspecificity, including citation of conflicting opinions, and iv) comment about species complexes.

This section is followed by a list of taxa in genera that we do not recognize. Those names represent taxa that have not been studied recently and have not been recognized in any recent treatment. However, many of those names are old and may gain priority over some recognized taxa once their identity is determined.

The next section is an alphabetical list of taxa of the ranks of species and below. The alphabetical list includes the confidence level, the taxon name and authority as well as a reference to the page of the corresponding entry in the systematic section. Thus, the alphabetical list provides a rapid gateway requiring no prior knowledge of higher taxonomy, but also serves as an index to the corresponding name in the systematic section that includes more detailed information.

The last section is a reference list where full bibliographic citations are given to all references for taxa included in the checklist. Titles and references have all been checked and verified, except in a few cases where we could not get hold of the publications.

Taxon confidence levels

Significantly, each accepted taxon is qualified using a three level ranking system that summarizes our knowledge about a taxon. The coding convention we are adopting largely follows that described by using one to three stars, which has been applied to recent regional checklists produced by the Early Land Plants Today initiative, e.g.,,,, ). The conventions are briefly outlined below coupled with samples illustrating how these conventions are applied in practice. The application of a confidence level to a taxon’s status and whether it represents a genuine “species” that is reached through community consensus may go towards refining species estimates using an evidence-based approach. The confidence levels coupled with the detailed annotations in the form of footnotes may also provide a rapid assessment of taxa and help aid and drive future research into specific taxonomic or nomenclatural problems and issues.

The coding convention:

* Serious doubts. There are doubts about the value of the taxon. It can be that there are conflicting views without any substantial evidence in any direction, conflicting views with substantiating evidence supporting one or both positions, or evidence pointing towards synonymization but it is premature to do it. We have adhered to the principle that it is better to keep a taxon with one star (and preferably a note) than to synonymize it too quickly. Example: Bazzania asymmetrica is conspecific with Bazzania macgregorii in, but kept them separate. Example: Nardia kamtschatica may be conspecific with Nardia assamica, but the type specimen could not be studied.

** Knowledge problem. The taxon is not well known by the person evaluating it. It may be a newly described species or a species originally not well described and not restudied recently. Example: Jungermannia erectii Ajit P.Singh et V.Nath was recently described and has not been independently studied by someone with a global overview. It is therefore difficult to evaluate.

*** Accepted. A good taxon as currently understood based on personal experience or on taxonomic revisions that have been convincingly performed. Nomenclature and/or taxonomic position may, however, be questioned. Elements may be excluded from the taxon, but the taxon with the current type will still be accepted. Example: H. Bischler-Causse revised the genus Marchantia worldwide in a series of publications. Although she had a broad species concept, she also recognized infraspecific taxa. The taxa that she recognized without doubt should be accepted unless new evidence against it exists. Adopting a narrower species concept so that many of her subspecies are elevated to species does not change her view of what a good taxon is.

Methodology

The foundation of the checklist is in the underlying data set from which it is derived. It was briefly described by, ). The working data sets now includes a bibliography of 25,000 publications; approximately 39,000 published liverwort names (including “accepted” taxa, synonyms, invalid and illegitimate names). The data quality and standards were outlined by. In summary, for authorities and for the citation abbreviations, we follow the standards set by the on-line version of Authors of Plant Names at the Royal Botanical Gardens, Kew Website (http://www.ipni.org), with whom we collaborate closely and provide with updated data records. Publications and journal abbreviations follow Taxonomic Literature: A selective guide to botanical publications and collections with dates, commentaries and types (Stafleu and Cowan 1976–2009) (http://www.sil.si.edu/digitalcollections/tl-2/search.cfm) and Botanico Periodicum Huntianum (http://huntbot.andrew.cmu.edu/HIBD/Departments/Databases.shtml).

There has been an intense systematic effort focusing on data quality. In all but a few cases nomenclatural data has been verified against original publications. Söderström and Hagborg have checked and confirmed almost every original publication (three publications have not been available) for correct author, title and journal/book citation, date of publication as far as possible, page number for the protologue, and if the name is validly and legitimately published according to the International Code of Nomenclature for algae, fungi and plants (Melbourne Code; ). Most significantly, the checklist has been community driven and collaborative. Broader participation by taxonomic specialists and regional experts has lead to the checklist containing high quality data (cf. ). Development of the checklist has included three international meetings (the first one in 2009) and generated 74 published notes on taxonomy and nomenclature under the auspices of the Early Land Plants Today initiative from 2012 to 2015. The series of meetings were instrumental in providing a framework for direct interaction with taxonomic experts, workshops reviewing names, and helping identify potential participants who were taxonomic experts in specific taxonomic groups, whether these were individual genera or entire families.

The notes, published in the journal Phytotaxa, have included 38 authors from 13 countries. These are the results of several years of revisions of taxonomic groups by taxonomic experts. Following these revisions and working with participants identifying nomenclatural and taxonomic problems that would impact the worldwide checklist, Söderström, Hagborg and von Konrat led editorial efforts in compiling manuscripts with baseline information and nomenclatural or taxonomic issues that required resolving. The work was then coordinated with the taxonomic experts who drove and led the process and often identified further issues. The series of notes on taxonomy and nomenclature of liverworts and hornworts provided updated nomenclature, corrected invalid and illegitimate names and described new taxa based on studies (mainly molecular) that did not draw nomenclatural conclusions. Those notes are all open access and any changes were effective immediately as the Melbourne Code of Botanical Nomenclature allows publishing on internet from 1 Jan. 2012.

This checklist builds on all published taxonomic and nomenclatural papers, that have come to our attention until June 30, 2015.

Summary statistics

We here present the first-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 8,078 taxa (species and below) in 7,486 species representing 398 genera, 92 families, 20 orders and 7 classes from the two phyla. The list includes 3,533 species with three stars, 2,988 species with two stars and 915 species with one star. The checklist also has extra utility in that it contains 3,106 references in the bibliography that serve as a powerful bibliographic resource for liverwort and hornwort systematic and taxonomic research.

Concluding remarks

The marked-up publication form of the current checklist by PhytoKeys provides a virtual instrument with a linked environment both internally (e.g. within an article) and externally (GBIF, IPNI, Tropicos, Wikispecies, etc.) that will undoubtedly help accelerate taxonomic research. The published checklist was the first phase in providing a worldwide list of accepted names. The next phase is to establish a generally recognized online repository to augment the huge underlying informational auxiliary data of over 25,000 publications, almost 39,000 published names, and the over 700,000 geographical observations. Several features of Web-based technology make it an essential tool, including 1) a vehicle to facilitate data access, 2) to unify the vastly scattered data on distributional information and nomenclature, 3) offering dynamic rather than static data in a searchable forum. The broader accessibility to the wealth of auxiliary data will help augment monographic and revisional work for many taxonomic groups, aid in identifying the need for increased floristic and survey work in many regions throughout the world, and have broad implications and applications beyond taxonomic research such as conservation science. The current Early Land Plants Today model has strong participation from taxonomic experts and an online resource will provide an opportunity to expand stakeholders to include ecologists, conservationists, scientists from other disciplines and general interest groups. However, such an effort can only be successful if it comes with sustained funding and infrastructure rather than depending on an ad hoc commitment by a few individuals, however dedicated.

The project will help augment monographic and revisional work for many taxonomic groups and aid in identifying the need for increased floristic and survey work in many regions throughout the world. Although there are many challenges ahead to obtain high quality data, quantifying global liverwort diversity is a tractable, multi-faceted and scientifically important goal, and everyone stands to gain by fostering this endeavour. The success of the project will lie on strong collaboration between institutions and the bryological community in general.

*** Anthocerotaceae Dumort.

by J.C. Villarreal and D.C. Cargill

Notes on nomenclature and taxonomy can also be found in and

*** Anthoceros L., Sp. Pl. 1: 1139, 1753. 1

*** Anthoceros adscendens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 24, 1832.

*** Anthoceros agrestis Paton, J. Bryol. 10 (3): 257, 1979, nom. conserv. 2

*** Anthoceros alpinus Steph., Sp. Hepat. (Stephani) 6: 425, 1923.

* Anthoceros angustifolius Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 582, 1846. 3

*** Anthoceros angustus Steph., Sp. Hepat. (Stephani) 5: 1001, 1916.

*** Anthoceros bharadwajii Udar et A.K.Asthana, Proc. Indian Natl. Sci. Acad., B 51 (4): 484, 1985.

** Anthoceros buettneri Steph., Sp. Hepat. (Stephani) 5: 997, 1916.

*** Anthoceros capricornii Cargill et G.A.M.Scott, J. Hattori Bot. Lab. 82: 55, 1997.

*** Anthoceros caucasicus Steph., Izv. Kavkazsk. Muz. 8: 87, 1914.

** Anthoceros cavernosus Steph., Sp. Hepat. (Stephani) 5: 998, 1916.

* Anthoceros chambensis Kashyap, J. Bombay Nat. Hist. Soc. 25 (2): 281, 1917.

** Anthoceros chungii Khanna, J. Indian Bot. Soc. 17 (5/6): 316, 1938.

** Anthoceros crispatus Griff., Not. pl. asiat. 2: 349, 1849.

** Anthoceros dimorphus Sim, Trans. Roy. Soc. South Africa 15 (1): 114, 1926.

*** Anthoceros erectus Kashyap, New Phytol. 14 (1): 9, 1915.

** Anthoceros expansus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 92, 2015. Bas.: Aspiromitus expansus Steph., Sp. Hepat. (Stephani) 5: 961, 1916.

* Anthoceros ferdinandi-muelleri Steph., Sp. Hepat. (Stephani) 5: 1007, 1916. 4

*** Anthoceros fragilis Steph., Sp. Hepat. (Stephani) 5: 1006, 1916.

*** Anthoceros fusiformis Austin, Bull. Torrey Bot. Club 6 (4): 28, 1875 [1876].

*** Anthoceros fusiformis var. taiwanensis J.Haseg., Acta Phytotax. Geobot. 44 (2): 100, 1993.

** Anthoceros gasongorii Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 10, 1916.

** Anthoceros granulatus Gottsche, Mexik. Leverm.: 275, 1863.

** Anthoceros harrisanus (Steph.) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 31, 1962. Bas.: Aspiromitus harrisanus Steph., Sp. Hepat. (Stephani) 5: 965, 1916.

* Anthoceros helmsii Steph., Hedwigia 32 (3): 142, 1893. 5

** Anthoceros jamesonii Taylor ex Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 171, 1855.

* Anthoceros javanicoides H.A.Mill., Phytologia 47 (4): 319, 1981. Nom. nov. pro Anthoceros javanicus Steph., Sp. Hepat. (Stephani) 5: 988, 1916, nom. illeg. 6

** Anthoceros jungermannioides Schwein., Spec. Fl. Amer. Crypt.: 25, 1821.

* Anthoceros kajumas (K.I.Goebel) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951. Bas.: Aspiromitus kajumas K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 69, 1928.

* Anthoceros koshii Khanna, J. Indian Bot. Soc. 15 (4): 237, 1936.

*** Anthoceros lamellatus Steph., Sp. Hepat. (Stephani) 5: 1000, 1916.

*** Anthoceros laminifer Steph., J. Linn. Soc., Bot. 29 (201): 266, 1892.

*** Anthoceros macounii M.Howe, Bull. Torrey Bot. Club 25 (1): 19, 1898.

*** Anthoceros macrosporus Steph., Sp. Hepat. (Stephani) 5: 1005, 1916.

** Anthoceros maritimus Steph., Sp. Hepat. (Stephani) 5: 984, 1916.

* Anthoceros megasporus Meijer, J. Hattori Bot. Lab. 18: 6, 1957.

** Anthoceros muscoides Colenso, Trans. & Proc. New Zealand Inst. 16: 361, 1884.

*** Anthoceros myriandroecius Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 134, 1911.

** Anthoceros natalensis Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 732, 1913.

*** Anthoceros neesii Prosk., Leberm. Eur. 2 (9): 1312, 1958. Nom. nov. pro Anthoceros punctatus α* monocarpus Nees, Naturgesch. Eur. Leberm. 4: 339, 1838.

** Anthoceros niger Steph., Sp. Hepat. (Stephani) 5: 1005, 1916.

** Anthoceros orizabensis (Steph.) Hässel, Candollea 45 (1): 211, 1990. Bas.: Aspiromitus orizabensis Steph., Sp. Hepat. (Stephani) 5: 965, 1916.

*** Anthoceros pandei Udar et A.K.Asthana, J. Indian Bot. Soc. 64: 305, 1985.

*** Anthoceros patagonicus Hässel, Candollea 45 (1): 207, 1990.

*** Anthoceros patagonicus subsp. gremmenii J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 32, 2013.

** Anthoceros peruvianus Steph., Sp. Hepat. (Stephani) 5: 999, 1916.

** Anthoceros pinnatus Steph., Bol. Soc. Brot. 4: 154 [182], 1885 [1886].

*** Anthoceros punctatus L., Sp. Pl. 1: 1139, 1753. 7

* Anthoceros pusillus Colenso, Trans. & Proc. New Zealand Inst. 18: 255, 1886.

*** Anthoceros rosulans J.Haseg., J. Hattori Bot. Lab. 60: 379, 1986.

*** Anthoceros sambesianus Steph., Sp. Hepat. (Stephani) 5: 996, 1916.

*** Anthoceros scariosus Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 230, 1869.

** Anthoceros schroederi Steph., 52 (5): 306, 1912.

** Anthoceros serratus Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 90, 1911.

** Anthoceros simulans M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 204, 1934.

** Anthoceros spongiosus Steph., Sp. Hepat. (Stephani) 5: 1003, 1916.

*** Anthoceros subtilis Steph., Sp. Hepat. (Stephani) 5: 1003, 1916.

*** Anthoceros telaganus Steph., Sp. Hepat. (Stephani) 5: 1005, 1916.

*** Anthoceros tristanianus J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 33, 2013.

*** Anthoceros tuberculatus Lehm. et Lindenb., Nov. Stirp. Pug. 4: 25, 1832.

** Anthoceros venosus Lindenb. et Gottsche, Syn. Hepat. 4: 584, 1846.

Excluded from the genus

* Anthoceros aethyopicus Gola, Ann. Bot. (Rome) 13 (1): 73, 1914. 8

* Anthoceros brunnthaleri Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 732, 1913. 9

* Anthoceros floribundus Steph., Sp. Hepat. (Stephani) 5: 977, 1916. 10

* Anthoceros mildbraedii Steph., Sp. Hepat. (Stephani) 6: 428, 1923. 11

* Anthoceros parvifrons Steph., 52 (5): 307, 1912. 12

* Anthoceros pseudocostus Steph., Sp. Hepat. (Stephani) 5: 977, 1916. 13

* Anthoceros rossoi Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 10, 1916. 14

*** Folioceros D.C.Bharadwaj, Geophytology 1 (1): 9, 1971.

*** Folioceros amboinensis (Schiffn.) Piippo, Acta Bot. Fenn. 148: 36, 1993. Bas.: Anthoceros amboinensis Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 45, 1890.

** Folioceros apiahynus (Steph.) Hässel, Candollea 45 (1): 215, 1990. Bas.: Anthoceros apiahynus Steph., Sp. Hepat. (Stephani) 5: 999, 1916.

** Folioceros argillaceus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 93, 2015. Bas.: Aspiromitus argillaceus Steph., Sp. Hepat. (Stephani) 5: 970, 1916.

*** Folioceros assamicus D.C.Bharadwaj, Geophytology 1 (1): 10, 1971.

** Folioceros dilatatus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 93, 2015. Bas.: Anthoceros dilatatus Steph., Bot. Jahrb. Syst. 8 (2): 95, 1886.

*** Folioceros dixitianus (Mahab.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975. Bas.: Aspiromitus dixitianus Mahab., Curr. Sci. 10 (12): 532, 1941.

*** Folioceros fuciformis (Mont.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975. Bas.: Anthoceros fuciformis Mont., Ann. Sci. Nat. Bot. (sér. 2) 20: 296, 1844.

*** Folioceros glandulosus (Lehm. et Lindenb.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975. Bas.: Anthoceros glandulosus Lehm. et Lindenb., Nov. Stirp. Pug. 4: 26, 1832.

*** Folioceros incurvus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975. Bas.: Anthoceros incurvus Steph., Hedwigia 32 (3): 143, 1893.

*** Folioceros indicus D.C.Bharadwaj, Geophytology 8 (1): 114, 1978.

*** Folioceros kashyapii S.C.Srivast. et A.K.Asthana, Bryologist 92 (2): 219, 1989.

*** Folioceros mangaloreus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975. Bas.: Aspiromitus mangaloreus Steph., Sp. Hepat. (Stephani) 5: 967, 1916.

*** Folioceros paliformis D.K.Singh, Bull. Bot. Surv. India 29: 176, 1987 [1989].

*** Folioceros physocladus D.C.Bharadwaj, Geophytology 8 (1): 115, 1978. Based on: Anthoceros physocladus Schiffn. ex Pandé, Proc. Indian Sci. Congr. Assoc. 47 (2): 98, 1960, nom. inval.

*** Folioceros pinnilobus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975. Bas.: Anthoceros pinnilobus Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 299, 1907.

*** Folioceros satpurensis D.C.Bharadwaj et K.P.Srivast., Geophytology 8 (1): 112, 1978. Based on: Anthoceros satpurensis K.P.Srivast., Proc. Indian Sci. Congr. Assoc. 47 (3): 337, 1960, nom. inval.

*** Folioceros udarii A.K.Asthana et S.C.Srivast., Cryptog. Bryol. Lichénol. 7 (2): 151, 1986.

*** Folioceros verruculosus (J.Haseg.) R.L.Zhu et M.J.Lai, Ann. Bot. Fenn. 48 (5): 383, 2011. Bas.: Anthoceros verruculosus J.Haseg., Acta Phytotax. Geobot. 44 (2): 103, 1993.

*** Dendrocerotaceae J.Haseg.

by J.C. Villarreal and D.C. Cargill

*** Dendroceros Nees, Syn. Hepat. 4: 579, 1846.

** subg. Apoceros R.M.Schust., Phytologia 63 (3): 200, 1987.

*** Dendroceros cavernosus J.Haseg., J. Hattori Bot. Lab. 47: 306, 1980.

*** Dendroceros difficilis Steph., Sp. Hepat. (Stephani) 5: 1009, 1917.

*** Dendroceros muelleri Steph., Hedwigia 28 (2): 133, 1889.

*** Dendroceros ogeramnangus Piippo, Acta Bot. Fenn. 148: 40, 1993.

*** Dendroceros pedunculatus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 14, 1909.

** Dendroceros subdifficilis S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 119, 1951.

** subg. Dendroceros

*** Dendroceros acutilobus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 18, 1909.

*** Dendroceros borbonicus Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 31, 1893 [1894].

*** Dendroceros crispus (Sw.) Nees, Syn. Hepat. 4: 581, 1846. Bas.: Anthoceros crispus Sw., Prodr. (Swartz): 146, 1788.

*** Dendroceros foliicola J.Haseg., J. Hattori Bot. Lab. 47: 296, 1980.

*** Dendroceros japonicus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 15, 1909.

*** Dendroceros javanicus (Nees) Nees, Syn. Hepat. 4: 582, 1846. Bas.: Anthoceros javanicus Nees, Enum. Pl. Crypt. Javae: 1, 1830.

*** Dendroceros subplanus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 20, 1909.

*** Dendroceros tubercularis S.Hatt., Bot. Mag. (Tokyo) 58 (685): 6, 1944.

*** Dendroceros validus Steph., Sp. Hepat. (Stephani) 5: 1016, 1917.

** Dendroceros vesconianus Gottsche, J. Bot. (Morot) 12: 150, 1898.

** Dendroceros wattsianus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 17, 1909.

Incertae sedis

* Dendroceros adglutinatus (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 580, 1846. Bas.: Monoclea adglutinata Hook.f. et Taylor, London J. Bot. 4: 96, 1845. 15

*** Dendroceros africanus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 18, 1909.

** Dendroceros allionii Steph., Sp. Hepat. (Stephani) 5: 1014, 1917.

** Dendroceros australis Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 17, 1909.

** Dendroceros breutelii Nees, Syn. Hepat. 4: 581, 1846.

** Dendroceros breutelii var. surinamensis Lindenb. et Gottsche, Linnaea 24 (6): 639, 1851 [1852].

*** Dendroceros cichoraceus (Mont.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 16, 1858. Bas.: Anthoceros cichoraceus Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 355, 1845.

* Dendroceros crassicostatus Steph., Sp. Hepat. (Stephani) 5: 1015, 1917.

** Dendroceros crassinervis (Nees) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 18, 1858. Bas.: Anthoceros crassinervis Nees, Syn. Hepat. 4: 589, 1846.

*** Dendroceros crispatus (Hook.) Nees, Syn. Hepat. 4: 579, 1846. Bas.: Monoclea crispata Hook., Bot. Misc. 1: 117, 1830.

** Dendroceros crispatus var. simplicior Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 574, 1885.

** Dendroceros cucullatus Steph., Sp. Hepat. (Stephani) 6: 429, 1923. 16

* Dendroceros exalatus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 14, 1909. 17

* Dendroceros gracilis Steph., Sp. Hepat. (Stephani) 5: 1015, 1917.

*** Dendroceros granulatus Mitt., Fl. vit.: 419, 1871 [1873].

** Dendroceros herasii M.Infante, J. Bryol. 32 (4): 285, 2010.

** Dendroceros humboldtensis Hürl., Bauhinia 1 (3): 254, 1960.

** Dendroceros paivae C.A.Garcia, Sérgio et J.C.Villarreal, Cryptog. Bryol. 33 (1): 5, 2012.

* Dendroceros rarus Steph., Sp. Hepat. (Stephani) 5: 1014, 1917.

* Dendroceros reticulus Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 37, 1950 [1951]. 18

** Dendroceros rigidus Steph., Sp. Hepat. (Stephani) 5: 1017, 1917.

*** Dendroceros seramensis J.Haseg., Acta Phytotax. Geobot. 37 (1/3): 10, 1986.

** Dendroceros subtropicus C.J.Wild, Trans. Nat. Hist. Soc. Queensland 1: 49, 1893.

** Dendroceros tahitensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 138, 1873.

*** Megaceros Campb., Ann. Bot. (Oxford) 21 (4): 484, 1907.

** Megaceros aneuriformis Steph., Sp. Hepat. (Stephani) 5: 949, 1916.

*** Megaceros austronesophilus Cargill et Seppelt, Austral. Syst. Bot. 26 (5): 372, 2013.

** Megaceros ciliatus K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 74, 1928.

*** Megaceros denticulatus (Lehm.) Steph., Sp. Hepat. (Stephani) 5: 956, 1916. Bas.: Anthoceros denticulatus Lehm., Nov. Stirp. Pug. 10: 25, 1857.

*** Megaceros flagellaris (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 951, 1916. Bas.: Anthoceros flagellaris Mitt., Fl. vit.: 419, 1871 [1873].

*** Megaceros gracilis (Reichardt) Steph., Sp. Hepat. (Stephani) 5: 955, 1916. Bas.: Anthoceros gracilis Reichardt, Verh. K.K. Zool.-Bot. Ges. Wien 16: 957, 1866.

** Megaceros leptohymenius (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 955, 1916. Bas.: Monoclea leptohymenia Hook.f. et Taylor, London J. Bot. 3: 575, 1844.

*** Megaceros pellucidus (Colenso) E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 115, 1972. Bas.: Anthoceros pellucidus Colenso, Trans. & Proc. New Zealand Inst. 17: 263, 1885.

** Megaceros tjibodensis Campb., Ann. Bot. (Oxford) 21 (4): 484, 1907.

Excluded from the genus 19

* Megaceros flavens (Spruce) Campb., Ann. Bot. (Oxford) 21 (4): 483, 1907. Bas.: Anthoceros flavens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 575, 1885.

* Megaceros jamesonii (Taylor) Steph., Biblioth. Bot. 87 (2): 268, 1916. Bas.: Dendroceros jamesonii Taylor, London J. Bot. 7: 285, 1848.

*** Nothoceros (R.M.Schust.) J.Haseg., J. Hattori Bot. Lab. 76: 32, 1994. Bas.: Megaceros subg. Nothoceros R.M.Schust., Phytologia 63 (3): 200, 1987.

*** Nothoceros aenigmaticus J.C.Villarreal et K.D.McFarland, Bryologist 113 (1): 109, 2010. Based on: Megaceros aenigmaticus R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 830, 1992, nom. inval.

*** Nothoceros canaliculatus (Pagán) J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007. Bas.: Dendroceros canaliculatus Pagán, Bryologist 45 (4): 111, 1942.

*** Nothoceros endiviifolius (Mont.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007. Bas.: Anthoceros endiviifolius Mont., Voy. Pole Sud, Bot. 1: 211, 1845.

** Nothoceros fuegiensis (Steph.) J.C.Villarreal, Bryologist 113 (1): 109, 2010. Bas.: Megaceros fuegiensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 91, 1911.

*** Nothoceros giganteus (Lehm. et Lindenb.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007. Bas.: Anthoceros giganteus Lehm. et Lindenb., Nov. Stirp. Pug. 4: 25, 1832.

*** Nothoceros minarum (Nees) J.C.Villarreal, Molec. Phylogen. Evol. 78: 34, 2014. Bas.: Anthoceros minarum Nees, Naturgesch. Eur. Leberm. 4: 340, 1838.

** Nothoceros renzagliensis J.C.Villarreal, L.V.Campos et Uribe, Syst. Bot. 37 (1): 32, 2012.

*** Nothoceros schizophyllus (Steph.) J.C.Villarreal, Molec. Phylogen. Evol. 78: 34, 2014. Bas.: Megaceros schizophyllus Steph., Sp. Hepat. (Stephani) 5: 949, 1916.

*** Nothoceros superbus J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 280, 2007.

*** Nothoceros vincentianus (Lehm. et Lindenb.) J.C.Villarreal, Bryologist 113 (1): 111, 2010. Bas.: Anthoceros vincentianus Lehm. et Lindenb., Nov. Stirp. Pug. 6: 16, 1834.

** Phaeomegaceros R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007.

*** Phaeomegaceros coriaceus (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007. Bas.: Anthoceros coriaceus Steph., Sp. Hepat. (Stephani) 5: 991, 1916.

*** Phaeomegaceros fimbriatus (Gottsche) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007. Bas.: Anthoceros fimbriatus Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 187, 1864.

*** Phaeomegaceros foveatus (J.Haseg.) J.C.Villarreal, Nova Hedwigia 91 (3/4): 352, 2010. Bas.: Phaeoceros foveatus J.Haseg., Bryol. Res. 7 (12): 374, 2001.

*** Phaeomegaceros hirticalyx (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007. Bas.: Aspiromitus hirticalyx Steph., Sp. Hepat. (Stephani) 5: 966, 1916.

*** Phaeomegaceros plicatus (Mitt.) J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 85, 2013. Bas.: Anthoceros plicatus Mitt., Rep. Challenger, Bot. 1 (2): 178, 1884.

*** Phaeomegaceros skottsbergii (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007. Bas.: Anthoceros skottsbergii Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 90, 1911.

*** Phaeomegaceros squamuliger (Spruce) J.C.Villarreal, Nova Hedwigia 91 (3/4): 351, 2010. Bas.: Anthoceros squamuliger Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 576, 1885.

** Phaeomegaceros squamuliger subsp. hasselii J.C.Villarreal, Cargill et Goffinet, Nova Hedwigia 91 (3/4): 352, 2010.

*** Phymatocerotaceae R.J.Duff, J.C.Villarreal, Cargill et Renzaglia

by J.C. Villarreal and D.C. Cargill

*** Phymatoceros Stotler, W.T.Doyle et Crand.-Stotl., Phytologia 87 (2): 113, 2005.

*** Phymatoceros bulbiculosus (Brot.) Stotler, W.T.Doyle et Crand.-Stotl., Phytologia 87 (2): 114, 2005. Bas.: Anthoceros bulbiculosus Brot., Fl. lusit. 2: 430, 1804 [1805].

*** Phymatoceros phymatodes (M.Howe) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 240, 2007. Bas.: Anthoceros phymatodes M.Howe, Bull. Torrey Bot. Club 25 (1): 12, 1898.

*** Notothyladaceae Müll.Frib. ex Prosk.

by J.C. Villarreal and D.C. Cargill

** Notothyladoideae Grolle

*** Notothylas Sull. ex A.Gray, Amer. J. Sci. Arts (ser. 2) 1 (1): 74, 1846.

** subg. Notothylas

*** Notothylas anaporata Udar et D.K.Singh, Rev. Bryol. Lichénol. 45 (2): 202, 1979.

*** Notothylas breutelii (Gottsche) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 21, 1858. Bas.: Anthoceros breutelii Gottsche, Syn. Hepat. 4: 583, 1846.

*** Notothylas depressispora J.Haseg., Acta Phytotax. Geobot. 30 (1/3): 26, 1979.

*** Notothylas dissecta Steph., Sp. Hepat. (Stephani) 5: 1020, 1917.

*** Notothylas himalayensis Udar et D.K.Singh, J. Bryol. 11 (3): 451, 1981.

*** Notothylas indica Kashyap, Proc. Lahore Philos. Soc. 4: 54, 1925.

*** Notothylas javanica (Sande Lac.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 20, 1858. Bas.: Blasia javanica Sande Lac., Syn. hepat. jav.: 94, 1856 [1857].

*** Notothylas orbicularis (Schwein.) Sull., Amer. J. Sci. Arts (ser. 2) 1 (1): 75, 1846. Bas.: Targionia orbicularis Schwein., Spec. Fl. Amer. Crypt.: 23, 1821.

*** Notothylas pandei Udar et V.Chandra, Geophytology 7 (2): 142, 1977.

** subg. Notothyloides A.K.Asthana et S.C.Srivast., Bryophyt. Biblioth. 42: 106, 1991.

*** Notothylas khasiana Udar et D.K.Singh, J. Indian Bot. Soc. 60: 112, 1981.

*** Notothylas pfleidereri Udar et D.K.Singh, Lindbergia 5 (1): 28, 1979.

Incertae sedis

*** Notothylas decurva (Mitt.) Steph., Cat. Afr. Pl. (Hiern) 2 (2): 320, 1901. Bas.: Anthoceros decurvus Mitt., Trans. Linn. Soc. London 23 (1): 58, 1860.

*** Notothylas flabellata Steph., Cat. Afr. Pl. (Hiern) 2 (2): 320, 1901.

*** Notothylas galapagensis M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 203, 1934.

** Notothylas irregularis Chantanaorr., Acta Bot. Hung. 56 (3/4): 270, 2014.

*** Notothylas kashyapii D.K.Singh, Indian J. Forest. 23 (4): 386, 2000.

*** Notothylas nepalensis D.K.Singh, J. Bombay Nat. Hist. Soc. 84 (3): 650, 1987 [1988].

*** Notothylas temperata J.Haseg., Acta Phytotax. Geobot. 30 (1/3): 20, 1979.

*** Notothylas udarii D.K.Singh et Semwal, Phytotaxonomy 1: 35, 2001.

* Notothylas verdoornii Khanna, Rev. Bryol. Lichénol. 6: 118, 1933.

*** Notothylas vitalii Udar et D.K.Singh, Misc. Bryol. Lichenol. 8 (9): 173, 1980.

** Notothylas yunnanensis T.Peng et R.L.Zhu, Phytotaxa 156 (3): 157, 2014.

** Phaeocerotoideae Hässel

* Mesoceros Piippo, Acta Bot. Fenn. 148: 30, 1993. 20

* Mesoceros mesophoros Piippo, Acta Bot. Fenn. 148: 30, 1993.

* Mesoceros porcatus Piippo, Haussknechtia, Beih. 9: 279, 1999.

** Paraphymatoceros Hässel, Phytologia 88 (2): 208, 2006.

*** Paraphymatoceros diadematus Hässel, Phytologia 88 (2): 209, 2006.

*** Paraphymatoceros hallii (Austin) Hässel, Phytologia 88 (2): 209, 2006. Bas.: Anthoceros hallii Austin, Bull. Torrey Bot. Club 6 (4): 26, 1875 [1876].

*** Paraphymatoceros pearsonii (M.Howe) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 94, 2015. Bas.: Anthoceros pearsonii M.Howe, Bull. Torrey Bot. Club 25 (1): 8, 1898.

*** Paraphymatoceros proskaueri (Stotler, Crand.-Stotl. et W.T.Doyle) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 94, 2015. Bas.: Phaeoceros proskaueri Stotler, Crand.-Stotl. et W.T.Doyle, Fieldiana, Bot. (n.ser.) 47: 232, 2008.

*** Phaeoceros Prosk., Bull. Torrey Bot. Club 78 (4): 346, 1951.

** Phaeoceros austroandinus Hässel, Candollea 44 (2): 721, 1989.

** Phaeoceros bolusii (Sim) S.W.Arnell, Hepat. South Africa: 401, 1963. Bas.: Anthoceros bolusii Sim, Trans. Roy. Soc. South Africa 15 (1): 114, 1926.

** Phaeoceros brevicapsulus (Steph.) Hässel, Candollea 44 (2): 725, 1989. Bas.: Anthoceros brevicapsulus Steph., Sp. Hepat. (Stephani) 5: 981, 1916.

*** Phaeoceros carolinianus (Michx.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951. Bas.: Anthoceros carolinianus Michx., Fl. bor.-amer. (Michaux) 2: 280, 1803.

*** Phaeoceros delicatus E.O.Campb. et Outred, New Zealand J. Bot. 33 (3): 285, 1995.

*** Phaeoceros dendroceroides (Steph.) Hässel, Beih. Nova Hedwigia 134: 487, 2009. Bas.: Anthoceros dendroceroides Steph., Sp. Hepat. (Stephani) 5: 984, 1916. 21

*** Phaeoceros engelii Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 248, 2008.

** Phaeoceros erectus Udar et D.K.Singh, Geophytology 11 (2): 257, 1981. Nom. nov. pro Anthoceros erectus Steph., Sp. Hepat. (Stephani) 5: 984, 1916, nom. illeg.

*** Phaeoceros evanidus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 245, 2008. Bas.: Anthoceros evanidus Steph., Sp. Hepat. (Stephani) 5: 990, 1916.

*** Phaeoceros exiguus (Steph.) J.Haseg., J. Hattori Bot. Lab. 60: 387, 1986. Bas.: Anthoceros exiguus Steph., Sp. Hepat. (Stephani) 5: 988, 1916.

*** Phaeoceros flexivalvis (Gottsche et Nees) Hässel, Candollea 44 (2): 728, 1989. Bas.: Anthoceros flexivalvis Gottsche et Nees, Syn. Hepat. 4: 586, 1846.

*** Phaeoceros fulvisporus (Steph.) J.Haseg., Trop. Bryol. 8: 52, 1993. Bas.: Anthoceros fulvisporus Steph., 52 (5): 306, 1912.

*** Phaeoceros gemmifer (Horik.) J.Haseg., J. Hattori Bot. Lab. 57: 252, 1984. Bas.: Anthoceros gemmifer Horik., Sci. Rep. Tôhoku Imp. Univ., Ser. 4, Biol. 4 (2): 426, 1929.

** Phaeoceros gualaquizanus (Steph.) Gradst., J. Hattori Bot. Lab. 45: 123, 1979. Bas.: Anthoceros gualaquizanus Steph., Sp. Hepat. (Stephani) 5: 979, 1916.

*** Phaeoceros himalayensis (Kashyap) Prosk. ex Bapna et G.G.Vyas, J. Hattori Bot. Lab. 25: 88, 1962. Bas.: Anthoceros himalayensis Kashyap, New Phytol. 14 (1): 8, 1915.

*** Phaeoceros huebschmannii Hässel, Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 301, 1986.

*** Phaeoceros inflatus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 246, 2008. Bas.: Anthoceros inflatus Steph., Sp. Hepat. (Stephani) 5: 990, 1916.

*** Phaeoceros kashyapii A.K.Asthana et S.C.Srivast., Bryophyt. Biblioth. 42: 129, 1991.

*** Phaeoceros laevis (L.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951. Bas.: Anthoceros laevis L., Sp. Pl. 1: 1139, 1753.

** Phaeoceros maranguensis (Steph.) Bapna, Univ. Udaipur Res. Stud. 3: 138, 1966. Bas.: Aspiromitus maranguensis Steph., Sp. Hepat. (Stephani) 5: 960, 1916.

* Phaeoceros microsporus (Steph.) Hässel, Candollea 44 (2): 721, 1989. Bas.: Aspiromitus microsporus Steph., Sp. Hepat. (Stephani) 5: 963, 1916. 22

*** Phaeoceros minutus (Mitt.) S.W.Arnell, Hepat. South Africa: 403, 1963. Bas.: Anthoceros minutus Mitt., J. Linn. Soc., Bot. 16 (91): 195, 1877.

*** Phaeoceros mohrii (Austin) Hässel, Candollea 44 (2): 721, 1989. Bas.: Anthoceros mohrii Austin, Bull. Torrey Bot. Club 6 (52): 304, 1879.

*** Phaeoceros oreganus (Austin) Hässel, Candollea 44 (2): 718, 1989. Bas.: Anthoceros oreganus Austin, Bull. Torrey Bot. Club 6 (4): 26, 1875 [1876].

*** Phaeoceros parvulus (Schiffn.) J.Haseg., J. Hattori Bot. Lab. 57: 248, 1984. Bas.: Anthoceros parvulus Schiffn., Österr. Bot. Z. 49 (11): 391, 1899.

*** Phaeoceros perpusillus Chantanaorr., Acta Bot. Hung. 51 (1/2): 30, 2009.

** Phaeoceros pichinchensis (Spruce) Hässel, Candollea 44 (2): 725, 1989. Bas.: Anthoceros pichinchensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 577, 1885.

** Phaeoceros propagulifer (Steph.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951. Bas.: Anthoceros propagulifer Steph., Sp. Hepat. (Stephani) 5: 1001, 1916.

* Phaeoceros striatisporus J.Haseg., J. Hattori Bot. Lab. 75, 268.23

*** Phaeoceros tenuis (Spruce) Hässel, Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 303, 1986. Bas.: Anthoceros tenuis Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcvi, 1889 [1890].

** Phaeoceros tigrinus (Gola) J.C.Villarreal, Phytotaxa 208 (1): 95, 2015. Bas.: Anthoceros tigrinus Gola, Ann. Bot. (Rome) 13 (1): 72, 1914.

*** Phaeoceros tuberosus (Taylor) Prosk., J. Indian Bot. Soc. 42A: 185, 1964. Bas.: Anthoceros tuberosus Taylor, London J. Bot. 5: 412, 1846.

*** Phaeoceros udarii A.K.Asthana et V.Nath, Proc. Natl. Acad. Sci. India, B 63 (4): 461, 1993.

** Phaeoceros wrightii (Steph.) Hässel, Candollea 44 (2): 722, 1989. Bas.: Anthoceros wrightii Steph., Sp. Hepat. (Stephani) 5: 999, 1916.

*** Leiosporocerotaceae Hässel ex Ochyra

by J.C. Villarreal and D.C. Cargill

*** Leiosporoceros Hässel, J. Bryol. 14 (2): 255, 1986.

*** Leiosporoceros dussii (Steph.) Hässel, J. Bryol. 14 (2): 255, 1986. Bas.: Anthoceros dussii Steph., Hedwigia 32 (3): 142, 1893.

*** Haplomitriaceae Dědeček

by S. Bartholomew-Began

The treatment of Haplomitriaceae follows. Morphogenetic and molecular studies support the infrageneric ranks of Haplomitrium.

*** Haplomitrium Nees, Naturgesch. Eur. Leberm. 1: 109, 1833 nom. conserv.

*** subg. Calobryum (Nees) R.M.Schust., Nova Hedwigia 13 (1/2): 40, 1967. Bas.: Calobryum Nees, Syn. Hepat. 4: 507, 1846.

*** Haplomitrium blumei (Nees) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963. Bas.: Monoclea blumei Nees, Enum. Pl. Crypt. Javae: 2, 1830.

*** Haplomitrium mnioides (Lindb.) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963. Bas.: Rhopalanthus mnioides Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 391, 1874.

*** subg. Haplomitrium

** sect. Archibryum (R.M.Schust.) J.J.Engel, Ann. Missouri Bot. Gard. 68 (4): 675, 1981. Bas.: Haplomitrium subg. Archibryum R.M.Schust., Nova Hedwigia 13 (1/2): 28, 1967.

*** Haplomitrium gibbsiae (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963. Bas.: Calobryum gibbsiae Steph., Sp. Hepat. (Stephani) 6: 76, 1917.

*** Haplomitrium intermedium Berrie, Proc. Linn. Soc. New South Wales (ser. 2) 87 (399): 191, 1963.

** sect. Haplomitrium

*** Haplomitrium hookeri (Lyell ex Sm.) Nees, Naturgesch. Eur. Leberm. 1: 111, 1833. Bas.: Jungermannia hookeri Lyell ex Sm., Engl. Bot. 35: tab. 2555, 1814.

*** Haplomitrium hookeri var. minutum (E.O.Campb.) Barthol.-Began, Bryophyt. Biblioth. 41: 230, 1991. Bas.: Steereomitrium minutum E.O.Campb., Mem. New York Bot. Gard. 45: 569, 1987.

*** Haplomitrium monoicum J.J.Engel, Ann. Missouri Bot. Gard. 68 (4): 668, 1981 [1982].

*** Haplomitrium ovalifolium R.M.Schust., Bryologist 74 (2): 136, 1971.

*** Treubiaceae Verd.

by R. Stotler and B.J. Crandall-Stotler

showed a clear molecular distinction between Treubia and Apotreubia, which is in accordance with their morphological differences and supports their recognition as separate genera.

*** Apotreubia S.Hatt. et Mizut., Bryologist 69 (4): 491, 1966 [1967].

*** Apotreubia hortoniae Konstant., Phytotaxa 76 (3): 33, 2013. Based on: Apotreubia hortoniae R.M.Schust. et Konstant., J. Hattori Bot. Lab. 78: 55, 1995, nom. inval.

*** Apotreubia nana (S.Hatt. et Inoue) S.Hatt. et Mizut., Bryologist 69 (4): 492, 1966 [1967]. Bas.: Treubia nana S.Hatt. et Inoue, J. Hattori Bot. Lab. 11: 99, 1954.

** Apotreubia pusilla (R.M.Schust.) Grolle, Acta Bot. Fenn. 125: 63, 1984. Bas.: Treubia pusilla R.M.Schust., Nova Hedwigia 15: 515, 1968.

*** Apotreubia yunnanensis Higuchi, Cryptog. Bryol. Lichénol. 19 (4): 321, 1998.

*** Treubia K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 1, 1890 [1891] nom. conserv.

*** Treubia insignis K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 1, 1890 [1891].

** Treubia insignis subsp. bracteata (Steph.) R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 241, 1969. Bas.: Treubia bracteata Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 302, 1896.

** Treubia insignis subsp. caledonica R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 243, 1969.

** Treubia insignis subsp. vitiensis R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 242, 1969.

*** Treubia lacunosa (Colenso) Prosk., Bryologist 58 (3): 199, 1955. Bas.: Noteroclada lacunosa Colenso, Trans. & Proc. New Zealand Inst. 18: 248, 1886.

** Treubia lacunosoides T.Pfeiff., W.Frey et M.Stech, Nova Hedwigia 75 (1/2): 249, 2002.

*** Treubia pygmaea R.M.Schust., Phytologia 56 (7): 460, 1985.

*** Treubia scapanioides R.M.Schust., J. Hattori Bot. Lab. 32: 246, 1969.

*** Treubia tahitensis (Nadeaud) Besch., J. Bot. (Morot) 12: 147, 1898. Bas.: Gottschea tahitensis Nadeaud, Énum. Pl. Tahiti: 7, 1873.

*** Treubia tasmanica R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 248, 1969.

*** Adelanthaceae Grolle

by K. Feldberg, J. Váňa and J. Heinrichs

The subfamily Jamesonielloideae was excluded from Lophoziaceae/Scapaniaceae by and this status was confirmed by e.g.. The family was described and defined by. Some taxonomic and nomenclatural notes can also be found in, ),.

*** Adelanthoideae K.Feldberg, Heinrichs et Váňa

*** Adelanthus Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 243, 1864 nom. conserv.

** sect. Adelanthus

*** Adelanthus falcatus (Hook.) Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 243, 1864. Bas.: Jungermannia falcata Hook., Musci Exot. 1: tab. 89, 1818.

*** Adelanthus occlusus (Hook.f. et Taylor) Carrington, Trans. Bot. Soc. Edinburgh 10: 381, 1870. Bas.: Jungermannia occlusa Hook.f. et Taylor, London J. Bot. 3: 369, 1844.

** sect. Calyptrocolea (R.M.Schust.) Grolle, J. Hattori Bot. Lab. 35: 331, 1972. Bas.: Calyptrocolea R.M.Schust., Rev. Bryol. Lichénol. 34 (3/4): 685, 1966.

*** Adelanthus aureomarginatus R.M.Schust., Phytologia 39 (4): 250, 1978.

*** Adelanthus gemmiparus (R.M.Schust.) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Biol. Sci. 11 (18): 241, 1970. Bas.: Calyptrocolea gemmipara R.M.Schust., Rev. Bryol. Lichénol. 34 (3/4): 695, 1966 [1967].

** Adelanthus lingulatus J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 26, 2013.

*** Adelanthus tenuis J.J.Engel et Grolle, J. Hattori Bot. Lab. 35: 333, 1972.

** sect. Lindenbergiani Grolle, J. Hattori Bot. Lab. 35: 331, 1972.

*** Adelanthus carabayensis (Mont.) Grolle, J. Hattori Bot. Lab. 35: 348, 1972. Bas.: Plagiochila carabayensis Mont., Ann. Sci. Nat. Bot. (sér. 4) 5: 348, 1856.

*** Adelanthus integerrimus Grolle, J. Hattori Bot. Lab. 35: 340, 1972.

*** Adelanthus lindenbergianus (Lehm.) Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 244, 1864. Bas.: Jungermannia lindenbergiana Lehm., Linnaea 4: 367, 1829.

** sect. Pittieri Grolle, J. Hattori Bot. Lab. 35: 331, 1972.

*** Adelanthus pittieri (Steph.) Grolle, J. Hattori Bot. Lab. 35: 337, 1972. Bas.: Tylimanthus pittieri Steph., Sp. Hepat. (Stephani) 6: 250, 1922.

*** Adelanthus squarrosus Grolle, J. Hattori Bot. Lab. 67: 243, 1989.

*** Pseudomarsupidium Herzog, Svensk Bot. Tidskr. 47 (1): 42, 1953.

*** Pseudomarsupidium aureocinctum (R.M.Schust.) J.J.Engel, Novon 17 (3): 312, 2007. Bas.: Adelanthus decipiens subsp. aureocinctus R.M.Schust., Phytologia 39 (4): 250, 1978.

*** Pseudomarsupidium borneensis (Grolle) Váňa, L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 60, 2012. Bas.: Adelanthus borneensis Grolle, J. Hattori Bot. Lab. 35: 362, 1972.

*** Pseudomarsupidium decipiens (Hook.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 443, 1963. Bas.: Jungermannia decipiens Hook., Brit. Jungermann.: tab. 50, 1813.

*** Pseudomarsupidium piliferum (Steph.) Herzog ex Grolle, Trans. Brit. Bryol. Soc. 4 (3): 443, 1963. Bas.: Marsupidium piliferum Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 602 (386), 1908.

*** Wettsteinia Schiffn., Ann. Jard. Bot. Buitenzorg, suppl. 2: 44, 1898.

*** Wettsteinia densiretis (Herzog) Grolle, J. Hattori Bot. Lab. 28: 99, 1965. Bas.: Tylimanthus densiretis Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 712, 1942.

*** Wettsteinia inversa (Sande Lac.) Schiffn., Ann. Jard. Bot. Buitenzorg, suppl. 2: 45, 1898. Bas.: Plagiochila inversa Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 289, 1864.

*** Wettsteinia rotundifolia (Horik.) Grolle, J. Hattori Bot. Lab. 28: 100, 1965. Bas.: Adelanthus rotundifolius Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 181, 1934.

*** Wettsteinia schusteriana Grolle, J. Hattori Bot. Lab. 28: 99, 1965.

** Jamesonielloideae Inoue

*** Cuspidatula Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1141 (124), 1901.

*** Cuspidatula contracta (Reinw., Blume et Nees) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1141 (124), 1901. Bas.: Jungermannia contracta Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 233, 1824 [1825].

*** Cuspidatula flaccida (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 142, 2010. Bas.: Anastrophyllum flaccidum Steph., Sp. Hepat. (Stephani) 6: 105, 1917.

*** Cuspidatula flexicaulis (Nees) Váňa et L.Söderstr., Phytotaxa 76 (3): 35, 2013. Bas.: Jungermannia flexicaulis Nees, Linnaea 6 (4): 604, 1831.

*** Cuspidatula kirkii (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 142, 2010. Bas.: Jamesoniella kirkii Steph., Hedwigia 34 (2): 47, 1895.

*** Cuspidatula monodon (Taylor) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1143 (126), 1901. Bas.: Jungermannia monodon Taylor, Nov. Stirp. Pug. 8: 7, 1844.

*** Cuspidatula orbicularis (Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 36, 2013. Bas.: Jamesoniella orbicularis Grolle, Feddes Repert. 82 (1): 42, 1971.

*** Cuspidatula robusta (Austin) Váňa et L.Söderstr., Phytotaxa 76 (3): 36, 2013. Bas.: Jungermannia robusta Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 219, 1869.

** Denotarisia Grolle, Feddes Repert. 82 (1): 6, 1971.

*** Denotarisia linguifolia (De Not.) Grolle, Feddes Repert. 82 (1): 6, 1971. Bas.: Plagiochila linguifolia De Not., Epat. Borneo: 13, 1874.

** Nothostrepta R.M.Schust., Phytologia 45 (5): 420, 1980.

*** Nothostrepta bifida (Steph.) R.M.Schust., Phytologia 45 (5): 420, 1980. Bas.: Plagiochila bifida Steph., Annuario Reale Ist. Bot. Roma 2: 86, 1885 [1886].

*** Nothostrepta longissima (Steph.) R.M.Schust., Phytologia 45 (5): 420, 1980. Bas.: Anastrophyllum longissimum Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 13, 1901.

** Pisanoa Hässel, Lindbergia 14 (3): 179, 1988 [1989].

*** Pisanoa chilensis Hässel, Lindbergia 14 (3): 179, 1988 [1989].

** Protosyzygiella (Inoue) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 334, 1980. Bas.: Syzygiella subg. Protosyzygiella Inoue, J. Hattori Bot. Lab. 29: 180, 1966.

*** Protosyzygiella pseudoconnexa (Herzog) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 334, 1980. Bas.: Plagiochila pseudoconnexa Herzog, Rev. Bryol. Lichénol. 21 (3/4): 259, 1952 [1953].

*** Syzygiella Spruce, J. Bot. 14: 234, 1876. 24

*** subg. Anomalae (Inoue) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010. Bas.: Syzygiella sect. Anomalae Inoue, J. Hattori Bot. Lab. 29: 183, 1966.

*** Syzygiella anomala (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 471 (190), 1902. Bas.: Plagiochila anomala Lindenb. et Gottsche, Syn. Hepat. 5: 646, 1847.

*** Syzygiella bilobata Inoue, J. Hattori Bot. Lab. 29: 186, 1966.

** Syzygiella ciliata Gradst. et A.R.Benitez, Nova Hedwigia 99 (1/2): 115, 2014.

*** Syzygiella concreta (Gottsche) Spruce, J. Bot. 14: 234, 1876. Bas.: Jungermannia concreta Gottsche, Mexik. Leverm.: 82, 1863.

*** Syzygiella manca (Mont.) Steph., Hedwigia 31 (1): 14, 1892. Bas.: Chiloscyphus mancus Mont., Syll. Gen. Sp. Crypt.: 63, 1856.

*** Syzygiella pectiniformis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 501, 1885.

*** Syzygiella tonduzana Steph., Sp. Hepat. (Stephani) 6: 118, 1917.

*** Syzygiella trigonifolia (Steph.) Herzog, Hedwigia 74 (2): 87, 1934. Bas.: Jamesoniella trigonifolia Steph., Biblioth. Bot. 87 (2): 185, 1916.

*** subg. Cryptochila (R.M.Schust.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010. Bas.: Cryptochila R.M.Schust., J. Hattori Bot. Lab. 26: 284, 1963.

*** Syzygiella acinacifolia (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010. Bas.: Jungermannia acinacifolia Hook.f. et Taylor, London J. Bot. 3: 367, 1844.

*** Syzygiella nigrescens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010. Bas.: Jamesoniella nigrescens Steph., Hedwigia 34 (2): 48, 1895.

*** Syzygiella paludosa (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010. Bas.: Jamesoniella paludosa Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 11, 1901.

*** Syzygiella pseudocclusa (E.A.Hodgs.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010. Bas.: Jamesoniella pseudocclusa E.A.Hodgs., Trans. Roy. Soc. New Zealand 85 (4): 583, 1958.

*** Syzygiella sonderi (Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010. Bas.: Jungermannia sonderi Gottsche, Linnaea 28 (5): 550, 1856 [1857].

*** Syzygiella spegazziniana (Spruce ex C.Massal.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010. Bas.: Jungermannia spegazziniana Spruce ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 216, 1885.

*** subg. Pseudoplagiochila Inoue, J. Hattori Bot. Lab. 29: 182, 1966.

*** Syzygiella ovalifolia Inoue, J. Hattori Bot. Lab. 29: 191, 1966.

* Syzygiella securifolia (Nees) Inoue, J. Hattori Bot. Lab. 46: 232, 1979. Bas.: Plagiochila securifolia Nees, Sp. Hepat. (Lindenberg) 2-4: 58, 1840. 25

*** Syzygiella subintegerrima (Reinw., Blume et Nees) Spruce, J. Linn. Soc., Bot. 30 (210): 362, 1895. Bas.: Jungermannia subintegerrima Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 238, 1824 [1825].

*** Syzygiella tasmanica (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010. Bas.: Jungermannia tasmanica Hook.f. et Taylor, London J. Bot. 5: 274, 1846.

*** subg. Roivainenia (Perss.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010. Bas.: Roivainenia Perss., Nova Hedwigia 3 (1): 43, 1961.

*** Syzygiella jacquinotii (Mont.) Hentschel, K.Feldberg, Váňa et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010. Bas.: Jungermannia jacquinotii Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 250, 1843.

*** subg. Syzygiella

*** Syzygiella autumnalis (DC.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010. Bas.: Jungermannia autumnalis DC., Fl. Franç. (DC. & Lamarck), 5 (6): 202, 1815.

*** Syzygiella campanulata Herzog, Rev. Bryol. Lichénol. 11 (1): 9, 1938 [1939].

*** Syzygiella colorata (Lehm.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010. Bas.: Jungermannia colorata Lehm., Linnaea 4: 366, 1829.

** Syzygiella colorata var. collenchymata J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 102, 2013.

*** Syzygiella contigua Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 470 (189), 1902. Nom. nov. pro Jungermannia contigua Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 118, 1864, nom. illeg.

*** Syzygiella elongella (Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010. Bas.: Jungermannia elongella Taylor, London J. Bot. 5: 274, 1846.

*** Syzygiella macrocalyx (Mont.) Spruce, J. Bot. 14: 234, 1876. Bas.: Jungermannia macrocalyx Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 248, 1843.

*** Syzygiella nipponica (S.Hatt.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010. Bas.: Jamesoniella nipponica S.Hatt., J. Jap. Bot. 19 (11): 350, 1943.

** Syzygiella oenops (Lindenb. et Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010. Bas.: Jungermannia oenops Lindenb. et Gottsche, Syn. Hepat. 5: 673, 1847.

*** Syzygiella perfoliata (Sw.) Spruce, J. Bot. 14: 234, 1876. Bas.: Jungermannia perfoliata Sw., Prodr. (Swartz): 143, 1788.

*** Syzygiella purpurascens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010. Bas.: Jamesoniella purpurascens Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 200, 1891 [1892].

*** Syzygiella rubricaulis (Nees) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 468 (187), 1902. Bas.: Jungermannia rubricaulis Nees, Fl. Bras. (Martius) 1 (1): 344, 1833.

*** Syzygiella setulosa Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 469 (188), 1902.

*** Syzygiella teres (Carrington et Pearson) Váňa, Phytotaxa 76 (3): 35, 2013. Bas.: Jungermannia teres Carrington et Pearson, Pap. & Proc. Roy. Soc. Tasmania 1887: 9, 1888.

*** Syzygiella undata (Mont.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010. Bas.: Jungermannia undata Mont., Ann. Sci. Nat. Bot. (sér. 4) 14: 183, 1860.

Incertae sedis

*** Syzygiella eatonii (Austin) Inoue, J. Jap. Bot. 37 (12): 359, 1962. Bas.: Plagiochila eatonii Austin, Trans. Connecticut Acad. Arts 8 (15): 257, 1891.

*** Syzygiella uleana Steph., Hedwigia 44 (4): 224, 1905.

** Vanaea (Inoue et Gradst.) Inoue et Gradst., Trop. Bryol. 1: 33, 1989. Bas.: Anastrophyllum subg. Vanaea Inoue et Gradst., Bull. Natl. Sci. Mus. Tokyo, B 14 (3): 88, 1988.

*** Vanaea plagiochiloides (Inoue et Gradst.) Inoue et Gradst., Trop. Bryol. 1: 33, 1989. Bas.: Anastrophyllum plagiochiloides Inoue et Gradst., Bull. Natl. Sci. Mus. Tokyo, B 14 (3): 88, 1988.

*** Anastrophyllaceae L.Söderstr., De Roo et Hedd.

by J. Váňa and L. Söderström

Anastrophyllaceae was described by from elements usually included in Lophoziaceae. Further taxonomic and nomenclatural notes can be found in, ). The complex of Chandonanthus/Plicanthus/Tetralophozia should be checked using molecular methods before generic and specific status of most of the species can be confirmed. The placement of Hattoria and Zantenia in the family is still provisional. The placement of Isopaches is also unclear, placed it in the family, but the study of showed that it can not be placed there.

*** Anastrepta (Lindb.) Schiffn., Hepat. (Engl.-Prantl): 85, 1893. Bas.: Jungermannia sect. Anastrepta Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 40, 1889.

*** Anastrepta orcadensis (Hook.) Schiffn., Hepat. (Engl.-Prantl): 85, 1893. Bas.: Jungermannia orcadensis Hook., Brit. Jungermann.: tab. 71, 1815.

*** Anastrophyllum (Spruce) Steph., Hedwigia 32 (3): 139, 1893. Bas.: Jungermannia subg. Anastrophyllum Spruce, J. Bot. 14: 235, 1876.

*** Anastrophyllum alpinum Steph., Sp. Hepat. (Stephani) 6: 103, 1917.

*** Anastrophyllum assimile (Mitt.) Steph., Hedwigia 32 (3): 140, 1893. Bas.: Jungermannia assimilis Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 93, 1860 [1861].

*** Anastrophyllum auritum (Lehm.) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1137 (120), 1901. Bas.: Jungermannia aurita Lehm., Linnaea 4: 368, 1829.

*** Anastrophyllum ciliatum Steph., Hedwigia 32 (3): 139, 1893.

*** Anastrophyllum donnianum (Hook.) Steph., Hedwigia 32 (3): 140, 1893. Bas.: Jungermannia donniana Hook., Brit. Jungermann.: tab. 39, 1813.

*** Anastrophyllum ellipticum Inoue, Bull. Natl. Sci. Mus. Tokyo, B 4 (1): 13, 1978. 26

** Anastrophyllum esenbeckii (Mont.) Steph., Hedwigia 32 (3): 140, 1893. Bas.: Jungermannia esenbeckii Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 247, 1843.

*** Anastrophyllum fissum Steph., Bull. Herb. Boissier 5 (10): 845, 1897.

*** Anastrophyllum joergensenii Schiffn., Hedwigia 49 (4): 396, 1910.

* Anastrophyllum lignicola D.B.Schill et D.G.Long, Ann. Bot. Fenn. 39 (2): 130, 2002. 27

*** Anastrophyllum michauxii (F.Weber) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 289, 1932 [1933]. Bas.: Jungermannia michauxii F.Weber, Hist. Musc. Hepat. Prodr.: 76, 1815.

*** Anastrophyllum nigrescens (Mitt.) Steph., Hedwigia 32 (3): 140, 1893. Bas.: Jungermannia nigrescens Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851.

*** Anastrophyllum obtusum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 285, 1950.

*** Anastrophyllum piligerum (Nees) Steph., Hedwigia 32 (3): 140, 1893. Bas.: Jungermannia piligera Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 414, 1824 [1825].

*** Anastrophyllum squarrosum Herzog, Ann. Bryol. 5: 72, 1932.

*** Anastrophyllum stellatum R.M.Schust., Phytologia 39 (4): 243, 1978.

*** Anastrophyllum tubulosum (Nees) Grolle, J. Hattori Bot. Lab. 28: 101, 1965. Bas.: Jungermannia tubulosa Nees, Enum. Pl. Crypt. Javae: 32, 1830.

*** Barbilophozia Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908.

** subg. Barbilophozia

*** Barbilophozia barbata (Schmidel ex Schreb.) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908. Bas.: Jungermannia barbata Schmidel ex Schreb., Spic. Fl. Lips.: 107, 1771.

*** Barbilophozia hatcheri (A.Evans) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908. Bas.: Jungermannia hatcheri A.Evans, Bull. Torrey Bot. Club 25 (8): 417, 1898.

*** Barbilophozia lycopodioides (Wallr.) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908. Bas.: Jungermannia lycopodioides Wallr., Comp. fl. Germ. 2 (III): 76, 1831.

*** Barbilophozia rubescens (R.M.Schust. et Damsh.) Kartt. et L.Söderstr., Ann. Bot. Fenn. 29 (2): 120, 1992. Bas.: Lophozia rubescens R.M.Schust. et Damsh., Phytologia 63 (5): 325, 1987.

** subg. Sudeticae (Schljakov) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010. Bas.: Lophozia sect. Sudeticae Schljakov, Pečen. Mchi Sev. SSSR 3: 113, 1980.

*** Barbilophozia sudetica (Nees ex Huebener) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010. Bas.: Jungermannia sudetica Nees ex Huebener, Hepaticol. germ.: 142, 1834. 28

*** Biantheridion (Grolle) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010]. Bas.: Jamesoniella sect. Biantheridion Grolle, Trans. Brit. Bryol. Soc. 4 (4): 662, 1964.

*** Biantheridion undulifolium (Nees) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010]. Bas.: Jungermannia schraderi β undulifolia Nees, Naturgesch. Eur. Leberm. 1: 306, 1833.

*** Chandonanthus Mitt., Handb. N. Zeal. fl. 2: 753, 1867.

*** Chandonanthus squarrosus (Menzies) Mitt., Handb. N. Zeal. fl. 2: 753, 1867. Bas.: Jungermannia squarrosa Menzies, Musci Exot. 1: tab. 78, 1818.

*** Crossocalyx Meyl., Bull. Soc. Vaud. Sci. Nat. 60 (249): 266, 1939.

*** Crossocalyx hellerianus (Nees ex Lindenb.) Meyl., Bull. Soc. Vaud. Sci. Nat. 60 (249): 266, 1939. Bas.: Jungermannia helleriana Nees ex Lindenb., Syn. hepat. eur: 64, 1829.

*** Crossocalyx tenuis (Harry Williams) Schljakov, Novosti Sist. Nizš. Rast. 15: 246, 1978. Bas.: Anastrophyllum tenue Harry Williams, Bryologist 71 (1): 34, 1968.

*** Gymnocolea (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835. Bas.: Jungermannia sect. Gymnocolea Dumort., Syll. Jungerm. Europ.: 52, 1831.

*** Gymnocolea borealis (Frisvoll et Moen) R.M.Schust., Lindbergia 12 (1): 7, 1986. Bas.: Lophozia borealis Frisvoll et Moen, Lindbergia 6 (2): 138, 1980 [1981].

** Gymnocolea fascinifera Potemkin, Arctoa 2: 76, 1993.

*** Gymnocolea inflata (Huds.) Dumort., Recueil Observ. Jungerm.: 17, 1835. Bas.: Jungermannia inflata Huds., Fl. Angl. (Hudson), ed. 2: 511, 1778.

* Gymnocolea inflata subsp. acutiloba (Schiffn.) R.M.Schust. et Damsh. ex L.Söderstr. et Váňa, Lindbergia 27 (1): 43, 2002. Bas.: Lophozia acutiloba Schiffn., Hedwigia 48 (3): 187, 1909. 29

*** Hamatostrepta Váňa et D.G.Long, Fieldiana, Bot. (n.ser.) 47: 134, 2008.

*** Hamatostrepta concinna Váňa et D.G.Long, Fieldiana, Bot. (n.ser.) 47: 134, 2008.

** Hattoria R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 69, 1961.

*** Hattoria yakushimensis (Horik.) R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 70, 1961. Bas.: Anastrophyllum yakushimense Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 149, 1934.

*** Isopaches H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 287, 1932 [1933].

*** Isopaches alboviridis (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 205, 1979. Bas.: Lophozia alboviridis R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 487, 1969.

*** Isopaches bicrenatus (Schmidel ex Hoffm.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 288, 1932 [1933]. Bas.: Jungermannia bicrenata Schmidel ex Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 11 (addenda), 1795 [1796].

*** Isopaches decolorans (Limpr.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 288, 1932 [1933]. Bas.: Jungermannia decolorans Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 57: 316, 1879 [1880].

*** Isopaches pumicicola (Berggr.) Bakalin, Arctoa 17: 162, 2008 [2009]. Bas.: Lophozia pumicicola Berggr., New Zealand Hepat.: 21, 1898.

*** Neoorthocaulis L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010.

*** Neoorthocaulis attenuatus (Mart.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010. Bas.: Jungermannia quinquedentata δ attenuata Mart., Fl. crypt. erlang.: 177, 1817.

*** Neoorthocaulis binsteadii (Kaal.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010. Bas.: Jungermannia binsteadii Kaal., Skr. Vidensk.-Selsk. Christiana, Math.-Naturvidensk. Kl. 1898 (9): 9, 1898.

*** Neoorthocaulis floerkei (F.Weber et D.Mohr) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010. Bas.: Jungermannia floerkei F.Weber et D.Mohr, Bot. Taschenb. (Weber): 410, 1807.

** Neoorthocaulis hyperboreus (R.M.Schust.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010. Bas.: Lophozia floerkei var. hyperborea R.M.Schust., Bull. Natl. Mus. Canada 164: 21, 1959.

*** Orthocaulis H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 293, 1932 [1933].

*** Orthocaulis atlanticus (Kaal.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 294, 1932 [1933]. Bas.: Jungermannia atlantica Kaal., Skr. Vidensk.-Selsk. Christiana, Math.-Naturvidensk. Kl. 1898 (9): 11, 1898.

* Orthocaulis cavifolius H.Buch et S.W.Arnell, Memoranda Soc. Fauna Fl. Fennica 26: 71, 1951.

** Plicanthus R.M.Schust., Nova Hedwigia 74 (3/4): 484, 2002.

*** Plicanthus birmensis (Steph.) R.M.Schust., Beih. Nova Hedwigia 119: 223, 2002. Bas.: Chandonanthus birmensis Steph., Bull. Soc. Roy. Bot. Belgique 38 (1): 43, 1899. 30

** Plicanthus difficilis (Steph.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 30, 2013. Bas.: Chandonanthus difficilis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 101, 1914.

** Plicanthus giganteus (Steph.) R.M.Schust., Nova Hedwigia 74 (3/4): 485, 2002. Bas.: Chandonanthus giganteus Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 124, 1911. 31

*** Plicanthus hirtellus (F.Weber) R.M.Schust., Nova Hedwigia 74 (3/4): 492, 2002. Bas.: Jungermannia hirtella F.Weber, Hist. Musc. Hepat. Prodr.: 50, 1815.

*** Schizophyllopsis Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013. Nom. nov. pro Anastrophyllum subg. Schizophyllum R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 739, 1969. 32

** Schizophyllopsis aristata (Herzog ex N.Kitag.) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013. Bas.: Anastrophyllum bidens var. aristatum Herzog ex N.Kitag., J. Hattori Bot. Lab. 33: 216, 1970.

*** Schizophyllopsis bidens (Reinw., Blume et Nees) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013. Bas.: Jungermannia bidens Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 208, 1824 [1825].

** Schizophyllopsis lanciloba (Steph.) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013. Bas.: Anastrophyllum lancilobum Steph., Sp. Hepat. (Stephani) 6: 107, 1917.

*** Schizophyllopsis papillosa (J.J.Engel et Braggins) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013. Bas.: Anastrophyllum papillosum J.J.Engel et Braggins, J. Bryol. 20 (2): 381, 1998.

*** Schizophyllopsis sphenoloboides (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 152 (1): 49, 2013. Bas.: Anastrophyllum sphenoloboides R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 741, 1969.

*** Schljakovia Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010].

*** Schljakovia kunzeana (Huebener) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010]. Bas.: Jungermannia kunzeana Huebener, Hepaticol. germ.: 115, 1834.

*** Schljakovianthus Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010].

*** Schljakovianthus quadrilobus (Lindb.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010]. Bas.: Jungermannia quadriloba Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 55, 1889.

*** Sphenolobopsis R.M.Schust. et N.Kitag., Nova Hedwigia 22: 152, 1971 [1972].

*** Sphenolobopsis pearsonii (Spruce) R.M.Schust., Nova Hedwigia 22: 153, 1971 [1972]. Bas.: Jungermannia pearsonii Spruce, J. Bot. 19: 33, 1881.

*** Sphenolobus (Lindb.) Berggr., New Zealand Hepat.: 22, 1898. Bas.: Jungermannia sect. Sphenolobus Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 369, 1874.

*** Sphenolobus austroamericanus (Váňa) Váňa, Phytotaxa 81 (1): 30, 2013. Bas.: Anastrophyllum austroamericanum Váňa, J. Hattori Bot. Lab. 48: 225, 1980.

*** Sphenolobus minutus (Schreb. ex D.Crantz) Berggr., New Zealand Hepat.: 22, 1898. Bas.: Jungermannia minuta Schreb. ex D.Crantz, Forts. Hist. Grönland: 285, 1770 (; non vidi).

*** Sphenolobus saxicola (Schrad.) Steph., Bull. Herb. Boissier (sér. 2) 2 (2): 168 (160), 1902. Bas.: Jungermannia saxicola Schrad., Syst. Samml. Crypt. Gew. 2: 4, 1797.

** Tetralophozia (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 13: 227, 1976. Bas.: Chandonanthus subg. Tetralophozia R.M.Schust., J. Hattori Bot. Lab. 23: 206, 1960 [1961].

*** Tetralophozia cavallii (Gola) Váňa, Trop. Bryol. 8: 102, 1993. Bas.: Blepharostomum cavallii Gola, Ann. Bot. (Rome) 6 (2): 274, 1907.

*** Tetralophozia filiformis (Steph.) Urmi, J. Bryol. 12 (3): 394, 1983. Bas.: Chandonanthus filiformis Steph., Sp. Hepat. (Stephani) 3: 644, 1909.

** Tetralophozia pilifera (Steph.) R.M.Schust., Nova Hedwigia 74 (3/4): 482, 2002. Bas.: Chandonanthus pilifer Steph., Sp. Hepat. (Stephani) 3: 644, 1909.

*** Tetralophozia setiformis (Ehrh.) Schljakov, Novosti Sist. Nizš. Rast. 13: 228, 1976. Bas.: Jungermannia setiformis Ehrh., Hannover. Mag. 22 (8): 142, 1784.

*** Zantenia (S.Hatt.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 29, 2013. Bas.: Anastrophyllum subg. Zantenia S.Hatt., Bot. Mag. (Tokyo) 79 (937): 342, 1966.

*** Zantenia borneensis (Herzog) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 29, 2013. Bas.: Anastrophyllum borneense Herzog, Trans. Brit. Bryol. Soc. 1 (4): 282, 1950.

*** Zantenia denticulata (Grolle) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013. Bas.: Anastrophyllum denticulatum Grolle, Nova Hedwigia 16: 148, 1968.

*** Zantenia karstenii (Schiffn.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013. Bas.: Anastrophyllum karstenii Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 268, 1893.

*** Zantenia prionophylla (S.Hatt.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013. Bas.: Anastrophyllum prionophyllum S.Hatt., Bot. Mag. (Tokyo) 79 (937): 342, 1966.

*** Cephaloziaceae Mig.

by J. Váňa with contributions by S.R. Gradstein (Odontoschisma)

Cephaloziaceae was recently studied by and. Some taxonomic and nomenclatural notes were published by, ) and.

** Alobielloideae R.M.Schust.

** Alobiella (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893. Bas.: Cephalozia subg. Alobiella Spruce, Cephalozia: 28, 1882.

*** Alobiella husnotii (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893. Bas.: Cephalozia husnotii Spruce, Cephalozia: 30, 1882.

** Alobiellopsis R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965.

*** Alobiellopsis acroscypha (Spruce) R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965. Bas.: Cephalozia acroscypha Spruce, Cephalozia: 30, 1882.

*** Alobiellopsis dominicensis (Spruce) Fulford, Mem. New York Bot. Gard. 11 (3): 350, 1968. Bas.: Alobiella dominicensis Spruce, J. Linn. Soc., Bot. 30 (210): 355, 1895.

*** Alobiellopsis heteromorpha (Lehm.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 682, 1969. Bas.: Jungermannia heteromorpha Lehm., Linnaea 4: 362, 1829.

*** Alobiellopsis parvifolia (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 679, 1969. Bas.: Alobiella parvifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 568 (352), 1908.

*** Alobiellopsis pillansii (Sim) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 683, 1969. Bas.: Cephalozia pillansii Sim, Trans. Roy. Soc. South Africa 15 (1): 87, 1926.

** Cephalozioideae Müll.Frib.

*** Cephalozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835. Bas.: Jungermannia sect. Cephalozia Dumort., Syll. Jungerm. Europ.: 60, 1831.

*** Cephalozia acuminata (Herzog) Váňa, Phytotaxa 112 (1): 11, 2013. Bas.: Hygrobiella acuminata Herzog, Trans. Brit. Bryol. Soc. 1 (4): 292, 1950.

** Cephalozia acutiloba (Inoue) Váňa, Phytotaxa 112 (1): 11, 2013. Bas.: Metahygrobiella acutiloba Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 10 (2): 155, 1967.

*** Cephalozia albula Steph., Hedwigia 32 (5): 318, 1893. Nom. nov. pro Jungermannia albula Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 93, 1860 [1861], nom. illeg.

*** Cephalozia ambigua C.Massal., Malpighia 21 (7/8): 310, 1907.

*** Cephalozia austrigena R.M.Schust. ex J.J.Engel, Novon 17 (3): 312, 2007. Based on: Cephalozia bicuspidata subsp. austrigena R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 712, 1974, nom. inval.

*** Cephalozia badia (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 483 (313), 1908. Bas.: Jungermannia badia Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 452, 1890.

*** Cephalozia bicuspidata (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835. Bas.: Jungermannia bicuspidata L., Sp. Pl. 1: 1132, 1753.

* Cephalozia bicuspidata subsp. lammersiana (Huebener) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 730, 1974. Bas.: Jungermannia lammersiana Huebener, Flora 15 (20): 306, 1832.

*** Cephalozia chilensis (J.J.Engel et R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 119: 29, 2002. Bas.: Metahygrobiella chilensis J.J.Engel et R.M.Schust., Brittonia 40 (2): 203, 1988.

*** Cephalozia conchata (Grolle et Váňa) Váňa, Syst. Bot. 40 (1): 38, 2015. Bas.: Jungermannia conchata Grolle et Váňa, Fragm. Florist. Geobot. 37 (1): 3, 1992.

*** Cephalozia crossii Spruce, Cephalozia: 46, 1882.

*** Cephalozia darjeelingensis Udar et D.Kumar, Geophytology 6 (1): 36, 1976.

*** Cephalozia drucei (R.M.Schust.) Váňa, Phytotaxa 112 (1): 11, 2013. Bas.: Metahygrobiella drucei R.M.Schust., J. Hattori Bot. Lab. 26: 273, 1963.

*** Cephalozia fuegiensis Váňa, Phytotaxa 112 (1): 12, 2013. Nom. nov. pro Hygrobiella dusenii Steph., Sp. Hepat. (Stephani) 6: 444, 1924.

*** Cephalozia hamatiloba Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 427 (303), 1908.

*** Cephalozia hamatiloba subsp. siamensis (N.Kitag.) Váňa, Acta Bot. Fenn. 177: 16, 2004. Bas.: Cephalozia siamensis N.Kitag., J. Hattori Bot. Lab. 32: 293, 1969.

*** Cephalozia lacinulata (J.B.Jack ex Gottsche et Rabenh.) Spruce, Cephalozia: 45, 1882. Bas.: Jungermannia lacinulata J.B.Jack ex Gottsche et Rabenh., Hepat. Eur., Leberm. 62-64: no. 624, 1877.

*** Cephalozia lucens (A.Evans) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 496 (326), 1908. Bas.: Jungermannia lucens A.Evans, Trans. Connecticut Acad. Arts 8 (15): 258, 1891.

*** Cephalozia macgregorii (Steph.) Váňa, Phytotaxa 112 (1): 12, 2013. Bas.: Hygrobiella macgregorii Steph., Hedwigia 34 (2): 45, 1895.

*** Cephalozia macounii (Austin) Austin, Hepat. bor.-amer.: 14, 1873. Bas.: Jungermannia macounii Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 222, 1869.

*** Cephalozia maxima Steph., Sp. Hepat. (Stephani) 6: 441, 1924.

*** Cephalozia mollusca (De Not.) Váňa, Phytotaxa 112 (1): 12, 2013. Bas.: Jungermannia mollusca De Not., Epat. Borneo: 16, 1874.

* Cephalozia neesiana Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 429 (305), 1908. 33

*** Cephalozia nishimurae (N.Kitag.) Váňa, Phytotaxa 112 (1): 12, 2013. Bas.: Hygrobiella nishimurae N.Kitag., Misc. Bryol. Lichenol. 9 (4): 69, 1982.

** Cephalozia pachygyna R.M.Schust. ex J.J.Engel, Novon 17 (3): 313, 2007.

** Cephalozia physocaula (Hook.f. et Taylor) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 485 (315), 1908. Bas.: Jungermannia physocaula Hook.f. et Taylor, London J. Bot. 3: 455, 1844.

** Cephalozia schusteriana J.J.Engel, Novon 17 (3): 313, 2007.

*** Cephalozia stolonacea (Herzog) Váňa, Phytotaxa 112 (1): 12, 2013. Bas.: Hygrobiella stolonacea Herzog, Trans. Brit. Bryol. Soc. 1 (4): 293, 1950.

*** Cephalozia tubulata (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877. Bas.: Jungermannia tubulata Hook.f. et Taylor, London J. Bot. 3: 463, 1844.

Excluded from the genus

* Cephalozia hians Steph., Sp. Hepat. (Stephani) 6: 441, 1924. 34

* Cephalozia indica Udar et D.Kumar, Geophytology 6 (1): 42, 1976. 35

* Cephalozia kodaikanalensis G.Asthana et Saumya Srivast., Geophytology 43 (1): 63, 2013. 36

* Cephalozia parvifolia Arnell, Rev. Bryol. 25 (1): 1, 1898. 37

* Cephalozia tricuspidata (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877. Bas.: Jungermannia tricuspidata Nees, Enum. Pl. Crypt. Javae: 31, 1830. 38

*** Fuscocephaloziopsis Fulford, Mem. New York Bot. Gard. 11 (3): 353, 1968.

** Fuscocephaloziopsis affinis (Lindb. ex Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013. Bas.: Cephalozia affinis Lindb. ex Steph., Bull. Herb. Boissier (sér. 2) 8 (4): 277 (291), 1908.

*** Fuscocephaloziopsis africana (Váňa) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013. Bas.: Cephalozia africana Váňa, Beih. Nova Hedwigia 90: 187, 1988.

*** Fuscocephaloziopsis albescens (Hook.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013. Bas.: Jungermannia albescens Hook., Brit. Jungermann.: tab. 72, 1815.

* Fuscocephaloziopsis albescens var. islandica (Nees) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013. Bas.: Jungermannia islandica Nees, Naturgesch. Eur. Leberm. 2: 29, 1836.

*** Fuscocephaloziopsis baldwinii (C.M.Cooke) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013. Bas.: Cephalozia baldwinii C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 35, 1904.

** Fuscocephaloziopsis biloba (Herzog) Fulford, Mem. New York Bot. Gard. 11 (3): 355, 1968. Bas.: Alobiella biloba Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 166, 1955.

*** Fuscocephaloziopsis catenulata (Huebener) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013. Bas.: Jungermannia catenulata Huebener, Hepaticol. germ.: 169, 1834.

*** Fuscocephaloziopsis catenulata subsp. nipponica (S.Hatt.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013. Bas.: Cephalozia nipponica S.Hatt., Bull. Tokyo Sci. Mus. 11: 74, 1944.

*** Fuscocephaloziopsis connivens (Dicks.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013. Bas.: Jungermannia connivens Dicks., Fasc. Pl. Crypt. Brit. 4: 19, 1801.

*** Fuscocephaloziopsis connivens subsp. fissa (Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013. Bas.: Cephalozia fissa Steph., Hedwigia 30 (5): 204, 1891.

*** Fuscocephaloziopsis connivens subsp. sandwicensis (Mont.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013. Bas.: Jungermannia sandwicensis Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 249, 1843.

*** Fuscocephaloziopsis crassifolia (Lindenb. et Gottsche) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013. Bas.: Jungermannia crassifolia Lindenb. et Gottsche, Syn. Hepat. 5: 685, 1847.

*** Fuscocephaloziopsis gollanii (Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013. Bas.: Cephalozia gollanii Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 428 (304), 1908.

*** Fuscocephaloziopsis leucantha (Spruce) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013. Bas.: Cephalozia leucantha Spruce, Cephalozia: 68, 1882.

*** Fuscocephaloziopsis loitlesbergeri (Schiffn.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013. Bas.: Cephalozia loitlesbergeri Schiffn., Österr. Bot. Z. 62 (1): 10, 1912.

*** Fuscocephaloziopsis lunulifolia (Dumort.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013. Bas.: Jungermannia lunulifolia Dumort., Syll. Jungerm. Europ.: 61, 1831. 39

*** Fuscocephaloziopsis macrostachya (Kaal.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013. Bas.: Cephalozia macrostachya Kaal., Rev. Bryol. 29 (1): 8, 1902.

*** Fuscocephaloziopsis macrostachya subsp. australis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013. Bas.: Cephalozia macrostachya subsp. australis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 754, 1974.

*** Fuscocephaloziopsis macrostachya subsp. macrostachya var. spiniflora (Schiffn.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013. Bas.: Cephalozia spiniflora Schiffn., Hedwigia 54 (6): 323, 1914.

*** Fuscocephaloziopsis monticola (J.D.Godfrey) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013. Bas.: Schofieldia monticola J.D.Godfrey, Bryologist 79 (3): 315, 1976.

*** Fuscocephaloziopsis pachycaulis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013. Bas.: Cephalozia pachycaulis R.M.Schust., Bryologist 96 (4): 623, 1993.

*** Fuscocephaloziopsis pleniceps (Austin) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013. Bas.: Jungermannia pleniceps Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 222, 1869.

** Fuscocephaloziopsis pleniceps var. caroliniana (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013. Bas.: Cephalozia pleniceps var. caroliniana R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 780, 1974.

*** Fuscocephaloziopsis pulvinata (Steph.) Fulford, Mem. New York Bot. Gard. 11 (3): 355, 1968. Bas.: Alobiella pulvinata Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 572 (356), 1908.

* Fuscocephaloziopsis schusteri (Sushil K.Singh et D.K.Singh) Váňa, Phytotaxa 183 (4): 291, 2014. Bas.: Cephalozia schusteri Sushil K.Singh et D.K.Singh, Lindbergia 32 (1): 1, 2007.

*** Fuscocephaloziopsis subintegra Gradst. et Váňa, Cryptog. Bryol. 25 (3): 274, 2004.

*** Fuscocephaloziopsis zoopsioides (Horik.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013. Bas.: Cephalozia zoopsioides Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 178, 1934.

*** Nowellia Mitt., Nat. hist. Azores: 321, 1870.

** sect. Acronowellia R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 816, 1974.

*** Nowellia reedii H.Rob., Bryologist 73 (1): 150, 1970.

*** Nowellia yunckeri Fulford, Mem. New York Bot. Gard. 11 (3): 329, 1968.

** sect. Metanowellia (Grolle) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 816, 1974. Bas.: Nowellia subg. Metanowellia Grolle, J. Hattori Bot. Lab. 31: 33, 1968.

*** Nowellia borneensis (De Not.) Schiffn., Hepat. (Engl.-Prantl): 98, 1893. Bas.: Jungermannia curvifolia var. borneensis De Not., Epat. Borneo: 19, 1874.

*** Nowellia dominicensis Steph., Sp. Hepat. (Stephani) 6: 443, 1924.

*** Nowellia evansii Grolle, J. Hattori Bot. Lab. 31: 33, 1968.

*** Nowellia langii Pearson, J. Linn. Soc., Bot. 46 (305): 25, 1922.

*** Nowellia pusilla Grolle, J. Hattori Bot. Lab. 31: 37, 1968.

*** Nowellia wrightii (Gottsche ex Spruce) Steph. ex Duss, Enum. musc. Antilles franç., Hép.: 21, 1903. Bas.: Cephalozia wrightii Gottsche ex Spruce, J. Linn. Soc., Bot. 30 (210): 354, 1895.

** sect. Nowellia

*** Nowellia aciliata (P.C.Chen et P.C.Wu) Mizut., Hikobia 11: 469, 1994. Bas.: Nowellia curvifolia var. aciliata P.C.Chen et P.C.Wu, Obs. fl. Hwangs.: 6, 1965.

*** Nowellia curvifolia (Dicks.) Mitt., Nat. hist. Azores: 321, 1870. Bas.: Jungermannia curvifolia Dicks., Fasc. Pl. Crypt. Brit. 2: 15, 1790.

** Odontoschismatoideae H.Buch ex Grolle

*** Odontoschisma (Dumort.) Dumort., Recueil Observ. Jungerm.: 19, 1835. Bas.: Pleuroschisma sect. Odontoschisma Dumort., Syll. Jungerm. Europ.: 68, 1831.

** sect. Cladopodiella (H.Buch) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014. Bas.: Cladopodiella H.Buch, Memoranda Soc. Fauna Fl. Fennica 1: 89, 1927.

*** Odontoschisma francisci (Hook.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 12, 2013. Bas.: Jungermannia francisci Hook., Brit. Jungermann.: tab. 49, 1813.

** sect. Denudata R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 833, 1974.

*** Odontoschisma brasiliense Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 585 (369), 1908.

** Odontoschisma cleefii Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014.

*** Odontoschisma denudatum (Mart.) Dumort., Recueil Observ. Jungerm.: 19, 1835. Bas.: Jungermannia scalaris var. denudata Mart., Fl. crypt. erlang.: 183, 1817.

** Odontoschisma denudatum subsp. naviculare (Steph.) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014. Bas.: Jamesoniella navicularis Steph., Sp. Hepat. (Stephani) 6: 101, 1917.

** Odontoschisma denudatum subsp. sandvicense (Ångstr.) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014. Bas.: Sphagnoecetis sandvicensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 22, 1872.

*** Odontoschisma elongatum (Lindb.) A.Evans, Rhodora 14 (157): 13, 1912. Bas.: Odontoschisma denudatum f. elongatum Lindb., Helsingf. Dagbl. 1874 (45, 16 Feb): 2, 1874.

*** Odontoschisma engelii Gradst. et Burghardt, Fieldiana, Bot. (n.ser.) 47: 194, 2008.

*** Odontoschisma longiflorum (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 419, 1877. Bas.: Sphagnoecetis longiflora Taylor, London J. Bot. 5: 281, 1846.

*** Odontoschisma macounii (Austin) Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 92, 1884. Bas.: Sphagnoecetis macounii Austin, Bull. Torrey Bot. Club 3 (3): 13, 1872.

*** Odontoschisma portoricense (Hampe et Gottsche) Steph., Hedwigia 27 (11/12): 296, 1888. Bas.: Sphagnoecetis portoricensis Hampe et Gottsche, Linnaea 25 (3): 343, 1852 [1853].

** Odontoschisma pseudogrosseverrucosum Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014.

*** Odontoschisma purpuratum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 297, 1950.

*** Odontoschisma soratamum Fulford, Mem. New York Bot. Gard. 11 (3): 338, 1968.

*** Odontoschisma variabile (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 419, 1877. Bas.: Sphagnoecetis variabilis Lindenb. et Gottsche, Syn. Hepat. 5: 688, 1847.

** Odontoschisma zhui Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1020, 2014.

** sect. Iwatsukia (N.Kitag.) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014. Bas.: Iwatsukia N.Kitag., J. Hattori Bot. Lab. 27: 178, 1964.

*** Odontoschisma bifidum (Fulford) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014. Bas.: Cladomastigum bifidum Fulford, Acta Bot. Venez. 2 (5/8): 80, 1967.

*** Odontoschisma jishibae (Steph.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 13, 2013. Bas.: Cephalozia jishibae Steph., Sp. Hepat. (Stephani) 6: 437, 1924.

*** Odontoschisma spinosum (Fulford) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014. Bas.: Cladomastigum spinosum Fulford, Mem. New York Bot. Gard. 23: 840, 1972.

** sect. Neesia Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014.

*** Odontoschisma fluitans (Nees) L.Söderstr. et Váňa, Phytotaxa 112 (1): 12, 2013. Bas.: Jungermannia fluitans Nees, Flora 6 (2): 30, 1823.

** sect. Odontoschisma

*** Odontoschisma grosseverrucosum Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 593 (377), 1908.

*** Odontoschisma sphagni (Dicks.) Dumort., Recueil Observ. Jungerm.: 19, 1835. Bas.: Jungermannia sphagni Dicks., Fasc. Pl. Crypt. Brit. 1: 6, 1785.

** Odontoschisma steyermarkii Gradst. et Ilk.-Borg., Nova Hedwigia 100 (1/2): 38, 2015 [2014 online].

Incertae sedis

* Odontoschisma obcordatum (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 586 (370), 1908. Bas.: Cephalozia obcordata Spruce, Cephalozia: 61, 1882. 40

** Schiffnerioideae R.M.Schust.

** Schiffneria Steph., Österr. Bot. Z. 44 (1): 1, 1894.

*** Schiffneria hyalina Steph., Österr. Bot. Z. 44 (1): 1, 1894.

** Trabacelluloideae R.M.Schust.

** Haesselia Grolle et Gradst., J. Hattori Bot. Lab. 64: 327, 1988.

*** Haesselia acuminata Gradst., Trop. Bryol. 1: 30, 1989.

*** Haesselia roraimensis Grolle et Gradst., J. Hattori Bot. Lab. 64: 327, 1988.

** Trabacellula Fulford, Acta Bot. Venez. 2 (5/8): 86, 1967.

*** Trabacellula tumidula Fulford, Acta Bot. Venez. 2 (5/8): 86, 1967.

*** Cephaloziellaceae Douin

by J. Váňa

The treatment of Cephaloziellaceae follows what was outlined in. Some nomenclatural and taxonomic notes can also be found in and, ). The placement of Phycolepidozia follows.

*** Allisoniella E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 80, 1965.

** sect. Allisoniella

*** Allisoniella nigra (Rodway) R.M.Schust., Nova Hedwigia 22: 137, 1971 [1972]. Bas.: Sphenolobus niger Rodway, Tasm. Bryoph.: 33, 1917.

** Allisoniella nigra var. acutiloba J.J.Engel, Novon 17 (3): 313, 2007.

** Allisoniella nigra subsp. novaezelandiae R.M.Schust., Nova Hedwigia 22: 143, 1971 [1972].

*** Allisoniella recurva R.M.Schust., Nova Hedwigia 22: 146, 1971 [1972].

*** Allisoniella subbipartita (C.Massal.) R.M.Schust. et J.J.Engel, Nova Hedwigia 22: 147, 1971 [1972]. Bas.: Cephalozia subbipartita C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 235, 1885.

*** Allisoniella tasmanica R.M.Schust., Nova Hedwigia 22: 145, 1971 [1972].

** sect. Protomarsupella (R.M.Schust.) R.M.Schust., Nova Hedwigia 22: 150, 1971. Bas.: Protomarsupella R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 264, 1966.

*** Allisoniella scottii (R.M.Schust.) R.M.Schust., Nova Hedwigia 22: 151, 1971 [1972]. Bas.: Protomarsupella scottii R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 267, 1966.

*** Amphicephalozia R.M.Schust., Nova Hedwigia 22: 131, 1971 [1972].

*** Amphicephalozia africana Váňa et M.Wigginton, J. Bryol. 30 (1): 55, 2008.

*** Amphicephalozia amplexicaulis R.M.Schust., Nova Hedwigia 22: 133, 1971 [1972].

*** Amphicephalozia geisslerae Pócs et Váňa, Polish Bot. J. 46 (2): 145, 2001.

*** Anastrophyllopsis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 81 (1): 15, 2013. Bas.: Anastrophyllum sect. Anastrophyllopsis R.M.Schust., Beih. Nova Hedwigia 119: 310, 2002.

*** Anastrophyllopsis involutifolia (Mont. ex Gottsche, Lindenb. et Nees) Váňa et L.Söderstr., Phytotaxa 81 (1): 15, 2013. Bas.: Jungermannia involutifolia Mont. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 1: 81, 1844.

*** Anastrophyllopsis revoluta (Steph.) Váňa et L.Söderstr., Phytotaxa 81 (1): 16, 2013. Bas.: Anastrophyllum revolutum Steph., Hedwigia 32 (3): 139, 1893.

*** Anastrophyllopsis subcomplicata (Lehm. et Lindenb.) Váňa et L.Söderstr., Phytotaxa 81 (1): 16, 2013. Bas.: Jungermannia subcomplicata Lehm. et Lindenb., Nov. Stirp. Pug. 7: 4, 1838.

** Cephalojonesia Grolle, Rev. Bryol. Lichénol. 37 (4): 763, 1970 [1971].

*** Cephalojonesia incuba Grolle et Vanden Berghen, Rev. Bryol. Lichénol. 37 (4): 764, 1970 [1971].

*** Cephalojonesia incuba subsp. mexicana Burghardt, Gradst. et Váňa, J. Hattori Bot. Lab. 100: 35, 2006.

** Cephalomitrion R.M.Schust., Nova Hedwigia 61 (3/4): 550, 1995.

*** Cephalomitrion aterrimum (Steph.) R.M.Schust., Nova Hedwigia 61 (3/4): 554, 1995. Bas.: Cephalozia aterrima Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 501 (331), 1908.

*** Cephaloziella (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 nom. conserv. Bas.: Cephalozia subg. Cephaloziella Spruce, Cephalozia: 62, 1882. 41

** subg. Cephaloziella

*** Cephaloziella aenigmatica R.M.Schust., Nova Hedwigia 63 (1/2): 20, 1996.

*** Cephaloziella anthelioides S.W.Arnell, Bot. Not. 105: 322, 1952.

*** Cephaloziella arctogena (R.M.Schust.) Konstant., Arctoa 3: 126, 1994. Bas.: Cephaloziella rubella var. arctogena R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 125, 1980.

** Cephaloziella arenaria (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963. Bas.: Cephalozia arenaria Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 504 (334), 1908.

*** Cephaloziella aspericaulis Jørg., Bergens Mus. Skr. (n.ser.) 16: 197, 1934.

*** Cephaloziella baumgartneri Schiffn., Verh. K.K. Zool.-Bot. Ges. Wien 56 (3): 273, 1906.

** Cephaloziella breviperianthia C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 208, 1981.

** Cephaloziella brinkmannii Douin, Mém. Soc. Bot. France 29: 75, 1920.

*** Cephaloziella capensis (Sim) S.W.Arnell, Bot. Not. 105: 326, 1952. Bas.: Cephalozia capensis Sim, Trans. Roy. Soc. South Africa 15 (1): 87, 1926.

*** Cephaloziella capillaris (Steph.) Douin, Mém. Soc. Bot. France 29: 59, 1920. Bas.: Cephalozia capillaris Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 496 (326), 1908.

*** Cephaloziella crassigyna (R.M.Schust.) R.M.Schust., Nova Hedwigia 61 (3/4): 556, 1995. Bas.: Cephaloziella aterrima var. crassigyna R.M.Schust., Nova Hedwigia 22: 211, 1971 [1972].

*** Cephaloziella crispata N.Kitag., J. Hattori Bot. Lab. 32: 301, 1969.

*** Cephaloziella densifolia R.M.Schust., Nova Hedwigia 22: 199, 1971 [1972].

** Cephaloziella densifolia var. dubia R.M.Schust., Nova Hedwigia 22: 202, 1971 [1972].

*** Cephaloziella divaricata (Sm.) Schiffn., Hepat. (Engl.-Prantl): 99, 1893. Bas.: Jungermannia divaricata Sm., Engl. Bot. 10: tab. 719, 1800.

** Cephaloziella divaricata var. scabra (M.Howe) Haynes, Bryologist 12 (4): 68, 1909. Bas.: Cephalozia divaricata var. scabra M.Howe, Mem. Torrey Bot. Club 7: 129, 1899. 42

** Cephaloziella dusenii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 49, 1900.

*** Cephaloziella elachista (J.B.Jack ex Gottsche et Rabenh.) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 336, 1900. Bas.: Jungermannia elachista J.B.Jack ex Gottsche et Rabenh., Hepat. Eur., Leberm. 58-59: no. 574, 1873.

** Cephaloziella elachista var. spinophylla (C.Gao) C.Gao, Fl. Bryoph. Sin. 9: 181, 2003. Bas.: Cephaloziella spinophylla C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 208, 1981.

** Cephaloziella elegans (Heeg) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 336, 1900. Bas.: Cephalozia elegans Heeg, Rev. Bryol. 20 (5): 82, 1893.

*** Cephaloziella exigua R.M.Schust., Nova Hedwigia 63 (1/2): 49, 1996.

*** Cephaloziella exiliflora (Taylor) Douin, Mém. Soc. Bot. France 29: 72, 1920. Bas.: Jungermannia exiliflora Taylor, London J. Bot. 5: 279, 1846.

** Cephaloziella fragillima (Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 409, 1976. Bas.: Cephalozia fragillima Spruce, Mem. Torrey Bot. Club 1 (3): 131, 1890.

*** Cephaloziella garsidei S.W.Arnell, Rev. Bryol. Lichénol. 23 (1/2): 173, 1954.

*** Cephaloziella grandiretis (R.M.Schust.) R.M.Schust., Nova Hedwigia 63 (1/2): 35, 1996. Bas.: Cephaloziella byssacea subsp. grandiretis R.M.Schust., Nova Hedwigia 22: 195, 1971 [1972].

*** Cephaloziella grimsulana (J.B.Jack ex Gottsche et Rabenh.) Lacout., Hépat. France: 52, 1905. Bas.: Jungermannia grimsulana J.B.Jack ex Gottsche et Rabenh., Hepat. Eur., Leberm. 53–55: no. 526, 1872.

*** Cephaloziella hampeana (Nees) Schiffn. ex Loeske, Moosfl. Harz.: 92, 1903. Bas.: Jungermannia hampeana Nees, Naturgesch. Eur. Leberm. 3: 560, 1838.

** Cephaloziella hebridensis Steph., Hedwigia 32 (5): 316, 1893.

*** Cephaloziella herzogiana (Pandé et K.P.Srivast.) Udar et D.Kumar, Geophytology 6 (1): 45, 1976. Bas.: Cephalozia herzogiana Pandé et K.P.Srivast., Feddes Repert. Spec. Nov. Regni Veg. 58: 75, 1955.

** Cephaloziella heteroica (C.M.Cooke) Douin, Mém. Soc. Bot. France 29: 76, 1920. Bas.: Cephalozia heteroica C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 38, 1904.

** Cephaloziella hyalina Douin, Mém. Soc. Bot. France 29: 77, 1920.

* Cephaloziella hyalina var. rappii (Douin) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 56, 1980. Bas.: Cephaloziella rappii Douin, Mém. Soc. Bot. France 29: 77, 1920.

*** Cephaloziella inaequalis R.M.Schust., Nova Hedwigia 22: 186, 1971 [1972].

*** Cephaloziella invisa R.M.Schust., Nova Hedwigia 63 (1/2): 30, 1996.

** Cephaloziella kilohanensis (C.M.Cooke) Douin, Mém. Soc. Bot. France 29: 85, 1920. Bas.: Cephalozia kilohanensis C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 37, 1904.

*** Cephaloziella longii Váňa, Folia Geobot. Phytotax. 27 (2): 193, 1992.

*** Cephaloziella lycopodioides (Sim) S.W.Arnell, Bot. Not. 105: 321, 1952. Bas.: Cephalozia lycopodioides Sim, Trans. Roy. Soc. South Africa 15 (1): 85, 1926.

** Cephaloziella mammillifera R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987.

*** Cephaloziella massalongi (Spruce) Müll.Frib., Lebermoose 2 (17): 191, 1913. Bas.: Cephalozia massalongi Spruce, Cephalozia: 71, 1882.

*** Cephaloziella muelleriana R.M.Schust., Nova Hedwigia 63 (1/2): 24, 1996.

*** Cephaloziella natalensis (Sim) S.W.Arnell, Hepat. South Africa: 338, 1963. Bas.: Cephalozia natalensis Sim, Trans. Roy. Soc. South Africa 15 (1): 86, 1926.

** Cephaloziella nicholsonii Douin, Rev. Bryol. 40 (6): 81, 1913.

*** Cephaloziella nothogena R.M.Schust., Nova Hedwigia 63 (1/2): 37, 1996.

** Cephaloziella obtusilobula R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 108, 1980.

** Cephaloziella patulifolia (Steph.) Douin, Mém. Soc. Bot. France 29: 70, 1920. Bas.: Cephalozia patulifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 509 (339), 1908.

*** Cephaloziella phyllacantha (C.Massal. et Carestia) Müll.Frib., Lebermoose 2 (17): 194, 1913. Bas.: Anthelia phyllacantha C.Massal. et Carestia, Nuovo Giorn. Bot. Ital. 12 (4): 340, 1880.

*** Cephaloziella polystratosa (R.M.Schust. et Damsh.) Konstant., Bot. Zhurn. (Moscow & Leningrad) 85 (10): 127, 2000. Bas.: Cephaloziella byssacea var. polystratosa R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987.

*** Cephaloziella pseudocrassigyna R.M.Schust. ex J.J.Engel, Novon 17 (3): 313, 2007.

*** Cephaloziella pulcherrima R.M.Schust., Nova Hedwigia 22: 203, 1971 [1972].

** Cephaloziella pulcherrima subsp. sphagnicola R.M.Schust., Nova Hedwigia 22: 205, 1971 [1972].

** Cephaloziella pungens Steph. ex Fulford, Mem. New York Bot. Gard. 11 (4): 410, 1976.

*** Cephaloziella rubella (Nees) Warnst., Krypt.-Fl. Brandenburg, Leber- & Torfmoose: 231, 1902. Bas.: Jungermannia rubella Nees, Naturgesch. Eur. Leberm. 2: 236, 1836.

*** Cephaloziella schelpei S.W.Arnell, Bot. Not. 110 (1): 18, 1957.

*** Cephaloziella spegazziniana (C.Massal.) Douin, Mém. Soc. Bot. France 29: 69, 1920. Bas.: Cephalozia spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 234, 1885.

*** Cephaloziella spinicaulis Douin, Mém. Soc. Bot. France 29: 62, 1920.

*** Cephaloziella spinigera (Lindb.) Jørg., Bergens Mus. Skr. (n.ser.) 16: 189, 1934. Bas.: Cephalozia spinigera Lindb., Musci Scand.: 4, 1879.

*** Cephaloziella stellulifera (Taylor ex Carrington et Pearson) Croz., Rev. Bryol. 30 (2): 31, 1903. Bas.: Jungermannia stellulifera Taylor ex Carrington et Pearson, Hepat. Brit. Exsicc. Fasc. I: no. 32, 1878.

*** Cephaloziella stephanii Schiffn. ex Douin, Mém. Soc. Bot. France 29: 85, 1920.

** Cephaloziella stolonifera R.M.Schust., Phytologia 39 (6): 426, 1978.

** Cephaloziella subtilis (Lindenb. et Gottsche) Steph., Hedwigia 32 (5): 318, 1893. Bas.: Jungermannia subtilis Lindenb. et Gottsche, Syn. Hepat. 5: 685, 1847.

*** Cephaloziella sumatrana Schiffn. ex Douin, Mém. Soc. Bot. France 29: 59, 1920.

*** Cephaloziella tabularis S.W.Arnell, Bot. Not. 105: 318, 1952.

*** Cephaloziella transvaalensis S.W.Arnell, Bot. Not. 110 (1): 19, 1957.

*** Cephaloziella triplicata S.W.Arnell, Bot. Not. 115: 203, 1962.

*** Cephaloziella umtaliensis S.W.Arnell, Bot. Not. 110 (1): 20, 1957.

*** Cephaloziella uncinata R.M.Schust., Meddel. Grønland 199 (1): 316, 1974.

** Cephaloziella uncinata var. brevigyna R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987.

** Cephaloziella uncinata var. sphagnicola R.M.Schust., Meddel. Grønland 199 (1): 323, 1974.

*** Cephaloziella vaginans Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 119, 1911.

*** Cephaloziella varians (Gottsche) Steph., Wiss. Ergebn. Schwed. Südpolar-Exped. [1901–1903] 4 (1): 4, 1905. Bas.: Jungermannia varians Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 452, 1890.

*** Cephaloziella verrucosa Steph., Hedwigia 32 (5): 318, 1893.

* Cephaloziella villaumei (Steph.) Váňa, Phytotaxa 112 (1): 3, 2013. Bas.: Cephalozia villaumei Steph., Sp. Hepat. (Stephani) 6: 437, 1924. 43

** Cephaloziella violacea Schljakov, Novosti Sist. Nizš. Rast. 15: 242, 1978.

** Cephaloziella welwitschii (Steph.) Douin, Mém. Soc. Bot. France 29: 58, 1920. Bas.: Cephalozia welwitschii Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 432 (308), 1908.

** subg. Dichiton (Mont.) Müll.Frib., Lebermoose 2 (27): 787, 1916. Bas.: Dichiton Mont., Syll. Gen. Sp. Crypt.: 52, 1856.

*** Cephaloziella calyculata (Durieu et Mont.) Müll.Frib., Lebermoose 2 (27): 787, 1916. Bas.: Jungermannia calyculata Durieu et Mont., Ann. Sci. Nat. Bot. (sér. 3) 11: 34, 1849.

*** Cephaloziella integerrima (Lindb.) Warnst., Krypt.-Fl. Brandenburg, Leber- & Torfmoose: 232, 1902. Bas.: Cephalozia integerrima Lindb., Acta Soc. Sci. Fenn. 10: 502, 1875.

** subg. Evansia (Douin et Schiffn.) Müll.Frib., Lebermoose 2 (27): 787, 1916. Bas.: Evansia Douin et Schiffn., Rev. Bryol. 40 (5): 66, 1913.

*** Cephaloziella antillana (Besch. et Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 405, 1976. Bas.: Cephalozia antillana Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxiii, 1889 [1890].

*** Cephaloziella dentata (Raddi) Steph., Bull. Herb. Boissier 5 (2): 78, 1897. Bas.: Jungermannia dentata Raddi, Jungermanniogr. Etrusca: 21, 1818.

*** Cephaloziella hirta (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963. Bas.: Cephalozia hirta Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 561 (345), 1908.

** Cephaloziella squarrosula (Trevis.) Steph., Hedwigia 32 (5): 318, 1893. Bas.: Cephalozia squarrosula Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877.

*** Cephaloziella subspinosa R.M.Schust., Nova Hedwigia 22: 210, 1971 [1972].

** subg. Prionolobus (Spruce) Müll.Frib., Lebermoose 2 (16): 110, 1912. Bas.: Cephalozia subg. Prionolobus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 508, 1885.

*** Cephaloziella acanthophora (S.Hatt.) Horik., Hikobia 1 (2): 79, 1951. Bas.: Prionolobus acanthophorus S.Hatt., Bull. Tokyo Sci. Mus. 11: 29, 1944.

*** Cephaloziella biokoensis Váňa et Frank Müll., Trop. Bryol. 24: 1, 2003.

*** Cephaloziella granatensis (J.B.Jack ex Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 411, 1976. Bas.: Cephalozia granatensis J.B.Jack ex Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 500 (330), 1908.

*** Cephaloziella grisea R.M.Schust., Phytologia 39 (6): 425, 1978.

* Cephaloziella meghalayensis Udar et Ad.Kumar, Lindbergia 8 (1): 34, 1982. 44

*** Cephaloziella microphylla (Steph.) Douin, Mém. Soc. Bot. France 29: 59, 1920. Bas.: Cephalozia microphylla Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 513 (343), 1908.

*** Cephaloziella tenuissima (Lehm.) Steph., Hedwigia 32 (5): 318, 1893. Bas.: Jungermannia tenuissima Lehm., Linnaea 4: 367, 1829.

*** Cephaloziella turneri (Hook.) Müll.Frib., Lebermoose 2 (17): 202, 1913. Bas.: Jungermannia turneri Hook., Brit. Jungermann.: tab. 29, 1812.

Incertae sedis

** Cephaloziella dentifolia Udar et Ad.Kumar, Lindbergia 8 (1): 30, 1982.

** Cephaloziella filum (Trevis.) Steph., Hedwigia 32 (5): 318, 1893. Bas.: Cephalozia filum Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877.

* Cephaloziella flexuosa C.Gao et K.C.Chang, Bull. Bot. Res., Harbin 4 (3): 88, 1984. 45

** Cephaloziella intricata Schiffn. ex Douin, Mém. Soc. Bot. France 29: 59, 1920.

** Cephaloziella levieri Schiffn. ex Douin, Mém. Soc. Bot. France 29: 80, 1920.

*** Cephaloziella pellucida R.M.Schust., Nova Hedwigia 63 (1/2): 57, 1996. 46

* Cephaloziella pygmaea (Spruce) Váňa, Phytotaxa 183 (4): 290, 2014. Bas.: Cephalozia pygmaea Spruce, Cephalozia: 69, 1882. 47

* Cephaloziella secundifolia Pearson, Ann. Bryol. 4: 106, 1931. 48

** Cephaloziella sinensis Douin, Rev. Bryol. 41 (1): 8, 1914.

** Cephaloziopsis (Spruce) Schiffn., Hepat. (Engl.-Prantl): 85, 1893. Bas.: Jungermannia sect. Cephaloziopsis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 511, 1885.

*** Cephaloziopsis intertexta (Gottsche) R.M.Schust., Nova Hedwigia 22: 183, 1971 [1972]. Bas.: Jungermannia intertexta Gottsche, Syn. Hepat. 1: 107, 1844.

** Chaetophyllopsis R.M.Schust., J. Hattori Bot. Lab. 23: 69, 1960 [1961].

*** Chaetophyllopsis whiteleggei (Carrington et Pearson) R.M.Schust. ex Hamlin, Rec. Domin. Mus. 7: 255, 1972. Bas.: Jungermannia whiteleggei Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1051, 1888.

** Cylindrocolea R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969.

** subg. Cylindrocolea

*** Cylindrocolea kiaeri (Austin) Váňa, Phytotaxa 112 (1): 2, 2013. Bas.: Jungermannia kiaeri Austin, Bull. Torrey Bot. Club 6 (3): 18, 1875.

*** Cylindrocolea sanctae-helenae M.Wigginton, Polish Bot. J. 58 (1): 107, 2013.

** sect. Cylindrocolea

*** Cylindrocolea andersonii R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 33, 1980.

*** Cylindrocolea brasiliensis D.P.Costa, N.D.Santos et Váňa, Bryologist 111 (4): 667, 2008.

*** Cylindrocolea chevalieri (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969. Bas.: Alobiella chevalieri Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 567 (351), 1908.

*** Cylindrocolea gittinsii (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 172, 1971 [1972]. Bas.: Cephaloziella gittinsii E.W.Jones, Trans. Brit. Bryol. Soc. 3 (3): 437, 1958.

*** Cylindrocolea madagascariensis (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969. Bas.: Lophozia madagascariensis Steph., Sp. Hepat. (Stephani) 6: 112, 1917.

*** Cylindrocolea nigerica (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 172, 1971 [1972]. Bas.: Cephaloziella nigerica E.W.Jones, Trans. Brit. Bryol. Soc. 3 (3): 435, 1958.

*** Cylindrocolea novae-caledoniae (Grolle) R.M.Schust., Nova Hedwigia 22: 161, 1971 [1972]. Bas.: Cephaloziella novae-caledoniae Grolle, Rev. Bryol. Lichénol. 29 (3/4): 208, 1960 [1961].

*** Cylindrocolea planifolia (Steph.) R.M.Schust., Nova Hedwigia 22: 164, 1971 [1972]. Bas.: Cephaloziella planifolia Steph., Hedwigia 32 (5): 317, 1893.

*** Cylindrocolea sprucei R.M.Schust., Nova Hedwigia 22: 163, 1971 [1972].

*** Cylindrocolea ugandica (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 171, 1971 [1972]. Bas.: Cephaloziella ugandica E.W.Jones, Trans. Brit. Bryol. Soc. 3 (3): 433, 1958.

** sect. Platycaulis R.M.Schust., Nova Hedwigia 22: 173, 1971.

*** Cylindrocolea recurvifolia (Steph.) Inoue, J. Jap. Bot. 47 (11): 348, 1972. Bas.: Cephalozia recurvifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 497 (327), 1908.

*** Cylindrocolea tagawae (N.Kitag.) R.M.Schust., Nova Hedwigia 22: 174, 1971 [1972]. Bas.: Cephaloziella tagawae N.Kitag., J. Hattori Bot. Lab. 32: 303, 1969.

** subg. Cylindroscyphus (Douin) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 20, 1980. Bas.: Cephaloziella subg. Cylindroscyphus Douin, Mém. Soc. Bot. France 29: 56, 1920.

*** Cylindrocolea abyssinica (Gola) Váňa, Phytotaxa 112 (1): 2, 2013. Bas.: Cephaloziella abyssinica Gola, Ann. Bot. (Rome) 13 (1): 68, 1914.

** Cylindrocolea obtusifolia Fulford, Mem. New York Bot. Gard. 11 (4): 401, 1976.

** Cylindrocolea reticulata Udar et Ad.Kumar, Lindbergia 8 (3): 181, 1982 [1983].

*** Cylindrocolea rhizantha (Mont.) R.M.Schust., Nova Hedwigia 22: 175, 1971 [1972]. Bas.: Jungermannia rhizantha Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 454, 1842.

*** Gottschelia Grolle, J. Hattori Bot. Lab. 31: 13, 1968.

*** Gottschelia maxima (Steph.) Grolle, J. Bryol. 25 (1): 6, 2003. Bas.: Tylimanthus maximus Steph., Sp. Hepat. (Stephani) 6: 249, 1922.

*** Gottschelia schizopleura (Spruce) Grolle, J. Hattori Bot. Lab. 31: 16, 1968. Bas.: Jungermannia schizopleura Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 517, 1885.

** Gymnocoleopsis (R.M.Schust.) R.M.Schust., Phytologia 39 (4): 243, 1978. Bas.: Gymnocolea subg. Gymnocoleopsis R.M.Schust., Bryologist 70 (1): 111, 1967.

*** Gymnocoleopsis capensis (S.W.Arnell) R.M.Schust., J. Hattori Bot. Lab. 78: 123, 1995. Bas.: Lophozia capensis S.W.Arnell, Svensk Bot. Tidskr. 47 (1): 112, 1953.

*** Gymnocoleopsis cylindriformis (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 78: 126, 1995. Bas.: Jungermannia cylindriformis Mitt., J. Linn. Soc., Bot. 15 (84): 196, 1876.

*** Herzogobryum Grolle, Rev. Bryol. Lichénol. 32 (3/4): 160, 1963 [1964]. Nom. nov. pro Chondrophyllum Herzog, Rev. Bryol. Lichénol. 21 (1/2): 46, 1952.

*** Herzogobryum atrocapillum (Hook.f. et Taylor) Grolle, Österr. Bot. Z. 113 (2): 228, 1966. Bas.: Gymnomitrion atrocapillum Hook.f. et Taylor, London J. Bot. 5: 258, 1846.

*** Herzogobryum filiforme R.M.Schust., Phytologia 45 (5): 422, 1980.

*** Herzogobryum molle Grolle, Österr. Bot. Z. 113 (2): 226, 1966.

*** Herzogobryum vermiculare (Schiffn.) Grolle, J. Hattori Bot. Lab. 28: 103, 1965. Bas.: Gymnomitrion vermiculare Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 2, 1890.

** Kymatocalyx Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 56, 1950.

*** Kymatocalyx africanus Váňa et M.Wigginton, Haussknechtia, Beih. 9: 158, 1999.

*** Kymatocalyx dominicensis (Spruce) Váňa, Österr. Bot. Z. 118 (5): 575, 1970. Bas.: Jungermannia dominicensis Spruce, J. Linn. Soc., Bot. 30 (210): 363, 1895.

*** Kymatocalyx madagascariensis (Steph.) Gradst. et Váňa, Haussknechtia, Beih. 9: 164, 1999. Bas.: Acrobolbus madagascariensis Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 460 (179), 1902.

*** Kymatocalyx rhizomaticus (Herzog) Gradst. et Váňa, Haussknechtia, Beih. 9: 166, 1999. Bas.: Stenorrhipis rhizomatica Herzog, Trans. Brit. Bryol. Soc. 1 (4): 290, 1950.

*** Lophonardia R.M.Schust., Phytologia 39 (4): 244, 1978.

*** Lophonardia jamesonii (Mont.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 19, 2013. Bas.: Jungermannia jamesonii Mont., Syll. Gen. Sp. Crypt.: 60, 1856.

*** Lophonardia laxifolia (Mont.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 20, 2013. Bas.: Sarcocyphos laxifolius Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 346, 1845.

*** Lophonardia tristaniana (S.W.Arnell) L.Söderstr. et Váňa, Phytotaxa 81 (1): 20, 2013. Bas.: Jungermannia tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 12, 1958.

*** Nothogymnomitrion R.M.Schust., J. Hattori Bot. Lab. 80: 43, 1996.

*** Nothogymnomitrion erosum (Carrington et Pearson) R.M.Schust., J. Hattori Bot. Lab. 80: 44, 1996. Bas.: Cesius erosus Carrington et Pearson, Pap. & Proc. Roy. Soc. Tasmania 1887: 8, 1888.

*** Obtusifolium S.W.Arnell, Ill. Moss Fl. Fennosc. Hep.: 309, 1956.

*** Obtusifolium obtusum (Lindb.) S.W.Arnell, Ill. Moss Fl. Fennosc. Hep.: 133, 1956. Bas.: Jungermannia obtusa Lindb., Musci Scand.: 7, 1879.

*** Oleolophozia L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010.

*** Oleolophozia perssonii (H.Buch et S.W.Arnell) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 51, 2010. Bas.: Lophozia perssonii H.Buch et S.W.Arnell, Bot. Not. 97: 382, 1944.

** Phycolepidozia R.M.Schust., Bull. Torrey Bot. Club 93 (6): 438, 1966 [1967].

** subg. Metaphycolepidozia Gradst., J.-P.Frahm et U.Schwarz, Taxon 63 (3): 506, 2014.

** Phycolepidozia indica Gradst., J.-P.Frahm et U.Schwarz, Taxon 63 (3): 499, 2014.

** subg. Phycolepidozia

*** Phycolepidozia exigua R.M.Schust., Bull. Torrey Bot. Club 93 (6): 440, 1966 [1967].

** Protolophozia (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 204, 1979. Bas.: Lophozia subg. Protolophozia R.M.Schust., Nova Hedwigia 15: 472, 1968.

** Protolophozia androgyna R.M.Schust. ex Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013. Based on: Lophozia androgyna R.M.Schust., Beih. Nova Hedwigia 119: 266, 2002, nom. inval.

** Protolophozia autoica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013. Bas.: Lophozia autoica R.M.Schust., Nova Hedwigia 15: 479, 1968.

** Protolophozia crispata (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013. Bas.: Lophozia crispata R.M.Schust., Nova Hedwigia 15: 474, 1968.

*** Protolophozia druceae (Grolle et E.A.Hodgs.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013. Bas.: Lophozia druceae Grolle et E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 112, 1972.

*** Protolophozia elongata (Steph.) Schljakov, Novosti Sist. Nizš. Rast. 16: 204, 1979. Bas.: Lophozia elongata Steph., Bull. Herb. Boissier (sér. 2) 2 (1): 41 (141), 1902.

*** Protolophozia herzogiana (E.A.Hodgs. et Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013. Bas.: Lophozia herzogiana E.A.Hodgs. et Grolle, Rev. Bryol. Lichénol. 31 (3/4): 152, 1962 [1963].

*** Protolophozia lancistipa (Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013. Bas.: Andrewsianthus lancistipus Grolle, Marion Prince Edw. Is: 230, 1971.

*** Protolophozia leucorhiza (Mitt.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013. Bas.: Jungermannia leucorhiza Mitt., J. Linn. Soc., Bot. 15 (82): 68, 1876.

*** Protolophozia longiflora (Herzog) L.Söderstr. et Váňa, Phytotaxa 76 (3): 51, 2013. Bas.: Orthocaulis longiflorus Herzog, Rev. Bryol. Lichénol. 23 (1/2): 32, 1954.

** Protolophozia monoica (E.A.Hodgs.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013. Bas.: Metahygrobiella monoica E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 76, 1965.

*** Protolophozia multicuspidata (Hook.f. et Taylor) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013. Bas.: Jungermannia multicuspidata Hook.f. et Taylor, London J. Bot. 3: 375, 1844.

** Protolophozia nivicola (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013. Bas.: Lophozia nivicola R.M.Schust., Nova Hedwigia 15: 477, 1968.

*** Protolophozia perssoniana (H.A.Mill.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013. Bas.: Lophozia perssoniana H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 508, 1963.

** Protolophozia subalpina (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013. Bas.: Lophozia autoica var. subalpina R.M.Schust., Nova Hedwigia 15: 482, 1968.

*** Protolophozia tasmanica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013. Bas.: Lophozia tasmanica R.M.Schust., Nova Hedwigia 15: 484, 1968.

*** Protolophozia verruculosa (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013. Bas.: Lophozia verruculosa R.M.Schust., Phytologia 39 (4): 242, 1978.

*** Lophoziaceae Cavers

by L. Söderström and J. Váňa

Lophoziaceae was broadly defined by excluding Anastrophyllaceae and Jamesonielloideae and also moving several traditionally included genera to other families. further refined the family. Nomenclatural and taxonomic notes can also be found in, ), and. The placement of Gerhildiella and Pseudocephaloziella in the family is provisional.

*** Andrewsianthus R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 66, 1961.

** subg. Andrewsianthus

*** Andrewsianthus aberrans (Nees et Mont.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 440, 1963. Bas.: Jungermannia aberrans Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 250, 1843.

*** Andrewsianthus bidens (Mitt. ex Steph.) R.M.Schust., Nova Hedwigia 15: 492, 1968. Bas.: Lophozia bidens Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 2 (1): 41 (141), 1902.

*** Andrewsianthus bilobus (Mitt.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 437, 1963. Bas.: Gymnanthe biloba Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 166, 1863.

*** Andrewsianthus carinatus Grolle, Marion Prince Edw. Is: 229, 1971.

*** Andrewsianthus cavifolius Grolle et Váňa, J. Hattori Bot. Lab. 38: 640, 1974.

*** Andrewsianthus chimbuensis R.M.Schust., Nova Hedwigia 15: 491, 1968.

*** Andrewsianthus kinabaluensis N.Kitag., J. Hattori Bot. Lab. 33: 205, 1970.

*** Andrewsianthus koponenii Váňa et Piippo, Ann. Bot. Fenn. 26 (3): 284, 1989.

*** Andrewsianthus mizutanii N.Kitag., J. Hattori Bot. Lab. 32: 307, 1969.

*** Andrewsianthus papillosus N.Kitag., J. Hattori Bot. Lab. 33: 207, 1970.

*** Andrewsianthus perigonialis (Hook.f. et Taylor) R.M.Schust., Beih. Nova Hedwigia 119: 336, 2002. Bas.: Jungermannia perigonialis Hook.f. et Taylor, London J. Bot. 3: 368, 1844.

*** Andrewsianthus puniceus (Nees) R.M.Schust. ex Grolle, Trans. Brit. Bryol. Soc. 4 (3): 439, 1963. Bas.: Jungermannia punicea Nees, Enum. Pl. Crypt. Javae: 32, 1830.

*** Andrewsianthus recurvifolius (Nees) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 710, 1969. Bas.: Jungermannia recurvifolia Nees, Enum. Pl. Crypt. Javae: 32, 1830.

*** Andrewsianthus sundaicus (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 710, 1969. Bas.: Anastrophyllum sundaicum Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 202, 1898.

*** Andrewsianthus zantenii Váňa, J. Hattori Bot. Lab. 38: 645, 1974.

** subg. Cephalolobus (R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 119: 326, 2002. Bas.: Cephalolobus R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 244, 1966.

*** Andrewsianthus hodgsoniae (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007. Bas.: Cephalolobus hodgsoniae R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 254, 1966.

*** Andrewsianthus marionensis (S.W.Arnell) Grolle, Marion Prince Edw. Is: 232, 1971. Bas.: Lophozia marionensis S.W.Arnell, Svensk Bot. Tidskr. 47 (3): 421, 1953.

*** Andrewsianthus scabrellus (C.Massal.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007. Bas.: Cephalozia scabrella C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 233, 1885.

** Andrewsianthus sphenoloboides (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007. Bas.: Cephalolobus sphenoloboides R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 251, 1966.

** Gerhildiella Grolle, Rev. Bryol. Lichénol. 34 (1/2): 187, 1966.

*** Gerhildiella rossneriana Grolle, Rev. Bryol. Lichénol. 34 (1/2): 187, 1966.

*** Heterogemma (Jørg.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010]. Bas.: Lophozia sect. Heterogemma Jørg., Bergens Mus. Skr. (n.ser.) 16: 146, 1934.

*** Heterogemma capitata (Hook.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010]. Bas.: Jungermannia capitata Hook., Brit. Jungermann.: tab. 80, 1816.

*** Heterogemma laxa (Lindb.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010]. Bas.: Jungermannia laxa Lindb., Acta Soc. Sci. Fenn. 10: 529, 1875.

*** Heterogemma patagonica (Herzog et Grolle) L.Söderstr. et Váňa, Phytotaxa 97 (2): 29, 2013. Bas.: Lophozia patagonica Herzog et Grolle, Rev. Bryol. Lichénol. 28 (3/4): 343, 1959 [1960].

*** Lophozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835. Bas.: Jungermannia sect. Lophozia Dumort., Syll. Jungerm. Europ.: 53, 1831. 49

*** Lophozia ascendens (Warnst.) R.M.Schust., Bryologist 55 (3): 180, 1952. Bas.: Sphenolobus ascendens Warnst., Hedwigia 57 (1/2): 63, 1916.

* Lophozia austrosibirica Bakalin, Ann. Bot. Fenn. 40 (1): 49, 2003.

** Lophozia ciliata Damsh., L.Söderstr. et H.Weibull, Lindbergia 25 (1): 3, 2000.

*** Lophozia guttulata (Lindb. et Arnell) A.Evans, Proc. Wash. Acad. Sci. 2: 302, 1900. Bas.: Jungermannia guttulata Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 51, 1889.

* Lophozia jamaicensis (Nees) Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 128 (155), 1901. Bas.: Jungermannia jamaicensis Nees, Syn. Hepat. 1: 105, 1844. 50

** Lophozia lacerata N.Kitag., Hikobia 3 (3): 172, 1963.

** Lophozia lantratoviae Bakalin, Ann. Bot. Fenn. 40 (1): 47, 2003.

** Lophozia murmanica Kaal., Rep. Second Norweg. Arctic Exped. 11: 34, 1906.

** Lophozia pacifica Bakalin, Bryologist 114 (2): 302, 2011.

* Lophozia pallida (Steph.) Grolle, J. Jap. Bot. 39 (6): 174, 1964. Bas.: Anastrophyllum pallidum Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1131 (114), 1901. 51

** Lophozia savicziae Schljakov, Novosti Sist. Nizš. Rast. 10: 299, 1973.

* Lophozia schusterana Schljakov, Novosti Sist. Nizš. Rast. 12: 320, 1975.

*** Lophozia silvicola H.Buch, Beret. 18 Skand. Naturforskarmøde: 228, 1929.

*** Lophozia silvicoloides N.Kitag., J. Hattori Bot. Lab. 28: 276, 1965.

* Lophozia subapiculata R.M.Schust. et Damsh., Meddel. Grønland 199 (1): 104, 1974. 52

** Lophozia udarii S.Srivast., S.C.Srivast. et K.K.Rawat, Nelumbo 55: 130, 2013. 53

*** Lophozia ventricosa (Dicks.) Dumort., Recueil Observ. Jungerm.: 17, 1835. Bas.: Jungermannia ventricosa Dicks., Fasc. Pl. Crypt. Brit. 2: 14, 1790. 54

* Lophozia wenzelii (Nees) Steph., Bull. Herb. Boissier (sér. 2) 2 (1): 35 (135), 1902. Bas.: Jungermannia wenzelii Nees, Naturgesch. Eur. Leberm. 2: 58, 1836. 55

Excluded from the genus

* Lophozia serpens (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835. Bas.: Jungermannia serpens Dumort., Syll. Jungerm. Europ.: 56, 1831. 56

** Lophoziopsis Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010].

*** Lophoziopsis excisa (Dicks.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010]. Bas.: Jungermannia excisa Dicks., Fasc. Pl. Crypt. Brit. 3: 11, 1793.

** Lophoziopsis excisa var. elegans (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010]. Bas.: Lophozia excisa var. elegans R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 522, 1969.

** Lophoziopsis excisa var. infuscata (R.M.Schust. et Damsh.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010]. Bas.: Lophozia excisa var. infuscata R.M.Schust. et Damsh., Meddel. Grønland 199 (1): 94, 1974.

** Lophoziopsis excisa var. succulenta (R.M.Schust. et Damsh.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010]. Bas.: Lophozia excisa var. succulenta R.M.Schust. et Damsh., Meddel. Grønland 199 (1): 96, 1974.

*** Lophoziopsis longidens (Lindb.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010]. Bas.: Jungermannia longidens Lindb., Helsingf. Dagbl. 1876 (323, 26 Nov.): 2, 1876.

** Lophoziopsis longidens subsp. arctica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 97 (2): 28, 2013. Bas.: Lophozia longidens subsp. arctica R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 539, 1969. 57

*** Lophoziopsis pellucida (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010]. Bas.: Lophozia pellucida R.M.Schust., Canad. J. Bot. 39 (4): 978, 1961.

** Lophoziopsis pellucida var. minor (R.M.Schust.) L.Söderstr. et Váňa, Phytotaxa 97 (2): 28, 2013. Bas.: Lophozia pellucida var. minor R.M.Schust., Canad. J. Bot. 39 (4): 984, 1961.

*** Lophoziopsis polaris (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010]. Bas.: Lophozia alpestris subsp. polaris R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 614, 1969.

** Lophoziopsis polaris var. sphagnorum (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010]. Bas.: Lophozia alpestris f. sphagnorum R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 619, 1969.

* Lophoziopsis propagulifera (Gottsche) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010]. Bas.: Jungermannia propagulifera Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 451, 1890. 58

** Lophoziopsis rubrigemma (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010]. Bas.: Lophozia rubrigemma R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 621, 1969.

** Pseudocephaloziella R.M.Schust., Phytologia 39 (4): 243, 1978.

*** Pseudocephaloziella epiphytica R.M.Schust., Phytologia 39 (4): 243, 1978.

*** Trilophozia (R.M.Schust.) Bakalin, Monogr. Lophozia: 34, 2005. Bas.: Tritomaria subg. Trilophozia R.M.Schust., Amer. Midl. Naturalist 49 (2): 382, 1953.

*** Trilophozia quinquedentata (Huds.) Bakalin, Monogr. Lophozia: 34, 2005. Bas.: Jungermannia quinquedentata Huds., Fl. Angl. (Hudson): 433, 1762.

** Trilophozia quinquedentata var. asymmetrica (Horik.) L.Söderstr. et Váňa, Phytotaxa 97 (2): 32, 2013. Bas.: Lophozia asymmetrica Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 153, 1934.

*** Tritomaria Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909. Based on: Tritomaria Schiffn., Ber. Naturwiss.-Med. Vereins Innsbruck 31 [Beilage]: 12, 1908.

*** Tritomaria exsecta (Schmidel) Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909. Bas.: Jungermannia exsecta Schmidel, Syst. Samml. Crypt. Gew. 2: 5, 1797, nom. conserv.

** Tritomaria exsecta subsp. novaezelandiae J.J.Engel, Bryologist 109 (1): 61, 2006.

*** Tritomaria exsectiformis (Breidl.) Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909. Bas.: Jungermannia exsectiformis Breidl., Mitt. Naturwiss. Vereins Steiermark 30: 321, 1894.

** Tritomaria exsectiformis subsp. arctica R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 661, 1969.

** Tritomaria exsectiformis subsp. camerunensis S.W.Arnell ex Váňa, Phytotaxa 167 (2): 216, 2014. Based on: Tritomaria camerunensis S.W.Arnell, Svensk Bot. Tidskr. 52 (1): 64, 1958, nom. inval.

*** Tritomaria ferruginea (Grolle) Váňa, Phytotaxa 81 (1): 24, 2013. Bas.: Andrewsianthus ferrugineus Grolle, Khumbu Himal 1 (4): 275, 1966.

* Tritomaria koreana Bakalin, S.S.Choi et B.Y.Sun, Arctoa 18: 163, 2009 [2010].

* Tritomaria mexicana Bakalin, Arctoa 17: 162, 2008 [2009].

*** Tritomaria scitula (Taylor) Jørg., Bergens Mus. Aarbok 1919/20 (7): 3, 1922. Bas.: Jungermannia scitula Taylor, London J. Bot. 5: 274, 1846.

*** Scapaniaceae Mig.

by J. Váňa

Scapaniaceae, in its classical concept, was recognized as monophyletic and nested within Lophoziaceae s. lat. (cf.,,, ). More recent molecular studies classified Scapaniaceae as a sister clade to Lophoziaceae, and the genus Schistochilopsis, a segregate of the genus Lophozia s. lat., more closely related to Scapaniaceae than to Lophoziaceae (cf., Vilnet et al. 2007). The genus Scapania was recently studied by morphological methods by, by molecular methods by Vilnet et al. (2007) and especially. Some taxonomic and nomenclatural notes can be found in,, ) and. The placement of Pseudotritomaria, Saccobasis and Schistochilopsis in the family follows.

*** Diplophyllum (Dumort.) Dumort., Recueil Observ. Jungerm.: 15, 1835 nom. conserv. Bas.: Jungermannia sect. Diplophyllum Dumort., Syll. Jungerm. Europ.: 44, 1831.

*** subg. Austrodiplophyllum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 18, 1968.

** Diplophyllum recurvifolium C.Massal., Atti Reale Ist. Veneto Sci. Lett. Arti 87 (2): 221, 1928.

*** Diplophyllum squarrosum Steph., Sp. Hepat. (Stephani) 4: 116, 1910.

*** Diplophyllum verrucosum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 19, 1968.

*** subg. Diplophyllum

*** sect. Diplophyllum

*** Diplophyllum albicans (L.) Dumort., Recueil Observ. Jungerm.: 16, 1835. Bas.: Jungermannia albicans L., Sp. Pl. 1: 1133, 1753.

*** sect. Protodiplophyllum (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 29, 2013. Bas.: Diplophyllum subg. Protodiplophyllum R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 192, 1974.

*** Diplophyllum africanum S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 527, 1956.

*** Diplophyllum andicola R.M.Schust., Phytologia 39 (4): 248, 1978.

*** Diplophyllum andrewsii A.Evans, Bryologist 25 (2): 28, 1922.

* Diplophyllum androgynum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 277, 1998.

* Diplophyllum angustifolium J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 262, 1998.

*** Diplophyllum apiculatum (A.Evans) Steph., Sp. Hepat. (Stephani) 4: 110, 1910. Bas.: Diplophylleia apiculata A.Evans, Bot. Gaz. 34 (5): 372, 1902.

** Diplophyllum dioicum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 20, 1968.

** Diplophyllum exiguum Steph., Sp. Hepat. (Stephani) 6: 500, 1924.

* Diplophyllum gemmiparum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 255, 1998.

* Diplophyllum incrassatum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 265, 1998.

*** Diplophyllum nanum Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 48, 1950 [1951].

* Diplophyllum novum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 274, 1998.

*** Diplophyllum obtusatum (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 215, 1974. Bas.: Diplophyllum apiculatum var. obtusatum R.M.Schust., Amer. Midl. Naturalist 49 (2): 432, 1953.

*** Diplophyllum obtusifolium (Hook.) Dumort., Recueil Observ. Jungerm.: 16, 1835. Bas.: Jungermannia obtusifolia Hook., Brit. Jungermann.: tab. 26, 1812. 59

** Diplophyllum obtusifolium subsp. domesticum (Gottsche) Váňa, Phytotaxa 76 (3): 29, 2013. Bas.: Jungermannia domestica Gottsche, Linnaea 28 (5): 548, 1856 [1857].

*** Diplophyllum serrulatum (Müll.Frib.) Steph., Sp. Hepat. (Stephani) 4: 112, 1910. Bas.: Diplophylleia serrulata Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 34, 1903.

*** Diplophyllum taxifolium (Wahlenb.) Dumort., Recueil Observ. Jungerm.: 16, 1835. Bas.: Jungermannia taxifolia Wahlenb., Fl. Lapp. (Wahlenberg): 389, 1812.

** Diplophyllum taxifolium var. mucronatum R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 203, 1974.

*** Diplophyllum trollii Grolle, Khumbu Himal 1 (4): 273, 1966.

*** Douinia (C.E.O.Jensen) H.Buch, Scapan. N.-Eur. Sib.: 13, 1928. Bas.: Diplophylleia subg. Douinia C.E.O.Jensen, Danmarks mosser: 145, 1915.

*** Douinia imbricata (M.Howe) Konstant. et Vilnet, Phytotaxa 76 (3): 31, 2013. Bas.: Scapania imbricata M.Howe, Bull. New York Bot. Gard. 2 (6): 104, 1901.

*** Douinia ovata (Dicks.) H.Buch, Scapan. N.-Eur. Sib.: 14, 1928. Bas.: Jungermannia ovata Dicks., Fasc. Pl. Crypt. Brit. 3: 11, 1793.

*** Douinia plicata (Lindb.) Konstant. et Vilnet, Phytotaxa 76 (3): 31, 2013. Bas.: Diplophyllum plicatum Lindb., Acta Soc. Sci. Fenn. 10: 235, 1872 [1873].

** Pseudotritomaria Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010].

*** Pseudotritomaria heterophylla (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010]. Bas.: Tritomaria heterophylla R.M.Schust., Canad. J. Bot. 36 (2): 272, 1958.

** Saccobasis H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 291, 1932 [1933].

*** Saccobasis polita (Nees) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 292, 1932 [1933]. Bas.: Jungermannia polita Nees, Naturgesch. Eur. Leberm. 2: 145, 1836.

** Saccobasis polymorpha (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 205, 1979. Bas.: Tritomaria polita subsp. polymorpha R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 700, 1969.

*** Scapania (Dumort.) Dumort., Recueil Observ. Jungerm.: 14, 1835 nom. conserv. Bas.: Radula sect. Scapania Dumort., Syll. Jungerm. Europ.: 38, 1831.

** subg. Ascapania Grolle, Khumbu Himal 1 (4): 268, 1966.

*** Scapania contorta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 101, 1860 [1861].

** subg. Gracilidae (H.Buch) Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012. Bas.: Scapania sect. Gracilidae H.Buch, Scapan. N.-Eur. Sib.: 106, 1928.

*** Scapania ampliata Steph., Bull. Herb. Boissier 5 (2): 106, 1897.

** Scapania ampliata subsp. queenslandica M.L.Hicks, J. Hattori Bot. Lab. 69: 130, 1991.

*** Scapania bolanderi Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 218, 1869.

*** Scapania gracilis Lindb., Morgonbladet (Helsinki) 1873 (286, 9 Dec): 2, 1873.

** Scapania macroparaphyllia T.Cao, C.Gao et J.Sun, Acta Phytotax. Sin. 42 (2): 180, 2004.

*** Scapania maxima Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 223, 1934.

*** Scapania nipponica (Amakawa et S.Hatt.) Amakawa, J. Hattori Bot. Lab. 30: 319, 1967. Bas.: Scapania bolanderi var. nipponica Amakawa et S.Hatt., J. Hattori Bot. Lab. 14: 83, 1955.

** Scapania paraphyllia T.Cao, C.Gao, J.Sun et B.R.Zuo, Acta Phytotax. Sin. 45 (3): 311, 2007.

*** Scapania subnimbosa Steph., Sp. Hepat. (Stephani) 4: 150, 1910.

** subg. Macroscapania R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 248, 1974.

* Scapania geppii Steph., Hedwigia 44 (1): 14, 1904.

*** Scapania portoricensis Hampe et Gottsche, Linnaea 25 (3): 342, 1852 [1853].

** Scapania portoricensis var. boliviensis (Steph.) Herzog, Ann. Bryol. 1: 110, 1928. Bas.: Scapania boliviensis Steph., Biblioth. Bot. 87 (2): 231, 1916.

** Scapania portoricensis var. organensis (Herzog) Herzog, Ann. Bryol. 1: 110, 1928. Bas.: Scapania organensis Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 27, 1925.

** Scapania portoricensis var. roraimensis Warnst., Hedwigia 63 (2): 109, 1921.

** subg. Plicaticalyx Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 36, 1903.

** sect. Planifoliae (Müll.Frib.) Potemkin, J. Hattori Bot. Lab. 85: 56, 1998. Bas.: Scapania Gruppe Planifoliae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 286, 1905.

*** Scapania davidii Potemkin, Ann. Bot. Fenn. 38 (2): 83, 2001.

** Scapania ferruginaeoides T.Cao, C.Gao et J.Sun, Guihaia 24 (1): 23, 2004.

** Scapania gaochii X.Fu ex T.Cao, Phytotaxa 97 (2): 26, 2013. Based on: Scapania gaochii X.Fu ex T.Cao, Acta Bot. Yunnan. 25 (5): 541, 2003, nom. inval.

*** Scapania harae Amakawa, J. Hattori Bot. Lab. 27: 5, 1964.

*** Scapania nimbosa Taylor, Nov. Stirp. Pug. 8: 6, 1844.

*** Scapania ornithopodioides (With.) Waddell, Moss Exch. Club Cat. Hepat.: 4, 1897. Bas.: Jungermannia ornithopodioides With., Bot. arr. veg. Gr. Brit. 2: 695, 1776.

*** Scapania rotundifolia W.E.Nicholson, Symb. Sin. 5: 31, 1930.

*** Scapania secunda Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 226, 1894.

*** Scapania zhukovae Potemkin, Arctoa 9: 121, 2000.

** sect. Plicaticalyx (Müll.Frib.) Potemkin, Ann. Bot. Fenn. 39 (4): 326, 2002. Bas.: Scapania subg. Plicaticalyx Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 36, 1903.

*** Scapania ciliatospinosa Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 222, 1934.

*** Scapania ferruginea (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 1: 72, 1844. Bas.: Jungermannia ferruginea Lehm. et Lindenb., Nov. Stirp. Pug. 4: 20, 1832.

*** Scapania hians Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 223, 1905.

** Scapania hians subsp. salishensis J.D.Godfrey et G.Godfrey, Bryologist 81 (3): 362, 1978.

*** Scapania orientalis Steph. ex Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 606, 1901.

*** Scapania pseudocontorta Potemkin, Arctoa 9: 115, 2000.

*** Scapania sinikkae Potemkin, Ann. Bot. Fenn. 38 (2): 85, 2001.

*** Scapania spiniloba Potemkin, Arctoa 9: 117, 2000.

** subg. Pseudomacrodiplophyllum Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012.

*** Scapania microdonta (Mitt.) Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 262, 1905. Bas.: Martinellius microdontus Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 196, 1891.

*** subg. Scapania

** sect. Aequilobae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 110, 1928. Bas.: Scapania Gruppe Aequilobae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 219, 1905.

*** Scapania aequiloba (Schwägr.) Dumort., Recueil Observ. Jungerm.: 14, 1835. Bas.: Jungermannia aequiloba Schwägr., Hist. Musc. Hepat. Prodr.: 24, 1814.

*** Scapania aspera M.Bernet et Bernet, Cat. hép. Suisse: 42, 1888.

** sect. Americanae Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012.

*** Scapania americana Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 44, 1903.

** sect. Apiculatae H.Buch, Scapan. N.-Eur. Sib.: 53, 1928.

*** Scapania apiculata Spruce, Hep. Pyr. Exsic.: no. 15, 1847.

*** Scapania carinthiaca J.B.Jack ex Lindb., Rev. Bryol. 7 (4): 77, 1880.

*** Scapania carinthiaca var. massalongi Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 598, 1901.

*** Scapania umbrosa (Schrad.) Dumort., Recueil Observ. Jungerm.: 14, 1835. Bas.: Jungermannia umbrosa Schrad., Syst. Samml. Crypt. Gew. 2: 5, 1797.

** sect. Ciliatae Grolle, Khumbu Himal 1 (4): 272, 1966.

*** Scapania bhutanensis Amakawa, Fl. E. Himalaya 2: 230, 1971.

*** Scapania ciliata Sande Lac., Prolus. fl. jap.: 209, 1867.

*** Scapania ciliata subsp. hawaiica (Müll.Frib.) Potemkin, Ann. Bot. Fenn. 39 (4): 321, 2002. Bas.: Scapania hawaiica Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 160, 1905.

*** Scapania hirosakiensis Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 120, 1905.

*** Scapania hollandiae W.S.Hong, Bryologist 83 (1): 56, 1980.

*** Scapania koponenii Potemkin, Ann. Bot. Fenn. 37 (1): 41, 2000.

*** Scapania lepida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 101, 1860 [1861].

*** Scapania sandei Schiffn. ex Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 612, 1901.

** sect. Compactae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 101, 1928. Bas.: Scapania Gruppe Compactae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 53, 1905.

*** Scapania compacta (Roth) Dumort., Recueil Observ. Jungerm.: 14, 1835. Bas.: Jungermannia compacta Roth, Tent. Fl. Germ. 3: 375, 1800.

*** Scapania kaurinii Ryan, Bot. Not. 42: 210, 1889.

*** Scapania spitsbergensis (Lindb.) Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 607, 1901. Bas.: Martinellius spitsbergensis Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 31, 1889.

** sect. Curtae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 55, 1928. Bas.: Scapania Gruppe Curtae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 245, 1905.

*** Scapania curta (Mart.) Dumort., Recueil Observ. Jungerm.: 14, 1835. Bas.: Jungermannia curta Mart., Fl. crypt. erlang.: 148, 1817.

** Scapania curta var. grandiretis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 393, 1974.

** Scapania curta var. isoloba R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 396, 1974.

*** Scapania diplophylloides Amakawa et S.Hatt., J. Hattori Bot. Lab. 9: 59, 1953.

*** Scapania esterhuyseniae S.W.Arnell, Bot. Not. 110 (1): 26, 1957.

*** Scapania fulfordiae W.S.Hong, Bryologist 83 (1): 46, 1980.

*** Scapania gamundiae R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 14, 1968.

*** Scapania helvetica Gottsche, Hepat. Eur., Leberm. 42-44: no. 426, 1868.

*** Scapania irrigua (Nees) Nees, Syn. Hepat. 1: 67, 1844. Bas.: Jungermannia irrigua Nees, Naturgesch. Eur. Leberm. 1: 193, 1833.

** Scapania irrigua subsp. rufescens (Loeske) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 471, 1974. Bas.: Scapania irrigua f. rufescens Loeske, Moosfl. Harz.: 71, 1903.

*** Scapania lingulata H.Buch, Meddel. Soc. Fauna Fl. Fenn. 42: 92, 1916.

** Scapania lingulata var. microphylla (Warnst.) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 415, 1974. Bas.: Scapania microphylla Warnst., Hedwigia 63 (2): 75, 1921.

** Scapania magadanica S.S.Choi, Bakalin et B.Y.Sun, Bot. Pacifica 1: 46, 2012.

*** Scapania mucronata H.Buch, Meddel. Soc. Fauna Fl. Fenn. 42: 91, 1916.

*** Scapania obcordata (Berggr.) S.W.Arnell, Ark. Bot. (n.ser.) 4 (6): 117, 1959. Bas.: Sarcocyphos obcordatus Berggr., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 13 (7): 96, 1875.

** Scapania parvifolia Warnst., Hedwigia 63 (2): 78, 1921.

*** Scapania praetervisa Meyl., Jahresber. Naturf. Ges. Graubündens (n.f.) 64: 364, 1926.

*** Scapania scandica (Arnell et H.Buch) Macvicar, Stud. handb. Brit. hepat. (ed. 2): 394, 1926. Bas.: Martinellius scandicus Arnell et H.Buch, Bot. Not. 74: 1, 1921.

** Scapania scandica var. argutedentata H.Buch, Scapan. N.-Eur. Sib.: 75, 1928.

** Scapania scandica var. dimorpha R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 453, 1974.

** Scapania scandica var. grandiretis (Schljakov) Schljakov, Pečen. Mchi Sev. SSSR 4: 152, 1981. Bas.: Scapania parvifolia var. grandiretis Schljakov, Novosti Sist. Nizš. Rast. 8: 332, 1971.

*** Scapania uliginosa (Lindenb.) Dumort., Recueil Observ. Jungerm.: 14, 1835. Bas.: Jungermannia undulata var. uliginosa Lindenb., Syn. hepat. eur: 58, 1829.

*** Scapania valdonii Váňa, Bedn.-Ochyra et Cykowska, Nova Hedwigia 89 (1/2): 126, 2009.

*** Scapania zemliae S.W.Arnell, Svensk Bot. Tidskr. 41: 215, 1947.

** sect. Cuspiduligerae H.Buch, Scapan. N.-Eur. Sib.: 125, 1928.

*** Scapania cuspiduligera (Nees) Müll.Frib., Lebermoose 2 (22): 472, 1915. Bas.: Jungermannia cuspiduligera Nees, Naturgesch. Eur. Leberm. 1: 180, 1833.

** Scapania cuspiduligera var. diplophyllopsis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 361, 1974.

** sect. Grolleoscapania Potemkin, Ann. Bot. Fenn. 39 (4): 329, 2002.

*** Scapania karl-muelleri Grolle, Khumbu Himal 1 (4): 270, 1966.

** sect. Hyperboreae Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 16, 2012.

*** Scapania hyperborea Jørg., Forh. Vidensk.-Selsk. Kristiania 1894 (8): 56, 1894.

*** Scapania paludicola Loeske et Müll.Frib., Lebermoose 2 (21): 425, 1915.

** Scapania paludicola var. viridigemma R.M.Schust., Bull. Natl. Mus. Canada 122: 20, 1950 [1951].

*** Scapania tundrae (Arnell) H.Buch, Scapan. N.-Eur. Sib.: 99, 1928. Bas.: Martinellius tundrae Arnell, Bot. Not. 74: 289, 1921.

** sect. Kaalaasia (H.Buch) Jørg., Bergens Mus. Skr. (n.ser.) 16: 210, 1934. Bas.: Scapania subg. Kaalaasia H.Buch, Scapan. N.-Eur. Sib.: 47, 1928.

*** Scapania calcicola (Arnell et J.Perss.) Ingham, Naturalist (Hull) 564: 11, 1904. Bas.: Martinellius calcicola Arnell et J.Perss., Rev. Bryol. 30 (6): 97, 1903.

*** Scapania gymnostomophila Kaal., Bot. Not. 49: 21, 1896.

*** Scapania ligulifolia R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 306, 1974.

*** Scapania pseudocalcicola R.M.Schust., Phytologia 63 (5): 327, 1987.

* Scapania sect. Muelleria Potemkin, Ann. Bot. Fenn. 39 (4): 320, 2002.

*** Scapania himalayica Müll.Frib. ex Herzog, Ann. Bryol. 12: 81, 1939.

*** Scapania schljakovii Potemkin, Ann. Bot. Fenn. 38 (2): 87, 2001.

** sect. Nemorosae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 152, 1928. Bas.: Scapania Gruppe Nemorosae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 155, 1905.

*** Scapania brevicaulis Taylor, London J. Bot. 5: 272, 1846.

*** Scapania crassiretis Bryhn, Rev. Bryol. 19 (1): 7, 1892.

* Scapania degenii Schiffn. ex Müll.Frib., Lebermoose 2 (22): 497, 1915. 60

* Scapania glaucoviridis Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 221, 1934. 61

** Scapania grossidens Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 146, 1905.

*** Scapania hedbergii S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 556, 1956.

*** Scapania integerrima Steph., Sp. Hepat. (Stephani) 4: 148, 1910.

*** Scapania matveyevae Potemkin, Arctoa 9: 97, 2000.

*** Scapania nemorea (L.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 160, 1963 [1964]. Bas.: Jungermannia nemorea L., Syst. Nat., ed. 10., 2: 1337, 1759.

** Scapania parvidens Steph., Hedwigia 44 (1): 15, 1904. 62

*** Scapania parvitexta Steph., Bull. Herb. Boissier 5 (2): 107, 1897.

*** Scapania rigida Nees, Syn. Hepat. 1: 69, 1844.

** sect. Scapania

*** Scapania gigantea Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 15, 1931.

** Scapania grandiloba Steph., Sp. Hepat. (Stephani) 6: 502, 1924.

*** Scapania komagadakensis Amakawa, J. Hattori Bot. Lab. 31: 96, 1968.

*** Scapania obscura (Arnell et C.E.O.Jensen) Schiffn., Österr. Bot. Z. 58 (10): 377, 1908. Bas.: Martinellius obscurus Arnell et C.E.O.Jensen, Moose Sarekgeb.: 91, 1907.

** Scapania paludosa (Müll.Frib.) Müll.Frib., Mitt. Bad. Bot. Vereins 4 (182/183): 287, 1902. Bas.: Scapania undulata var. paludosa Müll.Frib., Beih. Bot. Centralbl. 10 (4/5): 220, 1901.

*** Scapania rufidula Warnst., Hedwigia 63 (2): 94, 1921.

*** Scapania serrulata R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 539, 1974.

*** Scapania subalpina (Nees ex Lindenb.) Dumort., Recueil Observ. Jungerm.: 14, 1835. Bas.: Jungermannia subalpina Nees ex Lindenb., Syn. hepat. eur: 55, 1829.

** Scapania subalpina var. haynesiae Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (4): 638, 1946.

** Scapania subalpina var. muddiae C.D.Bird et W.S.Hong, Bryologist 83 (1): 51, 1980.

*** Scapania undulata (L.) Dumort., Recueil Observ. Jungerm.: 14, 1835. Bas.: Jungermannia undulata L., Sp. Pl. 1: 1132, 1753.

** sect. Scapaniella (H.Buch) Potemkin, J. Hattori Bot. Lab. 85: 43, 1998. Bas.: Scapaniella H.Buch, Scapan. N.-Eur. Sib.: 33, 1928.

*** Scapania glaucocephala (Taylor) Austin, Bull. Torrey Bot. Club 6 (16): 85, 1876. Bas.: Jungermannia glaucocephala Taylor, London J. Bot. 5: 277, 1846.

*** Scapania glaucocephala var. saxicola (R.M.Schust.) Potemkin, Bryologist 102 (1): 36, 1999. Bas.: Scapania saxicola R.M.Schust., Amer. Midl. Naturalist 49 (2): 448, 1953.

* Scapania scapanioides (C.Massal.) Grolle, Feddes Repert. 87 (3/4): 235, 1976. Bas.: Jungermannia scapanioides C.Massal., Hepaticol. ven.: 64, 1879. 63

** sect. Simmonsiae (R.M.Schust.) Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 16, 2012. Bas.: Scapania subsect. Simmonsiae R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 612, 1974.

*** Scapania simmonsii Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 51, 1906.

** sect. Sphaeriferae Konstant. et Potemkin, Ann. Bot. Fenn. 31 (2): 125, 1994.

*** Scapania sphaerifera H.Buch et Tuom., Memoranda Soc. Fauna Fl. Fennica 11: 227, 1936.

** sect. Stephaniae Potemkin, J. Hattori Bot. Lab. 85: 57, 1998. Based on: Scapania sect. Stephaniae Amakawa et S.Hatt., J. Hattori Bot. Lab. 12: 94, 1954. nom. inval.

*** Scapania griffithii Schiffn., Österr. Bot. Z. 49 (6): 204, 1899.

*** Scapania javanica Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 575, 1853.

* Scapania javanica var. scabra Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 397, 1955.

*** Scapania ligulata Steph., Hedwigia 44 (1): 14, 1904.

** Scapania ligulata subsp. stephanii (Müll.Frib.) Potemkin, Piippo et T.J.Kop., Ann. Bot. Fenn. 41 (6): 423, 2004. Bas.: Scapania stephanii Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 273, 1905.

** sect. Verrucosae Potemkin, J. Hattori Bot. Lab. 85: 54, 1998.

*** Scapania udarii S.C.Srivast. et A.Srivast., J. Indian Bot. Soc. 72: 237, 1993.

*** Scapania verrucosa Heeg, Rev. Bryol. 20 (5): 81, 1893.

*** Schistochilopsis (N.Kitag.) Konstant., Arctoa 3: 125, 1994. Bas.: Lophozia subg. Schistochilopsis N.Kitag., J. Hattori Bot. Lab. 28: 289, 1965.

*** Schistochilopsis cornuta (Steph.) Konstant., Arctoa 3: 125, 1994. Bas.: Schistochila cornuta Steph., Sp. Hepat. (Stephani) 4: 84, 1909.

*** Schistochilopsis grandiretis (Lindb. ex Kaal.) Konstant., Arctoa 3: 125, 1994. Bas.: Jungermannia grandiretis Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 322, 1893.

** Schistochilopsis hyperarctica Konstant. et L.Söderstr., Phytotaxa 162 (4): 240, 2014. Based on: Lophozia hyperarctica R.M.Schust., Canad. J. Bot. 39 (4): 967, 1961, nom. inval.

*** Schistochilopsis incisa (Schrad.) Konstant., Arctoa 3: 125, 1994. Bas.: Jungermannia incisa Schrad., Syst. Samml. Crypt. Gew. 2: 5, 1797.

* Schistochilopsis nakanishii (Inoue) Konstant., Arctoa 3: 125, 1994. Bas.: Lophozia nakanishii Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 9 (1): 37, 1966. 64

** Schistochilopsis opacifolia (Culm. ex Meyl.) Konstant., Arctoa 3: 125, 1994. Bas.: Lophozia opacifolia Culm. ex Meyl., Beitr. Kryptogamenfl. Schweiz 6 (4): 174, 1924. 65

*** Schistochilopsis setosa (Mitt.) Konstant., Arctoa 3: 125, 1994. Bas.: Jungermannia setosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 92, 1860 [1861].

*** Acrobolbaceae E.A.Hodgs.

by L. Briscoe and J.J. Engel

** Enigmella G.A.M.Scott et K.G.Beckm., J. Bryol. 17 (2): 297, 1992.

** Enigmella thallina G.A.M.Scott et K.G.Beckm., J. Bryol. 17 (2): 297, 1992.

** Acrobolboideae R.M.Schust. ex Briscoe

*** Acrobolbus Nees, Syn. Hepat. 1: 5, 1844.

** Acrobolbus africanus (Pearson) Briscoe, Phytotaxa 202 (1): 59, 2015. Bas.: Tylimanthus africanus Pearson, Forh. Vidensk.-Selsk. Kristiania 1887 (9): 14, 1887.

*** Acrobolbus anisodontus (Hook.f. et Taylor) Briscoe, Phytotaxa 202 (1): 59, 2015. Bas.: Jungermannia anisodonta Hook.f. et Taylor, London J. Bot. 4: 79, 1845.

** Acrobolbus antillanus R.M.Schust., J. Hattori Bot. Lab. 90: 143, 2001.

** Acrobolbus azoricus (Grolle et Perss.) Briscoe, Phytotaxa 202 (1): 59, 2015. Bas.: Tylimanthus azoricus Grolle et Perss., Svensk Bot. Tidskr. 60 (1): 169, 1966.

*** Acrobolbus caducifolius R.M.Schust., J. Hattori Bot. Lab. 90: 154, 2001.

*** Acrobolbus ciliatus (Mitt.) Schiffn., Hepat. (Engl.-Prantl): 86, 1893. Bas.: Gymnanthe ciliata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 100, 1860 [1861].

*** Acrobolbus cinerascens (Lehm. et Lindenb.) Bastow, Pap. & Proc. Roy. Soc. Tasmania 1887: 242, 1888. Bas.: Jungermannia cinerascens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 46, 1832.

*** Acrobolbus concinnus (Mitt.) Steph., Trans. & Proc. New Zealand Inst. 24: 399, 1892. Bas.: Gymnanthe concinna Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 230, 1860.

** Acrobolbus cuneifolius (Steph.) Briscoe, Phytotaxa 202 (1): 59, 2015. Bas.: Tylimanthus cuneifolius Steph., Bull. Herb. Boissier (sér. 2) 5 (12): 1138 (10), 1905.

*** Acrobolbus diversifolius R.M.Schust., J. Hattori Bot. Lab. 90: 150, 2001.

*** Acrobolbus epiphytus (Colenso) Briscoe, Phytotaxa 202 (1): 59, 2015. Bas.: Marsupidium epiphytum Colenso, Trans. & Proc. New Zealand Inst. 21: 64, 1889.

*** Acrobolbus flavicans (J.J.Engel et Grolle) Briscoe et J.J.Engel, Phytotaxa 202 (1): 59, 2015. Bas.: Marsupidium flavicans J.J.Engel et Grolle, J. Hattori Bot. Lab. 34: 438, 1971.

*** Acrobolbus gradsteinii (Grolle) Briscoe, Phytotaxa 202 (1): 59, 2015. Bas.: Marsupidium gradsteinii Grolle, J. Hattori Bot. Lab. 66: 337, 1989.

** Acrobolbus integrifolius (A.Evans) Briscoe, Phytotaxa 202 (1): 59, 2015. Bas.: Tylimanthus integrifolius A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891.

*** Acrobolbus knightii (Mitt.) Briscoe, Phytotaxa 202 (1): 59, 2015. Bas.: Marsupidium knightii Mitt., Handb. N. Zeal. fl. 2: 753, 1867.

** Acrobolbus kunkelii (Hässel et Solari) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015. Bas.: Tylimanthus kunkelii Hässel et Solari, Darwiniana 17: 574, 1972.

*** Acrobolbus laxus (Lehm. et Lindenb.) Briscoe, Phytotaxa 202 (1): 60, 2015. Bas.: Plagiochila laxa Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 68, 1840.

*** Acrobolbus limbatus (Steph.) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015. Bas.: Tylimanthus limbatus Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 25, 1911.

*** Acrobolbus lophocoleoides (Mitt.) Mitt., Handb. N. Zeal. fl. 2: 753, 1867. Bas.: Gymnanthe lophocoleoides Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 144, 1854.

** Acrobolbus madeirensis (Grolle et Perss.) Briscoe, Phytotaxa 202 (1): 60, 2015. Bas.: Tylimanthus madeirensis Grolle et Perss., Svensk Bot. Tidskr. 60 (1): 166, 1966.

** Acrobolbus mittenii Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 460 (179), 1902.

*** Acrobolbus ochrophyllus (Hook.f. et Taylor) R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 64, 1961. Bas.: Jungermannia ochrophylla Hook.f. et Taylor, London J. Bot. 3: 368, 1844.

*** Acrobolbus papillosus (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015. Bas.: Marsupidium papillosum J.J.Engel et Glenny, Nova Hedwigia 87 (3/4): 289, 2008.

*** Acrobolbus perpusillus (Colenso) Briscoe, Phytotaxa 202 (1): 60, 2015. Bas.: Tylimanthus perpusillus Colenso, Trans. & Proc. New Zealand Inst. 19: 286, 1887.

*** Acrobolbus perpusillus var. denticulatus (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015. Bas.: Marsupidium perpusillum var. denticulatum J.J.Engel et Glenny, Nova Hedwigia 87 (3/4): 284, 2008.

*** Acrobolbus plagiochiloides (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015. Bas.: Marsupidium plagiochiloides J.J.Engel et Glenny, Nova Hedwigia 87 (3/4): 284, 2008.

*** Acrobolbus pseudosaccatus (Grolle) Briscoe, Phytotaxa 202 (1): 60, 2015. Bas.: Tylimanthus pseudosaccatus Grolle, Nova Hedwigia 6 (3/4): 391, 1963.

*** Acrobolbus renifolius (Hässel et Solari) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015. Bas.: Tylimanthus renifolius Hässel et Solari, Darwiniana 17: 583, 1972.

** Acrobolbus ruwenzorensis (S.W.Arnell) Briscoe, Phytotaxa 202 (1): 60, 2015. Bas.: Tylimanthus ruwenzorensis S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 560, 1956.

*** Acrobolbus saccatus (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877. Bas.: Jungermannia saccata Hook., Musci Exot. 1: tab. 16, 1818.

*** Acrobolbus setulosus (Mitt.) Briscoe, Phytotaxa 202 (1): 60, 2015. Bas.: Gymnanthe setulosa Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 144, 1854.

*** Acrobolbus spinifolius R.M.Schust., J. Hattori Bot. Lab. 90: 137, 2001.

*** Acrobolbus sumatranus (Schiffn.) Briscoe, Phytotaxa 202 (1): 61, 2015. Bas.: Lophozia sumatrana Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 203, 1898.

*** Acrobolbus surculosus (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877. Bas.: Scapania surculosa Nees, Syn. Hepat. 1: 62, 1844.

*** Acrobolbus tenellus (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877. Bas.: Gymnanthe tenella Taylor, Nov. Stirp. Pug. 8: 1, 1844.

*** Acrobolbus tenellus var. diversifolius (E.A.Hodgs.) Briscoe, Phytotaxa 202 (1): 61, 2015. Bas.: Tylimanthus diversifolius E.A.Hodgs., Trans. Roy. Soc. New Zealand 85 (4): 575, 1958.

*** Acrobolbus urvilleanus (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877. Bas.: Plagiochila urvilleana Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 247, 1843.

*** Acrobolbus viridis (Mitt.) Briscoe et J.J.Engel, Phytotaxa 202 (1): 61, 2015. Bas.: Tylimanthus viridis Mitt., J. Linn. Soc., Bot. 15 (84): 197, 1876.

*** Acrobolbus wilsonii Nees, Syn. Hepat. 1: 5, 1844.

** Acrobolbus wilsonii var. andinus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 522, 1885.

Excluded from the genus

* Acrobolbus bispinosus (J.B.Jack et Steph.) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 459 (178), 1902. Bas.: Tylimanthus bispinosus J.B.Jack et Steph., Hedwigia 31 (1): 26, 1892. 66

** Austrolophozioideae R.M.Schust. ex Crand.-Stotl., Váňa et Stotler

** Austrolophozia R.M.Schust., J. Hattori Bot. Lab. 26: 282, 1963.

** Austrolophozia andina R.M.Schust., Nova Hedwigia 15: 495, 1968.

*** Austrolophozia camensis (Steph.) Grolle ex Hässel et Solari, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 3 (6): 240, 1970. Bas.: Tylimanthus camensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 24, 1911.

*** Austrolophozia paradoxa R.M.Schust., J. Hattori Bot. Lab. 26: 282, 1963.

*** Goebelobryum Grolle, J. Hattori Bot. Lab. 25: 135, 1962.

*** Goebelobryum grossitextum (Steph.) Grolle, J. Hattori Bot. Lab. 25: 137, 1962. Bas.: Marsupidium grossitextum Steph., Sp. Hepat. (Stephani) 6: 446, 1924.

*** Goebelobryum unguiculatum (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 25: 137, 1962. Bas.: Jungermannia unguiculata Hook.f. et Taylor, London J. Bot. 5: 279, 1846.

*** Goebelobryum vermiculare J.J.Engel et Glenny, Nova Hedwigia 95 (3/4): 320, 2012.

** Lethocoleoideae Grolle

*** Lethocolea Mitt., Handb. N. Zeal. fl. 2: 751, 1867 nom. conserv.

** Lethocolea congesta (Lehm.) S.W.Arnell, Bot. Not. 108: 311, 1955. Bas.: Jungermannia congesta Lehm., Linnaea 4: 365, 1829.

*** Lethocolea glossophylla (Spruce) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965. Bas.: Symphyomitra glossophylla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 503, 1885.

** Lethocolea indica G.Asthana et Maurya, Natl. Acad. Sci. Lett. 37 (6): 535, 2014.

*** Lethocolea javanica (Schiffn.) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965. Bas.: Symphyomitra javanica Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 193, 1898.

** Lethocolea naruto-toganensis Furuki, Bryologist 104 (2): 306, 2001.

*** Lethocolea pansa (Taylor) G.A.M.Scott et K.G.Beckm., Symp. Biol. Hung. 35: 212, 1987. Bas.: Jungermannia pansa Taylor, London J. Bot. 5: 275, 1846.

*** Lethocolea radicosa (Lehm. et Lindenb.) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965. Bas.: Jungermannia radicosa Lehm. et Lindenb., Nov. Stirp. Pug. 6: 35, 1834.

* Lethocolea repens S.Winkl., Mitt. Inst. Colombo-Alemán Invest. Ci. 3: 67, 1969.

** Saccogynidioideae Crand.-Stotl., Váňa et Stotler

*** Saccogynidium Grolle, J. Hattori Bot. Lab. 23: 43, 1960 [1961].

** sect. Decurvum Grolle, J. Hattori Bot. Lab. 23: 59, 1960.

*** Saccogynidium decurvum (Mitt.) Grolle, J. Hattori Bot. Lab. 23: 59, 1960 [1961]. Bas.: Lophocolea decurva Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 227, 1860.

** sect. Jugata Grolle, J. Hattori Bot. Lab. 23: 55, 1960.

** Saccogynidium rigidulum (Nees) Grolle, J. Hattori Bot. Lab. 23: 52, 1960 [1961]. Bas.: Jungermannia rigidula Nees, Enum. Pl. Crypt. Javae: 25, 1830.

** sect. Saccogynidium

*** Saccogynidium australe (Mitt.) Grolle, J. Hattori Bot. Lab. 23: 49, 1960 [1961]. Bas.: Saccogyna australis Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 145, 1854.

** Saccogynidium caldense (Ångstr.) Grolle, J. Hattori Bot. Lab. 23: 44, 1960 [1961]. Bas.: Chiloscyphus caldensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 80, 1876 [1877].

** Saccogynidium goebelii (Herzog) Grolle, J. Hattori Bot. Lab. 23: 52, 1960 [1961]. Bas.: Leioscyphus goebelii Herzog, Ann. Bryol. 5: 89, 1932.

** Saccogynidium muricellum (De Not.) Grolle, J. Hattori Bot. Lab. 36: 80, 1972 [1973]. Bas.: Chiloscyphus muricellus De Not., Epat. Borneo: 24, 1874.

*** Saccogynidium vasculosum (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 23: 46, 1960 [1961]. Bas.: Jungermannia vasculosa Hook.f. et Taylor, London J. Bot. 3: 461, 1844.

Incertae sedis

** Saccogynidium chiloscyphoides R.M.Schust., J. Hattori Bot. Lab. 26: 272, 1963.

** Saccogynidium irregularospinum C.Gao, T.Cao et M.J.Lai, Bryologist 104 (1): 129, 2001.

*** Antheliaceae R.M.Schust.

*** Anthelia (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835. Bas.: Jungermannia sect. Anthelia Dumort., Syll. Jungerm. Europ.: 63, 1831.

*** Anthelia julacea (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835. Bas.: Jungermannia julacea L., Sp. Pl., ed. 2: 1601, 1763.

** Anthelia juratzkana (Limpr.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877. Bas.: Jungermannia juratzkana Limpr., Hedwigia 15 (2): 18, 1876.

*** Arnelliaceae Nakai

placed Stephaniella and Stephaniellidium in Arnelliaceae following. However, argued based on further molecular evidence that Arnelliaceae should be retained as a monotypic family with a single species.

*** Arnellia Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 35, 1889.

*** Arnellia fennica (Gottsche) Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 35, 1889. Bas.: Jungermannia fennica Gottsche, Hepat. Eur., Leberm. 42-44: no. 418, 1868.

*** Balantiopsidaceae H.Buch

by J.J. Engel with contribution by J. Váňa (Neesioscyphus)

The placement of Pseudoisotachis in Balantiopsidaceae is only preliminary (cf. ).

*** Acroscyphella N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985. Nom. nov. pro Acroscyphus N.Kitag., Acta Phytotax. Geobot. 35 (1/3): 1, 1984.

*** Acroscyphella iwatsukii (N.Kitag.) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985. Bas.: Acroscyphus iwatsukii N.Kitag., Acta Phytotax. Geobot. 35 (1/3): 3, 1984.

*** Acroscyphella phoenicorhiza (Grolle) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985. Bas.: Neesioscyphus phoenicorhizus Grolle, Österr. Bot. Z. 111 (1): 27, 1964.

*** Acroscyphella tjiwideiensis (Sande Lac.) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985. Bas.: Chiloscyphus tjiwideiensis Sande Lac., Ned. Kruidk. Arch. 3: 418, 1854 [1855].

** Pseudoisotachis Váňa, Polish Bot. J. 58 (1): 55, 2013.

** Pseudoisotachis pocsii Váňa, Polish Bot. J. 58 (1): 55, 2013.

*** Balantiopsidoideae J.J.Engel et Váňa

*** Balantiopsis Mitt., Handb. N. Zeal. fl. 2: 751, 1867.

*** Balantiopsis asymmetrica (Herzog) J.J.Engel, Nova Hedwigia 16: 93, 1968. Bas.: Balantiopsis latifolia var. asymmetrica Herzog, Rev. Bryol. Lichénol. 23 (1/2): 53, 1954.

*** Balantiopsis bisbifida (Steph.) Steph., Sp. Hepat. (Stephani) 4: 101, 1910. Bas.: Isotachis bisbifida Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 24, 1901.

*** Balantiopsis brasiliensis Steph., Sp. Hepat. (Stephani) 4: 104, 1910.

*** Balantiopsis cancellata (Nees) Steph., Sp. Hepat. (Stephani) 4: 103, 1910. Bas.: Ptilidium cancellatum Nees, Syn. Hepat. 2: 251, 1845.

*** Balantiopsis ciliaris S.Hatt., J. Jap. Bot. 41 (5): 129, 1966.

** Balantiopsis ciliaris subsp. novoguineensis S.Hatt., J. Jap. Bot. 41 (5): 131, 1966.

*** Balantiopsis convexiuscula Berggr., New Zealand Hepat.: 44, 1898.

*** Balantiopsis crocea Herzog, Beih. Bot. Centralbl. 60B (1/2): 12, 1939.

*** Balantiopsis diplophylla (Hook.f. et Taylor) Mitt., Handb. N. Zeal. fl. 2: 753, 1867. Bas.: Jungermannia diplophylla Hook.f. et Taylor, London J. Bot. 3: 377, 1844.

** Balantiopsis diplophylla var. hockenii (Berggr.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 11, 1997. Bas.: Balantiopsis hockenii Berggr., New Zealand Hepat.: 46, 1898.

*** Balantiopsis erinacea (Hook.f. et Taylor) Mitt., Handb. N. Zeal. fl. 2: 753, 1867. Bas.: Jungermannia erinacea Hook.f. et Taylor, London J. Bot. 3: 462, 1844.

*** Balantiopsis lingulata R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 26, 1968.

*** Balantiopsis montana (Colenso) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 12, 1997. Bas.: Chiloscyphus montanus Colenso, Trans. & Proc. New Zealand Inst. 21: 62, 1889.

** Balantiopsis neocaledonica Pearson, J. Linn. Soc., Bot. 46 (305): 28, 1922.

** Balantiopsis paucidens Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 29, 1901.

*** Balantiopsis purpurata Mitt., Rep. Challenger, Bot. 1 (3, 1): 86, 1884.

*** Balantiopsis rosea Berggr., New Zealand Hepat.: 43, 1898.

*** Balantiopsis splendens (Steph.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 55, 1997. Bas.: Isotachis splendens Steph., Hedwigia 34 (2): 49, 1895.

*** Balantiopsis tumida Berggr., New Zealand Hepat.: 45, 1898.

*** Balantiopsis verrucosa J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 16, 1997.

*** Isotachidoideae Grolle

*** Isotachis Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 148, 1854.

*** Isotachis armata (Nees) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864. Bas.: Jungermannia armata Nees, Syn. Hepat. 1: 129, 1844.

*** Isotachis aubertii (Schwägr.) Mitt., J. Linn. Soc., Bot. 22 (146): 322, 1886. Bas.: Jungermannia aubertii Schwägr., Hist. Musc. Hepat. Prodr.: 19, 1814.

* Isotachis boliviensis Gottsche, Sp. Hepat. (Stephani) 3: 670, 1909.

** Isotachis chinensis C.Gao, T.Cao et J.Sun, Bryologist 105 (4): 694, 2002 [2003].

** Isotachis erythrorhiza (Lehm. et Lindenb.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 338, 1885. Bas.: Jungermannia erythrorhiza Lehm. et Lindenb., Nov. Stirp. Pug. 4: 44, 1832.

*** Isotachis fragilis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 67, 1911.

** Isotachis grandis Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1041, 1888.

*** Isotachis grossidens Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 69, 1911.

** Isotachis hastatistipula (Steph.) J.J.Engel, Phytotaxa 183 (4): 299, 2014. Bas.: Balantiopsis hastatistipula Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 98, 1914.

* Isotachis hians Steph., Sp. Hepat. (Stephani) 3: 665, 1909.

*** Isotachis humectata (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 3: 654, 1909. Bas.: Jungermannia humectata Hook.f. et Taylor, London J. Bot. 3: 462, 1844.

** Isotachis inflata Steph., Arch. Mus. Nac. Rio de Janeiro 13: 113, 1905.

*** Isotachis intortifolia (Hook.f. et Taylor) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864. Bas.: Jungermannia intortifolia Hook.f. et Taylor, London J. Bot. 3: 374, 1844.

** Isotachis japonica Steph., Sp. Hepat. (Stephani) 3: 652, 1909.

* Isotachis lacustris Herzog, Hedwigia 74 (2): 94, 1934.

*** Isotachis lopezii (R.M.Schust.) Gradst., Mem. New York Bot. Gard. 84: 66, 1999. Bas.: Ruizanthus lopezii R.M.Schust., Phytologia 39 (4): 241, 1978.

*** Isotachis lyallii Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 149, 1854.

*** Isotachis minima Pearson, Univ. Calif. Publ. Bot. 10 (4): 322, 1923.

*** Isotachis montana Colenso, Trans. & Proc. New Zealand Inst. 21: 68, 1889.

*** Isotachis multiceps (Lindenb. et Gottsche) Gottsche, Mexik. Leverm.: 105, 1863. Bas.: Jungermannia multiceps Lindenb. et Gottsche, Syn. Hepat. 5: 687, 1847. 67

** Isotachis multiceps var. fendleri Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 125, 1864.

** Isotachis nigella Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 107, 1952.

*** Isotachis obtusa Steph., Sp. Hepat. (Stephani) 6: 354, 1922.

** Isotachis olivacea R.M.Schust., J. Hattori Bot. Lab. 83: 207, 1997.

*** Isotachis plicata J.J.Engel, J. Hattori Bot. Lab. 83: 210, 1997.

** Isotachis pusilla Steph., Sp. Hepat. (Stephani) 3: 655, 1909.

** Isotachis riparia Rodway, Tasm. Bryoph.: 63, 1917.

*** Isotachis serrulata (Sw.) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864. Bas.: Jungermannia serrulata Sw., Prodr. (Swartz): 143, 1788.

*** Isotachis spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 220, 1885.

* Isotachis sprucei Beauverd, Sp. Hepat. (Stephani) 6: 572, 1924. Nom. nov. pro Isotachis trifida Steph., Sp. Hepat. (Stephani) 6: 356, 1922, nom. illeg.

*** Isotachis stephanii E.S.Salmon, Rev. Bryol. 28 (4): 75, 1901.

* Isotachis vexans Steph., Sp. Hepat. (Stephani) 3: 662, 1909.

*** Isotachis westlandica (E.A.Hodgs.) R.M.Schust., Nova Hedwigia 15: 455, 1968. Bas.: Rhizocaulia westlandica E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 78, 1965.

*** Neesioscyphus Grolle, Österr. Bot. Z. 111 (1): 19, 1964.

*** Neesioscyphus allionii (Steph.) Grolle, Rev. Bryol. Lichénol. 34 (1/2): 185, 1966. Bas.: Isotachis allionii Steph., Sp. Hepat. (Stephani) 6: 350, 1922.

*** Neesioscyphus argillaceus (Nees) Grolle, Österr. Bot. Z. 111 (1): 24, 1964. Bas.: Jungermannia argillacea Nees, Fl. Bras. (Martius) 1 (1): 338, 1833.

*** Neesioscyphus bicuspidatus (Steph.) Grolle, Rev. Bryol. Lichénol. 34 (1/2): 182, 1966. Bas.: Isotachis bicuspidata Steph., Symb. Antill. 2: 471, 1901.

*** Neesioscyphus carneus (Nees) Grolle, Österr. Bot. Z. 111 (1): 20, 1964. Bas.: Jungermannia carnea Nees, Fl. Bras. (Martius) 1 (1): 337, 1833.

*** Neesioscyphus homophyllus (Nees) Grolle, Österr. Bot. Z. 111 (2/3): 188, 1964. Bas.: Jungermannia homophylla Nees, Fl. Bras. (Martius) 1 (1): 336, 1833.

** Ruizanthoideae R.M.Schust. ex J.J.Engel et G.L.Merr.

*** Ruizanthus R.M.Schust., Phytologia 39 (4): 240, 1978.

*** Ruizanthus venezuelanus R.M.Schust., Phytologia 39 (4): 240, 1978.

*** Blepharidophyllaceae R.M.Schust. ex J.J.Engel

by J.J. Engel

*** Blepharidophyllum Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 151, 1873.

*** Blepharidophyllum densifolium (Hook.) Ångstr. ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 208, 1885. Bas.: Jungermannia densifolia Hook., Musci Exot. 1: tab. 36, 1818.

*** Blepharidophyllum vertebrale (Gottsche) Ångstr. ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 208, 1885. Bas.: Scapania vertebralis Gottsche, Syn. Hepat. 1: 72, 1844.

*** Clandarium (Grolle) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984]. Bas.: Blepharidophyllum subg. Clandarium Grolle, J. Hattori Bot. Lab. 28: 65, 1965.

*** Clandarium clandestinum (Mont.) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984]. Bas.: Plagiochila clandestina Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 247, 1843.

*** Clandarium gottscheanum (Grolle) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984]. Bas.: Blepharidophyllum gottscheanum Grolle, J. Hattori Bot. Lab. 28: 69, 1965.

*** Clandarium xiphophyllum (Grolle) R.M.Schust., Phytologia 56 (2): 68, 1984. Bas.: Blepharidophyllum xiphophyllum Grolle, J. Hattori Bot. Lab. 28: 65, 1965.

*** Calypogeiaceae Arnell

by M.A.M. Renner

Calypogeiaceae is shown to be monophyletic and the classification follows and.

*** Calypogeia Raddi, Jungermanniogr. Etrusca: 31, 1818 nom. conserv.

** subg. Asperifoliae (Warnst.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 115, 1969. Bas.: Calypogeia [unranked] Asperifoliae Warnst., Bryol. Z. 1 (7): 111, 1917.

*** Calypogeia arguta Nees et Mont., Naturgesch. Eur. Leberm. 3: 24, 1838.

** Calypogeia sullivantii Austin, Hepat. bor.-amer.: 19, 1873.

** subg. Calypogeia

*** Calypogeia andicola Bischl., Candollea 18: 79, 1962.

** Calypogeia annabonensis Steph., Sp. Hepat. (Stephani) 6: 447, 1924.

** Calypogeia azorica Bischl., Rev. Bryol. Lichénol. 37 (1): 116, 1970.

*** Calypogeia azurea Stotler et Crotz, Taxon 32 (1): 74, 1983.

*** Calypogeia bidentula (F.Weber) Nees, Syn. Hepat. 2: 199, 1845. Bas.: Jungermannia bidentula F.Weber, Hist. Musc. Hepat. Prodr.: 38, 1815.

** Calypogeia falcata Bischl., Candollea 18: 112, 1962.

*** Calypogeia fissa (L.) Raddi, Jungermanniogr. Etrusca: 33, 1818. Bas.: Mnium fissum L., Sp. Pl. 1: 1114, 1753, nom. conserv. 68

** Calypogeia fissa subsp. neogaea R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 169, 1969.

** Calypogeia goebelii (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 677 (409), 1908. Bas.: Kantius goebelii Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 260, 1893.

** Calypogeia goebelii var. siamensis N.Kitag., Beih. Nova Hedwigia 90: 165, 1988.

*** Calypogeia grandistipula (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 669 (401), 1908. Bas.: Kantius grandistipulus Steph., Hedwigia 34 (2): 52, 1895.

*** Calypogeia integristipula Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 662 (394), 1908.

*** Calypogeia laxa Gottsche et Lindenb., Syn. Hepat. 5: 713, 1847.

*** Calypogeia lechleri (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 680 (412), 1908. Bas.: Kantius lechleri Steph., Hedwigia 34 (2): 53, 1895. 69

** Calypogeia lechleri var. densifolia (Steph.) Bischl., Candollea 18: 101, 1962. Bas.: Kantius densifolius Steph., Hedwigia 34 (2): 52, 1895.

** Calypogeia longifolia Steph., Sp. Hepat. (Stephani) 6: 449, 1924.

** Calypogeia lophocoleoides Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 677 (409), 1908.

** Calypogeia mascarenensis Bischl., Rev. Bryol. Lichénol. 37 (1): 89, 1970.

** Calypogeia microstipula (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 670 (402), 1908. Bas.: Kantius microstipulus Steph., Hedwigia 34 (2): 53, 1895.

*** Calypogeia miquelii Mont. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 200, 1845.

*** Calypogeia muelleriana (Schiffn.) Müll.Frib., Beih. Bot. Centralbl. 10 (4/5): 217, 1901. Bas.: Kantius muellerianus Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 342, 1900. 70

** Calypogeia muelleriana subsp. blomquistii R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 187, 1969.

*** Calypogeia neesiana (C.Massal. et Carestia) Müll.Frib., Verh. Bot. Vereins Prov. Brandenburg 47: 320, 1905. Bas.: Kantius trichomanis var. neesianus C.Massal. et Carestia, Nuovo Giorn. Bot. Ital. 12 (4): 351, 1880.

** Calypogeia neesiana subsp. subalpina (Inoue) Inoue, Mem. Natl. Sci. Mus. (Tokyo) 4: 58, 1971. Bas.: Calypogeia subalpina Inoue, J. Jap. Bot. 37 (4): 103, 1962.

*** Calypogeia oblata Herzog, Svensk Bot. Tidskr. 51 (1): 189, 1957.

*** Calypogeia peruviana Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 47, 1838.

*** Calypogeia rhombifolia (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 667 (399), 1908. Bas.: Kantius rhombifolius Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 413, 1885.

** Calypogeia rhombifolia var. colombiana Bischl., Candollea 18: 104, 1962.

*** Calypogeia sphagnicola (Arnell et J.Perss.) Warnst. et Loeske, Verh. Bot. Vereins Prov. Brandenburg 47: 320, 1905. Bas.: Kantius sphagnicola Arnell et J.Perss., Rev. Bryol. 29 (2): 26, 1902. 71

*** Calypogeia subintegra (Gottsche, Lindenb. et Nees) Bischl., Candollea 18: 75, 1962. Bas.: Calypogeia peruviana β subintegra Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 712, 1847.

** Calypogeia subintegra var. dussiana (Steph.) Bischl., Candollea 18: 77, 1962. Bas.: Calypogeia dussiana Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 672 (404), 1908.

*** Calypogeia suecica (Arnell et J.Perss.) Müll.Frib., Beih. Bot. Centralbl. 17 (2): 224, 1904. Bas.: Kantius suecicus Arnell et J.Perss., Rev. Bryol. 29 (2): 29, 1902.

*** Calypogeia tenax (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 664 (396), 1908. Bas.: Kantius tenax Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 416, 1885.

** Calypogeia uncinulatula Herzog, Hedwigia 67 (6): 250, 1927.

Incertae sedis

*** Calypogeia aeruginosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861].

* Calypogeia amazonica (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 680 (412), 1908. Bas.: Kantius amazonicus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 415, 1885. 72

** Calypogeia angusta Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 663 (395), 1908.

** Calypogeia apiculata (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 668 (400), 1908. Bas.: Kantius apiculatus Steph., Hedwigia 34 (2): 51, 1895.

** Calypogeia asakawana S.Hatt. ex Inoue, J. Jap. Bot. 39 (4): 107, 1964.

** Calypogeia ceylanica S.Hatt. et Mizut., Candollea 23: 288, 1968.

** Calypogeia contracta Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 139, 1975.

** Calypogeia cuspidata (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 669 (401), 1908. Bas.: Kantius cuspidatus Steph., Bull. Herb. Boissier 5 (10): 846, 1897.

* Calypogeia decurrens (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 675 (407), 1908. Bas.: Kantius decurrens Steph., Hedwigia 34 (2): 52, 1895.

** Calypogeia formosana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 186, 1934.

** Calypogeia fujisana Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 135, 1975.

** Calypogeia granulata Inoue, J. Jap. Bot. 43 (10/11): 468, 1968.

** Calypogeia japonica Steph., Sp. Hepat. (Stephani) 6: 448, 1924.

** Calypogeia khasiana Ajit P.Singh et V.Nath, Taiwania 52 (4): 320, 2007.

** Calypogeia latissima Steph., Sp. Hepat. (Stephani) 6: 449, 1924.

*** Calypogeia lunata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861].

** Calypogeia marginella Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 106, 1860 [1861].

** Calypogeia obovata R.M.Schust., Phytologia 39 (4): 242, 1978.

* Calypogeia steyermarkii Fulford, Mem. New York Bot. Gard. 11 (3): 305, 1968.

** Calypogeia tosana (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 678 (410), 1908. Bas.: Kantius tosanus Steph., Hedwigia 34 (2): 54, 1895.

** Calypogeia udarii Sudipa Das et D.K.Singh, Nelumbo 53: 194, 2011.

** Eocalypogeia (R.M.Schust.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995. Bas.: Metacalypogeia subg. Eocalypogeia R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 107, 1969.

** Eocalypogeia quelpaertensis (S.Hatt. et Inoue) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995. Bas.: Metacalypogeia quelpaertensis S.Hatt. et Inoue, J. Hattori Bot. Lab. 25: 129, 1962.

** Eocalypogeia schusterana (S.Hatt. et Mizut.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995. Bas.: Metacalypogeia schusterana S.Hatt. et Mizut., Misc. Bryol. Lichenol. 4 (8): 121, 1967.

** Metacalypogeia (S.Hatt.) Inoue, J. Hattori Bot. Lab. 21: 231, 1959. Bas.: Calypogeia subg. Metacalypogeia S.Hatt., J. Hattori Bot. Lab. 18: 83, 1957.

*** Metacalypogeia alternifolia (Nees) Grolle, Österr. Bot. Z. 111 (2/3): 185, 1964. Bas.: Mastigobryum alternifolium Nees, Syn. Hepat. 2: 216, 1845.

** Metacalypogeia cordifolia (Steph.) Inoue, J. Hattori Bot. Lab. 21: 233, 1959. Bas.: Calypogeia cordifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 661 (393), 1908.

** Mizutania Furuki et Z.Iwats., J. Hattori Bot. Lab. 67: 291, 1989.

*** Mizutania riccardioides Furuki et Z.Iwats., J. Hattori Bot. Lab. 67: 291, 1989.

*** Mnioloma Herzog, Ann. Bryol. 3: 115, 1930.

** subg. Caracoma (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 833, 1995. Bas.: Calypogeia subg. Caracoma Bischl., Candollea 18: 26, 1962.

** Mnioloma bolivianum (Fulford) R.M.Schust., Beih. Nova Hedwigia 118: 509, 2000. Bas.: Calypogeia boliviana Fulford, Mem. New York Bot. Gard. 11 (3): 291, 1968.

*** Mnioloma caespitosum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995. Bas.: Kantius caespitosus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 412, 1885.

*** Mnioloma cellulosum (Spreng.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 836, 1995. Bas.: Jungermannia cellulosa Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (1): 232, 1827.

*** Mnioloma crenulatum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995. Bas.: Calypogeia crenulata Bischl., Candollea 18: 35, 1962.

*** Mnioloma cyclostipum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 843, 1995. Bas.: Kantius cyclostipus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 411, 1885.

** Mnioloma elliottii (Steph.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 841, 1995. Bas.: Calypogeia elliottii Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 663 (395), 1908.

** Mnioloma fissistipulum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995. Bas.: Calypogeia fissistipula Bischl., Candollea 18: 47, 1962.

*** Mnioloma fuscum (Lehm.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 848, 1995. Bas.: Jungermannia fusca Lehm., Linnaea 4: 360, 1829.

*** Mnioloma nephrostipum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995. Bas.: Kantius nephrostipus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 412, 1885.

** Mnioloma novaezelandiae J.J.Engel, Cryptog. Bryol. 27 (1): 111, 2006.

*** Mnioloma parallelogramum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995. Bas.: Kantius parallelogramus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 413, 1885.

** Mnioloma retusum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995. Bas.: Calypogeia retusa Bischl., Candollea 18: 33, 1962.

** Mnioloma stamatotonum M.A.M.Renner et E.A.Br., Fieldiana, Bot. (n.ser.) 47: 173, 2008.

*** Mnioloma venezuelanum (Fulford) R.M.Schust., Beih. Nova Hedwigia 118: 509, 2000. Bas.: Calypogeia venezuelana Fulford, Mem. New York Bot. Gard. 11 (3): 287, 1968.

** subg. Mnioloma, Fragm. Florist. Geobot. 40 (2): 833, 1995.

*** Mnioloma rhynchophyllum Herzog, Ann. Bryol. 3: 120, 1930.

*** Endogemmataceae Konstant., Vilnet et A.V.Troitsky

by N.A. Konstantinova

described the monotypic family Endogemmataceae based on molecular evidence after a re-evaluation of Solenostomataceae.

*** Endogemma Konstant., Vilnet et A.V.Troitsky, Folia Cryptog. Estonica 48: 132, 2011.

*** Endogemma caespiticia (Lindenb.) Konstant., Vilnet et A.V.Troitsky, Folia Cryptog. Estonica 48: 132, 2011. Bas.: Jungermannia caespiticia Lindenb., Syn. hepat. eur: 67, 1829.

*** Geocalycaceae H.Klinggr.

Placement of Geocalycaceae in Jungermanniinae follows.

*** Geocalyx Nees, Naturgesch. Eur. Leberm. 1: 97, 1833.

*** Geocalyx caledonicus Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 205 (265), 1908.

*** Geocalyx graveolens (Schrad.) Nees, Naturgesch. Eur. Leberm. 2: 397, 1836. Bas.: Jungermannia graveolens Schrad., Syst. Samml. Crypt. Gew. 2: 6, 1797.

** Geocalyx lancistipulus (Steph.) S.Hatt., J. Jap. Bot. 28 (8): 234, 1953. Bas.: Lophocolea lancistipula Steph., Sp. Hepat. (Stephani) 6: 281, 1922.

** Geocalyx orientalis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxix, 1889 [1890].

*** Gymnomitriaceae H.Klinggr.

by J. Váňa

The treatment of the family follows with some modifications. Nardioideae is included following and. Some re-arrangements in Apomarsupella, Gymnomitrion and Marsupella were done by and. Further notes on nomenclature and taxonomy can be found in. Herzogobryum and Nothogymnomitrion were removed from the family by. The placement of Acrolophozia, Nanomarsupella and Paramomitrion is provisional. Inclusion of Cryptocoleopsis follows.

*** Acrolophozia R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 259, 1966.

*** Acrolophozia fuegiana R.M.Schust., Nova Hedwigia 15: 499, 1968.

*** Acrolophozia pectinata R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 261, 1966.

*** Acrolophozia sulcata Hässel, J. Bryol. 11 (1): 108, 1980.

** Nanomarsupella R.M.Schust. ex A.Hagborg, L.Söderstr. et von Konrat, Phytotaxa 112 (1): 16, 2013. Based on: Marsupella subg. Nanomarsupella R.M.Schust., Phytologia 39 (4): 248, 1978.

*** Nanomarsupella xenophylla (R.M.Schust.) R.M.Schust. ex A.Hagborg, L.Söderstr. et von Konrat, Phytotaxa 112 (1): 16, 2013. Bas.: Marsupella xenophylla R.M.Schust., Phytologia 39 (4): 248, 1978.

** Paramomitrion R.M.Schust., J. Hattori Bot. Lab. 80: 134, 1996.

*** Paramomitrion paradoxum R.M.Schust., J. Hattori Bot. Lab. 80: 135, 1996.

*** Gymnomitrioideae T.Jensen

** Cryptocoleopsis Amakawa, J. Hattori Bot. Lab. 21: 274, 1959.

*** Cryptocoleopsis imbricata Amakawa, J. Hattori Bot. Lab. 21: 274, 1959.

*** Gymnomitrion Corda, Gen. hepat.: 651, 1829 nom. conserv.

*** Gymnomitrion adustum Nees, Naturgesch. Eur. Leberm. 1: 120, 1833.

*** Gymnomitrion africanum (Steph.) Horik., Acta Phytotax. Geobot. 13: 212, 1943. Bas.: Acolea africana Steph., Sp. Hepat. (Stephani) 6: 77, 1917.

*** Gymnomitrion alpinum (Gottsche ex Husn.) Schiffn., Österr. Bot. Z. 53 (7): 280, 1903. Bas.: Sarcocyphos alpinus Gottsche ex Husn., Hepaticol. gall. 1: 13, 1875.

*** Gymnomitrion asperulatum R.M.Schust., Acta Acad. Ped. Agr., Sect. Biol. 24: 114, 2003.

*** Gymnomitrion atrofilum Váňa, J. Hattori Bot. Lab. 41: 411, 1976.

*** Gymnomitrion bolivianum (Steph.) Váňa, Novon 20 (2): 225, 2010. Bas.: Anastrophyllum bolivianum Steph., Biblioth. Bot. 87 (2): 186, 1916.

*** Gymnomitrion brevissimum (Dumort.) Warnst., Hedwigia 53 (3): 196, 1913. Bas.: Acolea brevissima Dumort., Syll. Jungerm. Europ.: 76, 1831.

*** Gymnomitrion commutatum (Limpr.) Schiffn., Magyar Bot. Lapok 13: 304, 1914 [1915]. Bas.: Sarcocyphos commutatus Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 57: 314, 1879 [1880].

*** Gymnomitrion concinnatum (Lightf.) Corda, Gen. hepat.: 651, 1829. Bas.: Jungermannia concinnata Lightf., Fl. Scot. 2: 786, 1777, nom. conserv.

*** Gymnomitrion corallioides Nees, Naturgesch. Eur. Leberm. 1: 118, 1833.

*** Gymnomitrion crenatilobum Grolle, Khumbu Himal 1 (4): 278, 1966.

*** Gymnomitrion crenulatum Gottsche ex Carrington, Trans. Bot. Soc. Edinburgh 7 (3): 444, 1863.

*** Gymnomitrion crystallocaulon (Grolle) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015. Bas.: Marsupella crystallocaulon Grolle, Khumbu Himal 1 (4): 281, 1966.

** Gymnomitrion incompletum (Gottsche) R.M.Schust. ex Váňa, J. Hattori Bot. Lab. 40: 186, 1976. Bas.: Jungermannia incompleta Gottsche, Linnaea 28 (5): 551, 1856 [1857].

*** Gymnomitrion laceratum (Steph.) Horik., Acta Phytotax. Geobot. 13: 212, 1943. Bas.: Acolea lacerata Steph., Sp. Hepat. (Stephani) 6: 78, 1917.

*** Gymnomitrion miniatum Lindenb. et Gottsche, Syn. Hepat. 4: 617, 1846.

*** Gymnomitrion minutulum (Hässel) Váňa, Novon 20 (2): 225, 2010. Bas.: Marsupella minutula Hässel, J. Bryol. 11 (1): 123, 1980.

** Gymnomitrion moralesae Váňa, J. Hattori Bot. Lab. 48: 230, 1980.

*** Gymnomitrion mucronulatum (N.Kitag.) N.Kitag., Acta Phytotax. Geobot. 19 (2/3): 53, 1962. Bas.: Gymnomitrion concinnatum var. mucronulatum N.Kitag., Acta Phytotax. Geobot. 18 (2/3): 38, 1959.

** Gymnomitrion mucrophorum R.M.Schust., Bryologist 98 (2): 243, 1995.

*** Gymnomitrion nigrum (Grolle et Váňa) Váňa, Novon 20 (2): 225, 2010. Bas.: Marsupella nigra Grolle et Váňa, J. Hattori Bot. Lab. 40: 186, 1976.

*** Gymnomitrion noguchianum S.Hatt., J. Jap. Bot. 27 (2): 55, 1952.

*** Gymnomitrion obtusilobum N.Kitag., Bull. Univ. Mus. Univ. Tokyo 8: 229, 1975.

*** Gymnomitrion obtusum Lindb., Morgonbladet (Helsinki) 1877 (30, 6 Feb): 2, 1877.

*** Gymnomitrion pacificum Grolle, Trans. Brit. Bryol. Soc. 5 (1): 92, 1966.

*** Gymnomitrion revolutum (Nees) H.Philib., Rev. Bryol. 17 (3): 34, 1890. Bas.: Sarcocyphos revolutus Nees, Naturgesch. Eur. Leberm. 2: 419, 1836.

** Gymnomitrion revolutum subsp. novoguineanensis (R.M.Schust.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015. Bas.: Apomarsupella revoluta subsp. novoguineanensis R.M.Schust., J. Hattori Bot. Lab. 80: 90, 1996.

*** Gymnomitrion rubidum (Mitt.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015. Bas.: Jungermannia rubida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 90, 1860 [1861].

*** Gymnomitrion setaceum Grolle et Váňa, J. Hattori Bot. Lab. 41: 411, 1976.

*** Gymnomitrion sinense Müll.Frib., Rev. Bryol. Lichénol. 20 (1/2): 176, 1951.

** Gymnomitrion strictum (Berggr.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963. Bas.: Cesius strictus Berggr., New Zealand Hepat.: 2, 1898.

** Gymnomitrion strictum var. inaequale R.M.Schust., J. Hattori Bot. Lab. 80: 118, 1996.

*** Gymnomitrion subintegrum (S.W.Arnell) Váňa, Novon 20 (2): 225, 2010. Bas.: Marsupella subintegra S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 545, 1956.

*** Gymnomitrion truncatoapiculatum Herzog, Hedwigia 74 (2): 81, 1934.

*** Gymnomitrion verrucosum W.E.Nicholson, Symb. Sin. 5: 10, 1930.

*** Marsupella Dumort., Commentat. Bot. (Dumortier): 114, 1822.

*** Marsupella alata S.Hatt. et N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 79, 1960.

*** Marsupella andreaeoides (Lindb.) Müll.Frib., Feddes Repert. Spec. Nov. Regni Veg. 54 (2/3): 214, 1951. Bas.: Cesius andreaeoides Lindb., Meddel. Soc. Fauna Fl. Fenn. 14: 68, 1887.

*** Marsupella apiculata Schiffn., Österr. Bot. Z. 53 (6): 249, 1903.

*** Marsupella aquatica (Lindenb.) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 44 (8): 267, 1896 [1897]. Bas.: Jungermannia emarginata var. aquatica Lindenb., Syn. hepat. eur: 75, 1829.

*** Marsupella arctica (Berggr.) Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 26, 1906. Bas.: Sarcocyphos emarginatus var. arcticus Berggr., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 13 (7): 96, 1875.

*** Marsupella boeckii (Austin) Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 409, 1893. Bas.: Sarcocyphos boeckii Austin, Bull. Torrey Bot. Club 3 (3): 9, 1872.

*** Marsupella bolanderi (Austin) Underw., Zoe 1 (12): 365, 1891. Bas.: Sarcocyphos bolanderi Austin, Bull. Torrey Bot. Club 3 (3): 9, 1872.

*** Marsupella condensata (Ångstr. ex C.Hartm.) Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 420, 1893. Bas.: Gymnomitrion condensatum Ångstr. ex C.Hartm., Handb. Skand. fl. (ed. 10): 128, 1871.

*** Marsupella disticha Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 164 (25), 1901.

*** Marsupella emarginata (Ehrh.) Dumort., Recueil Observ. Jungerm.: 24, 1835. Bas.: Jungermannia emarginata Ehrh., Hannover. Mag. 22 (8): 141, 1784.

** Marsupella emarginata subsp. tubulosa (Steph.) N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 76, 1960. Bas.: Marsupella tubulosa Steph., Bull. Herb. Boissier 5 (2): 99, 1897.

** Marsupella emarginata subsp. tubulosa var. apertifolia (Steph.) N.Kitag., J. Hattori Bot. Lab. 26: 89, 1963. Bas.: Marsupella apertifolia Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 162 (23), 1901.

** Marsupella emarginata subsp. tubulosa var. patens N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 77, 1960.

** Marsupella emarginata subsp. tubulosa var. tubulosa (Steph.) N.Kitag. ex Váňa et L.Söderstr., Phytotaxa 183 (4): 288, 2014. Bas.: Marsupella tubulosa Steph., Bull. Herb. Boissier 5 (2): 99, 1897.

*** Marsupella funckii (F.Weber et D.Mohr) Dumort., Recueil Observ. Jungerm.: 24, 1835. Bas.: Jungermannia funckii F.Weber et D.Mohr, Bot. Taschenb. (Weber): 422, 1807.

*** Marsupella microphylla R.M.Schust., Phytologia 39 (4): 249, 1978.

** Marsupella minutissima N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 81, 1960.

*** Marsupella neesii Sande Lac. ex Schiffn., Consp. Hepat. Arch. Ind.: 70, 1898. Based on: Sarcocyphos neesii Nees ex Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 288, 1864, nom. inval.

*** Marsupella paroica R.M.Schust., Bryologist 60 (2): 145, 1957.

*** Marsupella profunda Lindb., Rev. Bryol. 14 (2): 19, 1887.

*** Marsupella pseudofunckii S.Hatt., J. Hattori Bot. Lab. 4: 63, 1950.

*** Marsupella sparsifolia (Lindb.) Dumort., Bull. Soc. Roy. Bot. Belgique 13: 128, 1874. Bas.: Sarcocyphos sparsifolius Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 280, 1868.

** Marsupella sparsifolia subsp. childii R.M.Schust., Phytotaxa 183 (4): 288, 2014. Based on: Marsupella sparsifolia subsp. childii R.M.Schust., J. Hattori Bot. Lab. 80: 61, 1996, nom. inval.

*** Marsupella sphacelata (Giesecke ex Lindenb.) Dumort., Recueil Observ. Jungerm.: 24, 1835. Bas.: Jungermannia sphacelata Giesecke ex Lindenb., Syn. hepat. eur: 76, 1829.

*** Marsupella spiniloba R.M.Schust. et Damsh., Phytologia 63 (5): 326, 1987.

*** Marsupella sprucei (Limpr.) Bernet, Cat. hép. Suisse: 33, 1888. Bas.: Sarcocyphos sprucei Limpr., Flora 64 (5): 72, 1881.

** Marsupella stableri Spruce, Rev. Bryol. 8 (6): 96, 1881.

*** Marsupella stoloniformis N.Kitag., J. Hattori Bot. Lab. 30: 201, 1967.

** Marsupella stoloniformis subsp. vermiformis R.M.Schust., J. Hattori Bot. Lab. 80: 72, 1996.

*** Marsupella yakushimensis (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 80, 1944. Bas.: Sphenolobus yakushimensis Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 156, 1934.

*** Poeltia Grolle, Khumbu Himal 1 (4): 280, 1966.

*** Poeltia campylata Grolle, Khumbu Himal 1 (4): 280, 1966.

*** Prasanthus Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 62, 1889.

** Prasanthus jamalicus Potemkin, Ann. Bot. Fenn. 29 (4): 319, 1992.

*** Prasanthus suecicus (Gottsche) Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 62, 1889. Bas.: Gymnomitrion suecicum Gottsche, Fl. Danica 16 (48): 20, 1871.

*** Nardioideae Váňa

*** Nardia Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 nom. conserv.

*** Nardia arnelliana Grolle, Bot. Mag. (Tokyo) 77 (914): 297, 1964.

*** Nardia assamica (Mitt.) Amakawa, J. Hattori Bot. Lab. 26: 23, 1963. Bas.: Jungermannia assamica Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861].

*** Nardia breidleri (Limpr.) Lindb., Helsingf. Dagbl. 1880 (311, 15 Nov.): 2, 1880. Bas.: Alicularia breidleri Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 57: 311, 1879 [1880].

*** Nardia compressa (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 694, 1821. Bas.: Jungermannia compressa Hook., Brit. Jungermann.: tab. 58, 1813.

*** Nardia flagelliformis Inoue, J. Jap. Bot. 46 (1): 1, 1971.

*** Nardia geoscyphus (De Not.) Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 371, 1874. Bas.: Alicularia geoscyphus De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 18: 486, 1859.

* Nardia geoscyphus var. dioica Bakalin, Arctoa 18: 87, 2009 [2010].

* Nardia geoscyphus var. suberecta (Lindb. ex Kaal.) Váňa, Phytotaxa 76 (3): 37, 2013. Bas.: Nardia haematosticta var. suberecta Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 395, 1893.

*** Nardia grollei Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 491, 2009.

*** Nardia insecta Lindb., Helsingf. Dagbl. 1878 (315, 18 Nov.): 2, 1878.

*** Nardia japonica Steph., Bull. Herb. Boissier 5 (2): 101, 1897.

* Nardia kamtschatica Arnell et C.E.O.Jensen, Hedwigia 67 (1/2): 111, 1927. 73

* Nardia leptocaulis C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 205, 1981.

*** Nardia lescurii (Austin) Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 115, 1884. Bas.: Alicularia lescurii Austin, Hepat. bor.-amer.: 4, 1873.

** Nardia minutifolia Furuki, Bryol. Res. 9 (3): 73, 2006.

** Nardia nuda (Lindenb. et Gottsche) Váňa, Folia Geobot. Phytotax. 8 (2): 193, 1973. Bas.: Jungermannia nuda Lindenb. et Gottsche, Syn. Hepat. 5: 668, 1847.

*** Nardia poeltii Váňa, J. Hattori Bot. Lab. 36: 73, 1972 [1973].

*** Nardia scalaris Gray, Nat. Arr. Brit. Pl. 1: 694, 1821.

* Nardia scalaris var. botryoidea (R.M.Schust.) Váňa, Phytotaxa 76 (3): 38, 2013. Bas.: Nardia scalaris subsp. botryoidea R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 862, 1969.

* Nardia scalaris var. harae (Amakawa) Váňa, Phytotaxa 76 (3): 38, 2013. Bas.: Nardia harae Amakawa, J. Jap. Bot. 32 (2): 38, 1957.

*** Nardia subclavata (Steph.) Amakawa, J. Jap. Bot. 32 (2): 40, 1957. Bas.: Jungermannia subclavata Steph., Sp. Hepat. (Stephani) 6: 93, 1917.

*** Nardia succulenta (A.Rich.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 519, 1885. Bas.: Jungermannia succulenta A.Rich., Nov. Stirp. Pug. 4: 43, 1832.

*** Nardia unispiralis Amakawa, J. Jap. Bot. 32 (6): 167, 1957.

** Gyrothyraceae R.M.Schust.

Placement of Gyrothyraceae in Jungermanniinae follows.

** Gyrothyra M.Howe, Bull. Torrey Bot. Club 24 (4): 201, 1897.

*** Gyrothyra underwoodiana M.Howe, Bull. Torrey Bot. Club 24 (4): 202, 1897.

*** Harpanthaceae Arnell

and resolved Harpanthus as an independent lineage which is recognized here as a monogeneric family as originally construed by.

*** Harpanthus Nees, Naturgesch. Eur. Leberm. 2: 351, 1836.

*** Harpanthus drummondii (Taylor) Grolle, Österr. Bot. Z. 112 (3): 274, 1965. Bas.: Chiloscyphus drummondii Taylor, London J. Bot. 5: 283, 1846.

*** Harpanthus flotovianus (Nees) Nees, Naturgesch. Eur. Leberm. 2: 353, 1836. Bas.: Jungermannia flotoviana Nees, Flora 16 (26): 408, 1833.

*** Harpanthus scutatus (F.Weber et D.Mohr) Spruce, Trans. Bot. Soc. Edinburgh 3 (1/4): 209, 1850. Bas.: Jungermannia scutata F.Weber et D.Mohr, Bot. Taschenb. (Weber): 408, 1807.

** Hygrobiellaceae Konstant. et Vilnet

by N. Konstantinova

The family Hygrobiellaceae was validated by.

*** Hygrobiella Spruce, Cephalozia: 73, 1882.

** Hygrobiella intermedia Bakalin et Vilnet, Pl. Syst. Evol. 300 (10): 2286, 2014.

*** Hygrobiella laxifolia (Hook.) Spruce, Cephalozia: 74, 1882. Bas.: Jungermannia laxifolia Hook., Brit. Jungermann.: tab. 59, 1813.

** Hygrobiella squamosa Bakalin et Vilnet, Pl. Syst. Evol. 300 (10): 2286, 2014.

*** Jackiellaceae R.M.Schust.

by J. Váňa

The monogeneric status of Jackiella is supported by a molecular study based on three loci by.

*** Jackiella Schiffn., Hep. Fl. Buitenzorg: 211, 1900.

** Jackiella angustifolia Herzog, Trans. Brit. Bryol. Soc. 1 (4): 296, 1950.

** Jackiella ceylanica Schiffn. ex Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 212 (272), 1908.

*** Jackiella curvata E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand 85 (4): 571, 1958.

*** Jackiella javanica Schiffn., Hep. Fl. Buitenzorg: 212, 1900.

** Jackiella javanica var. cavifolia Schiffn., Hep. Fl. Buitenzorg: 213, 1900.

** Jackiella javanica var. cordifolia Schiffn., Hep. Fl. Buitenzorg: 213, 1900.

** Jackiella renifolia Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 218, 1900 [1901].

** Jackiella sinensis (W.E.Nicholson) Grolle, Österr. Bot. Z. 111 (2/3): 186, 1964. Bas.: Aplozia sinensis W.E.Nicholson, Symb. Sin. 5: 12, 1930.

** Jackiella singapurensis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 218, 1900 [1901].

** Jackiella singapurensis var. philippinensis N.Kitag., Misc. Bryol. Lichenol. 9 (1): 9, 1981.

Excluded from the genus

* Jackiella unica Steph., Sp. Hepat. (Stephani) 6: 318, 1922. 74

*** Jungermanniaceae Rchb.

by J. Váňa

The treatment of Jungermanniaceae follows. Many old names in Jungermannia are still neither synonymized nor transferred and we do not know their value. Some of them may prove to be older names of currently accepted taxa. We list those doubtful names in a separate section below.

** Delavayelloideae Grolle

** Delavayella Steph., Hedwigia 33 (1): 4, 1894.

** Delavayella serrata Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 211, 1894.

** Delavayella serrata var. purpurea P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 38, 1955.

** Liochlaena Nees, Syn. Hepat. 2: 150, 1845.

*** Liochlaena lanceolata Nees, Syn. Hepat. 2: 150, 1845.

*** Liochlaena subulata (A.Evans) Schljakov, Pečen. Mchi Sev. SSSR 4: 71, 1981. Bas.: Jungermannia subulata A.Evans, Trans. Connecticut Acad. Arts 8 (15): 258, 1891.

** Jungermannioideae Dumort.

** Eremonotus Lindb. et Kaal. ex Pearson, Hepat. Br. Isl. 1 (6-15): 200, 1900.

*** Eremonotus myriocarpus (Carrington) Lindb. et Kaal. ex Pearson, Hepat. Br. Isl. 1 (6-15): 201, 1900. Bas.: Jungermannia myriocarpa Carrington, Hepat. Brit. Exsicc. Fasc. II: no. 96, 1879.

*** Jungermannia L., Sp. Pl. 1: 1131, 1753. 75

*** Jungermannia atrovirens Dumort., Syll. Jungerm. Europ.: 51, 1831.

*** Jungermannia borealis Damsh. et Váňa, Lindbergia 4 (1/2): 5, 1977.

* Jungermannia erectii Ajit P.Singh et V.Nath, Hepat. Khasi Jaintia Hills: E. Himal.: 117, 2007. 76

*** Jungermannia exsertifolia Steph., Sp. Hepat. (Stephani) 6: 86, 1917.

** Jungermannia exsertifolia subsp. cordifolia (Dumort.) Váňa, Folia Geobot. Phytotax. 8 (3): 268, 1973. Bas.: Aplozia cordifolia Dumort., Bull. Soc. Roy. Bot. Belgique 13: 59, 1874.

*** Jungermannia gollanii Steph., Sp. Hepat. (Stephani) 6: 86, 1917.

** Jungermannia konstantinovae Bakalin et Vilnet, Arctoa 18: 161, 2009 [2010].

*** Jungermannia ovatotrigona (Steph.) Grolle, Feddes Repert. 82 (1): 90, 1971. Bas.: Jamesoniella ovatotrigona Steph., Biblioth. Bot. 87 (2): 184, 1916.

*** Jungermannia polaris Lindb., Öfvers. Kongl. Vetensk.-Akad. Förh. 23 (10): 560, 1866 [1867].

*** Jungermannia pumila With., Arr. Brit. Pl., ed. 3, 3: 883, 1796.

Taxa doubtfully belonging to the genus 77

* Jungermannia amentacea Bertol., Mem. Reale Accad. Sci. Ist. Bologna (ser. 2) 1: 19, 1862.

* Jungermannia brasiliensis Raddi, Critt. Brasil.: 15, 1822. 78

* Jungermannia chinensis Osbeck, Dagb. Ostind. Resa: 221, 1757. 79

* Jungermannia cordata Vill., Hist. Pl. Dauphiné (Villars) 3: 923, 1789.

* Jungermannia crenulata Schmidel, Jungerm. Char.: 20, 1760.

* Jungermannia creutzeri Kremer, Monogr. hépat. Moselle: 26, 1837.

* Jungermannia digitata C.F.W.Meissn. ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 326, 1827.

* Jungermannia dubioides H.A.Mill., Phytologia 47 (4): 322, 1981. Nom. nov. pro Jungermannia dubia Nees, Prodr. Fl. Norfolk.: 5, 1833, nom. illeg.

* Jungermannia fernandeziana Mitt., Rep. Challenger, Bot. 1 (3, 1): 85, 1884.

* Jungermannia hexagona Schwägr., Hist. Musc. Hepat. Prodr.: 18, 1814.

* Jungermannia holandriana Kremer, Monogr. hépat. Moselle: 25, 1837.

* Jungermannia incerta Gottsche, Abh. Naturwiss. Vereins Bremen 7: 344, 1882. 80

* Jungermannia lateriflora Hampe ex Gottsche, Mexik. Leverm.: 82, 1863.

* Jungermannia lescuriana Austin, Rep. (Annual) Regents Univ. State New York State Cab. Nat. Hist. 19: 67, 1866. 81

* Jungermannia longiretis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxv, 1889 [1890]. 82

* Jungermannia mastigophora Spreng. Neue Entdeck. Pflanzenk. 2: 99, 1821. 83

* Jungermannia michelii Mérat, Nouv. fl. env. Paris (ed. 2) 1: 219, 1821. 84

* Jungermannia minima Scop., Fl. Carniol. (ed. 2) 2: 350, 1772.

* Jungermannia odorata With., Bot. arr. veg. Gr. Brit. 2: 693, 1776.

* Jungermannia peltata Schmidel, Jungerm. Char.: 14, 1760.

* Jungermannia quadridigitata Griff., Not. pl. asiat. 2: 314, 1849. 85

* Jungermannia sauteri De Not. ex Rabenh., Hedwigia 1 (20): 121, 1857.

* Jungermannia secunda Hampe ex Gottsche, Mexik. Leverm.: 82, 1863.

* Jungermannia stereocaulis Bory, Voy. Uranie, Bot. 4: 130, 1827. 86

* Jungermannia submersa Kremer, Monogr. hépat. Moselle: 36, 1837.

* Jungermannia sullivantiana Austin, Rep. (Annual) Regents Univ. State New York State Cab. Nat. Hist. 19: 66, 1866. 87

* Jungermannia supina Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 86, 1795 [1796].

* Jungermannia tenuis Ehrh., Beitr. Naturk. (Ehrhart) 4: 45, 1789. 88

* Jungermannia uncifolia Steph., Hedwigia 34 (2): 51, 1895.

* Jungermannia vernicosa Cass. ex Mérat, Nouv. fl. env. Paris (ed. 2) 1: 221, 1821.

** Mesoptychioideae R.M.Schust.

*** Mesoptychia (Lindb.) A.Evans, Ottawa Naturalist 17: 15, 1903. Bas.: Jungermannia sect. Mesoptychia Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 39, 1889.

*** Mesoptychia badensis (Gottsche ex Rabenh.) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012. Bas.: Jungermannia badensis Gottsche ex Rabenh., Hepat. Eur., Leberm. 9-10: no. 95, 1859.

*** Mesoptychia bantriensis (Hook.) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012. Bas.: Jungermannia bantriensis Hook., Brit. Jungermann.: tab. 41, 1813.

** Mesoptychia bantriensis subsp. wallfischii (Ştefănuţ) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012. Bas.: Leiocolea bantriensis subsp. wallfischii Ştefănuţ, Hornwort Liverwort Romania: 21, 2008.

* Mesoptychia chichibuensis (Inoue) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012. Bas.: Lophozia chichibuensis Inoue, J. Jap. Bot. 36 (2): 41, 1961.

* Mesoptychia collaris (Nees) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012. Bas.: Jungermannia collaris Nees, Fl. crypt. erlang.: xv, 1817.

* Mesoptychia fitzgeraldiae (Paton et A.R.Perry) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012. Bas.: Leiocolea fitzgeraldiae Paton et A.R.Perry, J. Bryol. 18 (3): 470, 1995.

*** Mesoptychia gillmanii (Austin) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012. Bas.: Jungermannia gillmanii Austin, Bull. Torrey Bot. Club 3 (3): 12, 1872.

*** Mesoptychia heterocolpos (Thed. ex Hartm.) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012. Bas.: Jungermannia heterocolpos Thed. ex Hartm., Handb. Skand. fl. (ed. 3): 328, 1838.

** Mesoptychia heterocolpos var. arctica (S.W.Arnell) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012. Bas.: Leiocolea arctica S.W.Arnell, Svensk Bot. Tidskr. 44 (2): 374, 1950.

** Mesoptychia heterocolpos var. harpanthoides (Bryhn et Kaal.) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012. Bas.: Lophozia harpanthoides Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 31, 1906.

*** Mesoptychia igiana (S.Hatt.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012. Bas.: Lophozia igiana S.Hatt., J. Jap. Bot. 31 (7): 201, 1956.

* Mesoptychia mamatkulovii (Duda) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012. Bas.: Lophozia mamatkulovii Duda, Trans. Brit. Bryol. Soc. 6 (1): 82, 1970.

*** Mesoptychia mayebarae (S.Hatt.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012. Bas.: Cephalozia mayebarae S.Hatt., J. Hattori Bot. Lab. 3: 37, 1948 [1950].

*** Mesoptychia morrisoncola (Horik.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012. Bas.: Lophozia morrisoncola Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 150, 1934.

* Mesoptychia polymorpha Stotler, Crand.-Stotl. et Bakalin, Polish Bot. J. 58 (1): 82, 2013.

*** Mesoptychia rutheana (Limpr.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012. Bas.: Jungermannia rutheana Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 61: 207, 1884.

** Mesoptychia rutheana var. laxa (Schiffn. ex Burrell) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012. Bas.: Lophozia schultzii var. laxa Schiffn. ex Burrell, J. Bot. 49: 217, 1911.

*** Mesoptychia sahlbergii (Lindb. et Arnell) A.Evans, Ottawa Naturalist 17: 15, 1903. Bas.: Jungermannia sahlbergii Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 40, 1889.

*** Mesoptychia subcrispa (Herzog) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012. Bas.: Lophozia subcrispa Herzog, Ann. Naturhist. Mus. Wien 53 (1): 362, 1942 [1943].

*** Mesoptychia turbinata (Raddi) L.Söderstr. et Váňa, Phytotaxa 65: 55, 2012. Bas.: Jungermannia turbinata Raddi, Jungermanniogr. Etrusca: 18, 1818.

* Mesoptychia ussuriensis (Bakalin) L.Söderstr. et Váňa, Phytotaxa 65: 55, 2012. Bas.: Leiocolea ussuriensis Bakalin, Arctoa 17: 103, 2008 [2009].

** Rivulariella D.H.Wagner, Phytoneuron 2013 (10): 2, 2013.

** Rivulariella gemmipara (A.Evans) D.H.Wagner, Phytoneuron 2013 (10): 2, 2013. Bas.: Chiloscyphus gemmiparus A.Evans, Bryologist 41 (3): 50, 1938.

** Notoscyphaceae Crand.-Stotl., Váňa et Stotler

circumscribed Notoscyphaceae as monogeneric based on molecular and morphological evidence.

*** Notoscyphus Mitt., Fl. vit.: 407, 1871 [1873].

*** Notoscyphus lutescens (Lehm. et Lindenb.) Mitt., Fl. vit.: 407, 1871 [1873]. Bas.: Jungermannia lutescens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 16, 1832.

** Saccogynaceae Heeg

recognized Saccogynaceae as being monogeneric.

*** Saccogyna Dumort., Commentat. Bot. (Dumortier): 113, 1822 nom. conserv.

* Saccogyna ligulata Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 207 (267), 1908.

** Saccogyna subacuta Steph., Sp. Hepat. (Stephani) 6: 316, 1922.

*** Saccogyna viticulosa (L.) Dumort., Syll. Jungerm. Europ.: 74, 1831. Bas.: Jungermannia viticulosa L., Sp. Pl. 1: 1131, 1753.

Excluded from the genus

* Saccogyna tridens Steph., Sp. Hepat. (Stephani) 6: 317, 1922. 89

*** Solenostomataceae Stotler et Crand.-Stotl.

by J. Váňa

Solenostomataceae was first defined by and refined to the current circumscription in. The placement of Aponardia is provisional following and the placement of Arctoscyphus, Cryptocolea and Diplocolea in the family is also provisional. Taxonomic and nomenclatural notes can also be found in and, ).

** Aponardia (R.M.Schust.) Váňa, Phytotaxa 65: 46, 2012. Bas.: Nardia subg. Aponardia R.M.Schust., Beih. Nova Hedwigia 119: 360, 2002.

*** Aponardia huerlimannii (Váňa et Grolle) Váňa, Phytotaxa 65: 46, 2012. Bas.: Nardia huerlimannii Váňa et Grolle, Österr. Bot. Z. 118 (3): 233, 1970.

** Arctoscyphus Hässel, Lindbergia 16 (4): 133, 1990 [1992].

** Arctoscyphus fuegiensis (C.Massal.) Hässel, Cryptog. Bryol. Lichénol. 17 (3): 164, 1996. Bas.: Leioscyphus repens var. fuegiensis C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 212, 1885.

** Arctoscyphus ronsmithii Hässel, Lindbergia 16 (4): 133, 1990 [1992].

** Cryptocolea R.M.Schust., Amer. Midl. Naturalist 49 (2): 414, 1953.

*** Cryptocolea imbricata R.M.Schust., Amer. Midl. Naturalist 49 (2): 417, 1953.

** Diplocolea Amakawa, J. Jap. Bot. 37 (9): 274, 1962.

*** Diplocolea sikkimensis Amakawa, J. Jap. Bot. 37 (9): 274, 1962.

*** Solenostoma Mitt., J. Proc. Linn. Soc., Bot. 8 (29): 51, 1864 [1865] nom. conserv. 90

** subg. Eucalyx (Lindb.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 38, 2015. Bas.: Nardia sect. Eucalyx Lindb., Acta Soc. Sci. Fenn. 10: 525, 1875.

*** Solenostoma bilobum (S.Hatt. ex Amakawa) Potemkin et Nyushko, Liverworts and Hornworts of Russia 1: 286, 2009. Bas.: Plectocolea biloba S.Hatt. ex Amakawa, J. Jap. Bot. 32 (7): 216, 1957.

*** Solenostoma emarginatum (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010. Bas.: Plectocolea emarginata Amakawa, J. Jap. Bot. 33 (11): 340, 1958.

*** Solenostoma flagellatum (S.Hatt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009. Bas.: Plectocolea flagellata S.Hatt., J. Hattori Bot. Lab. 3: 13, 1948 [1950].

*** Solenostoma hokkaidense (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010. Bas.: Jungermannia hokkaidensis Váňa, J. Hattori Bot. Lab. 35: 314, 1972.

*** Solenostoma obovatum (Nees) C.Massal., Epat. erb. critt. ital.: 17, 1903. Bas.: Jungermannia obovata Nees, Naturgesch. Eur. Leberm. 1: 332, 1833.

*** Solenostoma obscurum (A.Evans) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1013, 1969. Bas.: Nardia obscura A.Evans, Rhodora 21 (249): 159, 1919.

*** Solenostoma schusteranum (J.D.Godfrey et G.Godfrey) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010. Bas.: Jungermannia schusterana J.D.Godfrey et G.Godfrey, J. Hattori Bot. Lab. 46: 109, 1979.

*** Solenostoma subtilissimum (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1027, 1969. Bas.: Nardia subtilissima Schiffn., Ann. K. K. Naturhist. Hofmus. 23: 136, 1909.

** subg. Metasolenostoma Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 38, 2015.

*** Solenostoma fusiforme (Steph.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 944, 1969. Bas.: Nardia fusiformis Steph., Bull. Herb. Boissier 5 (2): 99, 1897.

*** Solenostoma gracillimum (Sm.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 972, 1969. Bas.: Jungermannia gracillima Sm., Engl. Bot. 32: tab. 2238, 1811.

*** Solenostoma handelii (Schiffn.) Müll.Frib., Beitr. Kryptogamenfl. Schweiz 10 (2): 38, 1947. Bas.: Nardia handelii Schiffn., Ann. K. K. Naturhist. Hofmus. 23: 135, 1909.

** Solenostoma lignicola (Schiffn.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010. Bas.: Nardia lignicola Schiffn., Ann. K. K. Naturhist. Hofmus. 23: 137, 1909.

*** Solenostoma limbatifolium (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009. Bas.: Jungermannia limbatifolia Amakawa, J. Hattori Bot. Lab. 31: 112, 1968.

** Solenostoma philippinense Váňa, Phytotaxa 152 (1): 43, 2013. Based on: Solenostoma gracillimum subsp. camiguinense Bakalin, Polish Bot. J. 58 (1): 134, 2013.

*** Solenostoma rubrum (Gottsche) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 975, 1969. Bas.: Jungermannia rubra Gottsche, Bot. Gaz. 13 (5): 113, 1888.

*** Solenostoma suborbiculatum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009. Bas.: Jungermannia suborbiculata Amakawa, J. Hattori Bot. Lab. 31: 112, 1968.

** subg. Plectocolea Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864.

*** Solenostoma balfourii (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010. Bas.: Jungermannia balfourii Váňa, Folia Geobot. Phytotax. 9 (3): 279, 1974.

*** Solenostoma borneense (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010. Bas.: Jungermannia borneensis Amakawa, J. Hattori Bot. Lab. 33: 160, 1970.

*** Solenostoma callithrix (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 486 (48), 1901. Bas.: Jungermannia callithrix Lindenb. et Gottsche, Syn. Hepat. 5: 673, 1847.

*** Solenostoma champawatense (S.N.Srivast. et Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009. Bas.: Jungermannia champawatensis S.N.Srivast. et Amakawa, Proc. Natl. Acad. Sci. India, B 61 (2): 205, 1991.

*** Solenostoma comatum (Nees) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 73, 1981. Bas.: Jungermannia comata Nees, Enum. Pl. Crypt. Javae: 78, 1830.

*** Solenostoma comatum var. novae-guineae (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010. Bas.: Jungermannia comata var. novae-guineae Váňa, J. Hattori Bot. Lab. 37: 187, 1973.

*** Solenostoma crenuliforme (Austin) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 494 (56), 1901. Bas.: Jungermannia crenuliformis Austin, Bull. Torrey Bot. Club 3 (3): 10, 1872.

*** Solenostoma decolor (Schiffn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010. Bas.: Jungermannia decolor Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 10, 1890.

*** Solenostoma dusenii (Steph.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010. Bas.: Nardia dusenii Steph., Hedwigia 30 (5): 209, 1891.

*** Solenostoma erectum (Amakawa) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 66, 1981. Bas.: Plectocolea erecta Amakawa, J. Jap. Bot. 32 (10): 307, 1957.

* Solenostoma flagellalioides C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 205, 1981.

*** Solenostoma flavialbicans (Amakawa et Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009. Bas.: Jungermannia flavialbicans Amakawa et Grolle, J. Hattori Bot. Lab. 31: 108, 1968.

*** Solenostoma fossombronioides (Austin) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1027, 1969. Bas.: Jungermannia fossombronioides Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 220, 1869.

*** Solenostoma glaucum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009. Bas.: Jungermannia glauca Amakawa, Fl. E. Himalaya: 511, 1966.

* Solenostoma gongshanense (C.Gao et J.Sun) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009. Bas.: Jungermannia gongshanensis C.Gao et J.Sun, Bull. Bot. Res., Harbin 27 (2): 140, 2007.

*** Solenostoma haskarlianum (Nees) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009. Bas.: Alicularia haskarliana Nees, Syn. Hepat. 1: 12, 1844.

*** Solenostoma hattorianum (Amakawa) Potemkin et Nyushko, Liverworts and Hornworts of Russia 1: 287, 2009. Bas.: Plectocolea hattoriana Amakawa, J. Jap. Bot. 33 (11): 341, 1958.

*** Solenostoma hirticalyx (Steph.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010. Bas.: Jungermannia hirticalyx Steph., Sp. Hepat. (Stephani) 6: 87, 1917.

*** Solenostoma horikawanum (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010. Bas.: Plectocolea horikawana Amakawa, J. Jap. Bot. 32 (7): 219, 1957.

*** Solenostoma hyalinum (Lyell) Mitt., Nat. hist. Azores: 319, 1870. Bas.: Jungermannia hyalina Lyell, Brit. Jungermann.: tab. 63, 1814.

*** Solenostoma infuscum (Mitt.) Hentschel, Pl. Syst. Evol. 268 (1/4): 152, 2007. Bas.: Plectocolea infusca Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 196, 1891.

*** Solenostoma infuscum var. ovicalyx (Steph.) Potemkin et Sofronova, Liverworts and Hornworts of Russia 1: 288, 2009. Bas.: Solenostoma ovicalyx Steph., Sp. Hepat. (Stephani) 6: 82, 1917.

** Solenostoma kurilense (Bakalin) Váňa, Phytotaxa 152 (1): 40, 2013. Bas.: Plectocolea flagellata var. kurilensis Bakalin, Arctoa 18: 90, 2009 [2010].

* Solenostoma lixingjiangii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009. Bas.: Jungermannia lixingjiangii C.Gao et X.L.Bai, Philipp. Scientist 38: 128, 2001.

*** Solenostoma marginatum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 983, 1969. Bas.: Plectocolea marginata S.Hatt., J. Hattori Bot. Lab. 3: 40, 1948 [1950].

*** Solenostoma micranthum (Mitt.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010. Bas.: Plectocolea micrantha Mitt., Fl. vit.: 405, 1871 [1873].

** Solenostoma montanum (Steph.) Váňa, Phytotaxa 65: 44, 2012. Bas.: Nardia montana Steph., Hedwigia 28 (3): 164, 1889.

* Solenostoma multicarpum (C.Gao et J.Sun) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009. Bas.: Jungermannia multicarpa C.Gao et J.Sun, Bull. Bot. Res., Harbin 27 (2): 139, 2007.

** Solenostoma nilgiriense (A.Alam, Ad.Kumar et S.C.Srivast.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009. Bas.: Jungermannia nilgiriensis A.Alam, Ad.Kumar et S.C.Srivast., Bull. Bot. Surv. India 49 (1/4): 220, 2007.

*** Solenostoma obliquifolium (Schiffn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010. Bas.: Nardia obliquifolia Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 191, 1898.

*** Solenostoma onraedtii (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010. Bas.: Jungermannia onraedtii Váňa, Folia Geobot. Phytotax. 9 (3): 282, 1974.

* Solenostoma orbicularifolium (Piippo ex C.Gao et Bai) Váňa, Phytotaxa 222 (2): 199, 2015. Bas.: Jungermannia orbicularifolia Piippo ex C.Gao et Bai, Philipp. Scientist 38: 129, 2001.

*** Solenostoma otianum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 984, 1969. Bas.: Plectocolea otiana S.Hatt., J. Jap. Bot. 28 (6): 183, 1953.

*** Solenostoma ovalifolium (Amakawa) Váňa, Phytotaxa 152 (1): 40, 2013. Bas.: Plectocolea infusca var. ovalifolia Amakawa, J. Jap. Bot. 34 (4): 115, 1959.

*** Solenostoma paroicum (Schiffn.) R.M.Schust., Amer. Midl. Naturalist 49 (2): 402, 1953. Bas.: Nardia paroica Schiffn., Lotos 58: 320, 1910.

*** Solenostoma plagiochilaceum (Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009. Bas.: Jungermannia plagiochilacea Grolle, J. Hattori Bot. Lab. 58: 197, 1985.

*** Solenostoma polyrhizoides (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009. Bas.: Jungermannia polyrhizoides Grolle ex Amakawa, J. Hattori Bot. Lab. 29: 262, 1966.

*** Solenostoma radicellosum Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864 [1865].

*** Solenostoma renauldii (Steph.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010. Bas.: Jungermannia renauldii Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 201, 1891 [1892].

*** Solenostoma rigidulum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 983, 1969. Bas.: Plectocolea rigidula S.Hatt., J. Jap. Bot. 27 (2): 53, 1952.

*** Solenostoma rosulans (Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009. Bas.: Nardia rosulans Steph., Bull. Herb. Boissier 5 (2): 101, 1897.

*** Solenostoma rotundatum Amakawa, J. Jap. Bot. 31 (2): 50, 1956.

*** Solenostoma rubripunctatum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 898, 1969. Bas.: Plectocolea rubripunctata S.Hatt., J. Hattori Bot. Lab. 3: 41, 1948 [1950].

* Solenostoma rupicola (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009. Bas.: Jungermannia rupicola Amakawa, J. Hattori Bot. Lab. 22: 23, 1960. 91

*** Solenostoma sikkimense (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009. Bas.: Jungermannia sikkimensis Schiffn. ex Steph., Sp. Hepat. (Stephani) 6: 92, 1917.

*** Solenostoma tetragonum (Lindenb.) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 509, 2009. Bas.: Jungermannia tetragona Lindenb., Bot. Zeitung (Berlin) 6 (25): 462, 1848.

* Solenostoma tetragonum var. kodaikanalense A.Alam, D.Sharma et So.Yadav, Phytotaxonomy 12: 70, 2012.

*** Solenostoma torticalyx (Steph.) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 69, 1981. Bas.: Jungermannia torticalyx Steph., Sp. Hepat. (Stephani) 6: 94, 1917.

*** Solenostoma truncatum (Nees) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 509, 2009. Bas.: Jungermannia truncata Nees, Enum. Pl. Crypt. Javae: 29, 1830.

** Solenostoma truncatum var. setulosum (Herzog) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009. Bas.: Plectocolea setulosa Herzog, J. Hattori Bot. Lab. 14: 33, 1955.

*** Solenostoma tuberculiferum (Herzog) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010. Bas.: Aplozia tuberculifera Herzog, Ann. Bryol. 5: 84, 1932.

*** Solenostoma unispire (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010. Bas.: Plectocolea unispiris Amakawa, J. Jap. Bot. 29 (6): 178, 1954.

*** Solenostoma virgatum (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009. Bas.: Plectocolea virgata Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 197, 1891.

*** Solenostoma vulcanicola (Schiffn.) Nyushko ex Potemkin et Sofronova, Liverworts and Hornworts of Russia 1: 289, 2009. Bas.: Nardia vulcanicola Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 191, 1898.

* Solenostoma zangmuii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009. Bas.: Jungermannia zangmuii C.Gao et X.L.Bai, Philipp. Scientist 38: 135, 2001.

** subg. Solenostoma

*** Solenostoma amoenum (Lindenb. et Gottsche) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010. Bas.: Jungermannia amoena Lindenb. et Gottsche, Syn. Hepat. 5: 674, 1847.

*** Solenostoma amplexifolium (Hampe) Váňa et Schäf.-Verw., Acta Bot. Hung. 51 (3/4): 408, 2009. Bas.: Plagiochila amplexifolia Hampe, Nov. Stirp. Pug. 7: 6, 1838.

** Solenostoma appalachianum R.M.Schust. ex Bakalin, Arctoa 23: 127, 2014. Based on: Solenostoma appalachianum R.M.Schust., Rhodora 60 (717): 246, 1958, nom. inval. 92

*** Solenostoma appressifolium (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 494, 2009. Bas.: Jungermannia appressifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861]. 93

*** Solenostoma appressifolium var. minor (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009. Bas.: Jungermannia clavellata var. minor Amakawa, J. Hattori Bot. Lab. 26: 25, 1963.

*** Solenostoma appressifolium var. nigricans (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009. Bas.: Jungermannia decolyana var. nigricans Amakawa, J. Hattori Bot. Lab. 30: 185, 1967.

*** Solenostoma ariadne (Taylor) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009. Bas.: Jungermannia ariadne Taylor, Nov. Stirp. Pug. 8: 9, 1844.

*** Solenostoma atrobrunneum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009. Bas.: Jungermannia atrobrunnea Amakawa, J. Hattori Bot. Lab. 30: 192, 1967.

*** Solenostoma atrorevolutum (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009. Bas.: Jungermannia atrorevoluta Grolle ex Amakawa, J. Hattori Bot. Lab. 29: 255, 1966.

*** Solenostoma atrovirens Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 493 (55), 1901.

*** Solenostoma baueri (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 495 (57), 1901. Bas.: Aplozia baueri Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 195, 1898.

*** Solenostoma bengalense (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009. Bas.: Jungermannia bengalensis Amakawa, J. Hattori Bot. Lab. 31: 112, 1968.

** Solenostoma breviflorum Kashyap et R.S.Chopra, Liverworts W. Himal. 2: 85, 1932.

*** Solenostoma caeleste (Inoue et Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010. Bas.: Jungermannia caelestis Inoue et Váňa, Stud. Cryptog. Papua N. Guinea: 16, 1979.

* Solenostoma caoi (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009. Bas.: Jungermannia caoi C.Gao et X.L.Bai, Philipp. Scientist 38: 152, 2001.

*** Solenostoma caucasicum (Váňa) Konstant., Arctoa 1: 123, 1992. Bas.: Jungermannia caucasica Váňa, Preslia 42: 96, 1970.

*** Solenostoma chenianum (C.Gao, Y.H.Wu et Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009. Bas.: Jungermannia cheniana C.Gao, Y.H.Wu et Grolle, Nova Hedwigia 77 (1/2): 190, 2003.

*** Solenostoma clavellatum Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 491 (53), 1901.

*** Solenostoma confertissimum (Nees) Schljakov, Novosti Sist. Nizš. Rast. 17: 239, 1980. Bas.: Jungermannia confertissima Nees, Naturgesch. Eur. Leberm. 1: 291, 1833.

*** Solenostoma coniflorum (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 497 (59), 1901. Bas.: Jungermannia coniflora Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 10, 1890.

*** Solenostoma crassulum (Nees et Mont.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 497 (59), 1901. Bas.: Jungermannia crassula Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 54, 1836.

** Solenostoma cryptogynum R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007. Based on: Solenostoma cryptogynum R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002, nom. inval.

*** Solenostoma cyclops (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 945, 1969. Bas.: Jungermannia cyclops S.Hatt., J. Hattori Bot. Lab. 3: 5, 1948 [1950].

*** Solenostoma diversiclavellatum (Amakawa et Grolle) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009. Bas.: Jungermannia diversiclavellata Amakawa et Grolle, J. Hattori Bot. Lab. 31: 107, 1968.

*** Solenostoma dulongense Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 497, 2009.

*** Solenostoma exsertum (A.Evans) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 490 (52), 1901. Bas.: Nardia exserta A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891.

*** Solenostoma faurieanum (Beauverd) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 945, 1969. Bas.: Jungermannia faurieana Beauverd, Sp. Hepat. (Stephani) 6: 571, 1924.

*** Solenostoma flagellare (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009. Bas.: Jungermannia flagellaris Amakawa, J. Hattori Bot. Lab. 29: 258, 1966.

*** Solenostoma flavorevolutum (Váňa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009. Bas.: Jungermannia flavorevoluta Váňa, J. Hattori Bot. Lab. 36: 63, 1972 [1973].

*** Solenostoma grollei (D.G.Long et Váňa) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009. Bas.: Gottschelia grollei D.G.Long et Váňa, J. Bryol. 29 (3): 167, 2007.

*** Solenostoma grosseverrucosum (Amakawa et S.Hatt.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015. Bas.: Horikawaella grosseverrucosa Amakawa et S.Hatt., Bull. Univ. Mus. Univ. Tokyo 8: 216, 1975.

*** Solenostoma heterolimbatum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009. Bas.: Jungermannia heterolimbata Amakawa, J. Hattori Bot. Lab. 30: 183, 1967.

*** Solenostoma hewsoniae (Amakawa et Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010. Bas.: Jungermannia hewsoniae Amakawa et Grolle, J. Hattori Bot. Lab. 31: 108, 1968.

*** Solenostoma hiugaense Amakawa, J. Jap. Bot. 31 (2): 47, 1956.

*** Solenostoma indrodayanum (Sushil K.Singh et D.K.Singh) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009. Bas.: Jungermannia indrodayana Sushil K.Singh et D.K.Singh, Cryptog. Bryol. 28 (2): 103, 2007.

*** Solenostoma inundatum (Hook.f. et Taylor) Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 490 (52), 1901. Bas.: Jungermannia inundata Hook.f. et Taylor, London J. Bot. 3: 559, 1844.

*** Solenostoma javanicum (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 494 (56), 1901. Bas.: Aplozia javanica Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 193, 1898.

** Solenostoma kanaii (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009. Bas.: Jungermannia kanaii Amakawa, J. Hattori Bot. Lab. 30: 194, 1967.

*** Solenostoma kashyapii (S.C.Srivast., S.Srivast. et D.Sharma) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009. Bas.: Jungermannia kashyapii S.C.Srivast., S.Srivast. et D.Sharma, Lindbergia 28 (3): 131, 2003.

*** Solenostoma lanigerum (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009. Bas.: Jungermannia lanigera Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861].

*** Solenostoma macrocarpum (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009. Bas.: Jungermannia macrocarpa Schiffn. ex Steph., Sp. Hepat. (Stephani) 6: 87, 1917.

*** Solenostoma mamatkulovii (Váňa et Zerov) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010. Bas.: Jungermannia mamatkulovii Váňa et Zerov, Preslia 49: 181, 1977.

* Solenostoma microphyllum C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 206, 1981.

* Solenostoma microrevolutum (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009. Bas.: Jungermannia microrevoluta C.Gao et X.L.Bai, Philipp. Scientist 38: 146, 2001.

*** Solenostoma mildbraedii (Steph.) R.M.Schust., Beih. Nova Hedwigia 119: 387, 2002. Bas.: Jungermannia mildbraedii Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 113, 1911.

*** Solenostoma niveum (Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010. Bas.: Jungermannia nivea Grolle, Misc. Bryol. Lichenol. 6 (1): 1, 1971.

* Solenostoma novazelandiae R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002. 94

*** Solenostoma ohbae (Amakawa) C.Gao, Bryofl. Xizang: 495, 1985. Bas.: Jungermannia ohbae Amakawa, Bull. Univ. Mus. Univ. Tokyo 8: 218, 1975.

*** Solenostoma orbiculatum (Colenso) R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002. Bas.: Gymnomitrion orbiculatum Colenso, Trans. & Proc. New Zealand Inst. 18: 236, 1886.

*** Solenostoma parvitextum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009. Bas.: Jungermannia parvitexta Amakawa, J. Hattori Bot. Lab. 30: 187, 1967.

*** Solenostoma patoniae (Grolle, D.B.Schill et D.G.Long) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009. Bas.: Gottschelia patoniae Grolle, D.B.Schill et D.G.Long, J. Bryol. 25 (1): 3, 2003.

*** Solenostoma pocsii (Váňa) Bakalin, Arctoa 18: 160, 2009 [2010]. Bas.: Jungermannia pocsii Váňa, Folia Geobot. Phytotax. 10 (4): 365, 1975.

*** Solenostoma poeltii (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009. Bas.: Jungermannia poeltii Amakawa, J. Hattori Bot. Lab. 29: 258, 1966.

*** Solenostoma pseudocyclops (Inoue) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 506, 2009. Bas.: Jungermannia pseudocyclops Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 9 (1): 37, 1966.

** Solenostoma pseudopyriflorum Bakalin et Vilnet, Arctoa 18: 159, 2009 [2010].

*** Solenostoma purpuratum (Mitt.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 489 (51), 1901. Bas.: Jungermannia purpurata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861].

*** Solenostoma pyriflorum Steph., Sp. Hepat. (Stephani) 6: 83, 1917. 95

*** Solenostoma pyriflorum var. gracillimum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 506, 2009. Bas.: Jungermannia pyriflora var. gracillima Amakawa, Fl. E. Himalaya 2: 228, 1971.

** Solenostoma pyriflorum var. major (S.Hatt.) Bakalin, Arctoa 17: 230, 2008 [2009]. Bas.: Jungermannia monticola f. major S.Hatt., J. Hattori Bot. Lab. 3: 8, 1948 [1950].

*** Solenostoma pyriflorum var. minutissimum (Amakawa) Bakalin, Arctoa 16: 208, 2007 [2008]. Bas.: Jungermannia pyriflora var. minutissima Amakawa, J. Hattori Bot. Lab. 22: 61, 1960. 96

*** Solenostoma raujeanum (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009. Bas.: Jungermannia raujeana Grolle ex Amakawa, J. Hattori Bot. Lab. 29: 262, 1966.

*** Solenostoma riclefii Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009. Nom. nov. pro Jungermannia grollei Amakawa, J. Hattori Bot. Lab. 29: 260, 1966.

*** Solenostoma sanguinolentum (Griff.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 489 (51), 1901. Bas.: Jungermannia sanguinolenta Griff., Not. pl. asiat. 2: 302, 1849.

*** Solenostoma schaulianum (Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009. Bas.: Jungermannia schauliana Steph., Sp. Hepat. (Stephani) 6: 90, 1917.

*** Solenostoma shimizuanum (S.Hatt. ex Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010. Bas.: Jungermannia shimizuana S.Hatt. ex Váňa, J. Hattori Bot. Lab. 35: 315, 1972.

*** Solenostoma speciosum (Horik.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009. Bas.: Anastrophyllum speciosum Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 147, 1934.

** Solenostoma speciosum subsp. villosum (R.M.Schust.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 246, 2009. Bas.: Scaphophyllum speciosum subsp. villosum R.M.Schust., Bryologist 101 (3): 434, 1998.

*** Solenostoma sphaerocarpum (Hook.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 499 (61), 1901. Bas.: Jungermannia sphaerocarpa Hook., Brit. Jungermann.: tab. 74, 1815.

*** Solenostoma stephanii (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 496 (58), 1901. Bas.: Aplozia stephanii Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 195, 1898.

*** Solenostoma stoloniferum (Steph.) S.W.Arnell, Hepat. South Africa: 316, 1963. Bas.: Nardia stolonifera Steph., Hedwigia 31 (3): 128, 1892.

*** Solenostoma strictum (Schiffn.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010. Bas.: Aplozia stricta Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 194, 1898.

*** Solenostoma subacutum (Herzog) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015. Bas.: Anastrophyllum subacutum Herzog, Ann. Bryol. 12: 75, 1939.

*** Solenostoma subrubrum (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009. Bas.: Jungermannia subrubra Schiffn. ex Steph., Sp. Hepat. (Stephani) 6: 93, 1917.

* Solenostoma sunii Bakalin et Vilnet, Bot. Pacifica 3 (2): 15, 2014.

* Solenostoma totopapillosum (E.A.Hodgs.) R.M.Schust., Bryologist 100 (3): 366, 1997. Bas.: Jungermannia totopapillosa E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 111, 1972. 97

*** Solenostoma udarii (S.C.Srivast. et P.Singh) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009. Bas.: Jungermannia udarii S.C.Srivast. et P.Singh, Recent Stud. Indian Bryoph.: 152, 1995.

* Solenostoma ventroversum (Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009. Bas.: Jungermannia ventroversa Grolle, Khumbu Himal 1 (4): 284, 1966. 98

*** Solenostoma zantenii (Amakawa) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009. Bas.: Jungermannia zantenii Amakawa, J. Hattori Bot. Lab. 31: 110, 1968.

* Solenostoma zengii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009. Bas.: Jungermannia zengii C.Gao et X.L.Bai, Philipp. Scientist 38: 151, 2001.

*** Southbyaceae Váňa, Crand.-Stotl., Stotler et D.G.Long

by J. Váňa

Southbyaceae has traditionally been placed in Arnelliaceae (e.g. ) or kept as a separate family (e.g. ) that has never been validated. However, it was re-established, rearranged and validated in the study by.

*** Gongylanthus Nees, Naturgesch. Eur. Leberm. 2: 405, 1836.

*** Gongylanthus dusenii Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 385 (41), 1906.

*** Gongylanthus ericetorum (Raddi) Nees, Naturgesch. Eur. Leberm. 2: 407, 1836. Bas.: Calypogeia ericetorum Raddi, Jungermanniogr. Etrusca: 31, 1818.

*** Gongylanthus granatensis (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 385 (41), 1906. Bas.: Lindigina granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 138, 1864.

*** Gongylanthus himalayensis Grolle, Khumbu Himal 1 (4): 285, 1966.

*** Gongylanthus javanicus Grolle, J. Jap. Bot. 40 (7): 206, 1965.

*** Gongylanthus liebmanianus (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 388 (44), 1906. Bas.: Gymnanthe liebmaniana Lindenb. et Gottsche, Syn. Hepat. 5: 712, 1847.

*** Gongylanthus limbatus (Herzog) Grolle et Váňa, Folia Geobot. Phytotax. 9 (2): 198, 1974. Bas.: Aplozia limbata Herzog, Beih. Bot. Centralbl. 61B (3): 561, 1942.

*** Gongylanthus muelleri (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 388 (44), 1906. Bas.: Lindigia muelleri Gottsche, Mexik. Leverm.: 121, 1863.

** Gongylanthus oniscoides (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 386 (42), 1906. Bas.: Calypogeia oniscoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 448, 1885.

*** Gongylanthus richardsii E.W.Jones, Trans. Brit. Bryol. Soc. 4 (4): 650, 1964.

*** Southbya Spruce, Ann. Mag. Nat. Hist. (ser. 2) 3 (18): 501, 1849.

*** Southbya gollanii Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 381 (37), 1906.

*** Southbya nigrella (De Not.) Henriq., Bol. Soc. Brot. 4: 244, 1886 [1887]. Bas.: Jungermannia nigrella De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 1: 315, 1838.

*** Southbya organensis Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 53, 1950.

*** Southbya tophacea (Spruce) Spruce, Ann. Mag. Nat. Hist. (ser. 2) 3 (18): 501, 1849. Bas.: Jungermannia tophacea Spruce, Hep. Pyr. Exsic.: no. 23, 1847.

** Stephaniellaceae R.M.Schust.

by J. Váňa

In the absence of molecular data, the phylogenetic affinities of Stephaniella and Stephaniellidium remain equivocal, but there is no evidence to support their placement in either the Arnelliaceae or Southbyaceae where they have been placed recently. Thus the family as construed by is retained.

*** Stephaniella J.B.Jack, Hedwigia 33 (1): 11, 1894.

* Stephaniella boliviensis Steph., Biblioth. Bot. 87 (2): 182, 1916.

*** Stephaniella hamata Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1024 (87), 1901.

*** Stephaniella paraphyllina J.B.Jack, Hedwigia 33 (1): 11, 1894.

*** Stephaniella rostrata U.Schmitt, Österr. Bot. Z. 115 (2): 124, 1968.

*** Stephaniella uncifolia S.Winkl., Österr. Bot. Z. 115 (2): 124, 1968.

** Stephaniellidium S.Winkl. ex Grolle, Acta Bot. Fenn. 121: 38, 1983. Based on: Stephaniellidium S.Winkl., Mitt. Inst. Colombo-Alemán Invest. Ci. 3: 60, 1969.

*** Stephaniellidium sleumeri (Müll.Frib.) S.Winkl. ex Grolle, Acta Bot. Fenn. 121: 38, 1983. Bas.: Stephaniella sleumeri Müll.Frib., Rev. Bryol. Lichénol. 20 (1/2): 177, 1951.

*** Trichotemnomataceae R.M.Schust.

*** Trichotemnoma R.M.Schust., Nova Hedwigia 15: 440, 1968.

*** Trichotemnoma corrugatum (Steph.) R.M.Schust., Nova Hedwigia 15: 440, 1968. Bas.: Blepharostoma corrugatum Steph., Hedwigia 32 (5): 315, 1893.

*** Blepharostomataceae W.Frey et M.Stech

The recognition of Blepharostomataceae follows following reconsideration of molecular and morphological data available at that time.

*** Blepharostoma (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835. Bas.: Jungermannia sect. Blepharostoma Dumort., Syll. Jungerm. Europ.: 65, 1831.

** Blepharostoma arachnoideum M.Howe, Mem. Torrey Bot. Club 7: 140, 1899.

** Blepharostoma indicum G.Asthana, M.Saxena et Maurya, J. Bryol. 35 (4): 267, 2013.

** Blepharostoma minor Horik., Hikobia 1 (2): 104, 1951 [1952].

*** Blepharostoma trichophyllum (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835. Bas.: Jungermannia trichophylla L., Sp. Pl. 1: 1135, 1753.

** Blepharostoma trichophyllum subsp. brevirete (Bryhn et Kaal.) R.M.Schust., Bull. Natl. Mus. Canada 164: 16, 1959. Bas.: Blepharostoma trichophyllum var. brevirete Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 46, 1906.

*** Brevianthaceae J.J.Engel et R.M.Schust.

by L. Söderström, J. Váňa, R. Stotler, B.J. Crandall-Stotler and J.J. Engel

The treatment of Brevianthaceae follows.

*** Brevianthus J.J.Engel et R.M.Schust., Phytologia 47 (4): 317, 1981.

*** Brevianthus flavus (Grolle) J.J.Engel et R.M.Schust., Phytologia 47 (4): 318, 1981. Bas.: Jackiella flava Grolle, J. Hattori Bot. Lab. 33: 222, 1970.

** Brevianthus flavus subsp. crenulatus J.J.Engel, Nova Hedwigia 93 (3/4): 406, 2011.

* Brevianthus hypocanthidium M.A.M.Renner et J.J.Engel, PhytoKeys 50: 46, 2015.

*** Tetracymbaliella Grolle, Nova Hedwigia 3 (1): 48, 1961.

** Tetracymbaliella comptonii (Pearson) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 164, 1963 [1964]. Bas.: Chiloscyphus comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 23, 1922.

*** Tetracymbaliella cymbalifera (Hook.f. et Taylor) Grolle, Nova Hedwigia 3 (1): 50, 1961. Bas.: Jungermannia cymbalifera Hook.f. et Taylor, Bot. Antarct. Voy. I (Fl. Antarct. 1): 151, 1845.

*** Tetracymbaliella decipiens (Gottsche) Grolle, Nova Hedwigia 3 (1): 49, 1961. Bas.: Chiloscyphus decipiens Gottsche, Syn. Hepat. 2: 176, 1845.

*** Tetracymbaliella subsimplex (Austin) J.J.Engel, Phytotaxa 207 (2): 185, 2015. Bas.: Polyotus subsimplex Austin, Bull. Torrey Bot. Club 6 (7): 46, 1875.

** Chonecoleaceae R.M.Schust. ex Grolle

** Chonecolea Grolle, Rev. Bryol. Lichénol. 25 (3/4): 294, 1956 [1957].

** Chonecolea acutiloba (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 321, 1980. Bas.: Clasmatocolea acutiloba Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 63, 1964.

** Chonecolea andina Grolle et Váňa, J. Hattori Bot. Lab. 48: 229, 1980.

** Chonecolea doellingeri (Nees) Grolle, Rev. Bryol. Lichénol. 25 (3/4): 295, 1956 [1957]. Bas.: Jungermannia doellingeri Nees, Syn. Hepat. 1: 104, 1844.

** Chonecolea ruwenzorensis E.W.Jones, J. Bryol. 13 (4): 498, 1986.

** Chonecolea schusteri Udar et Ad.Kumar, Bryologist 85 (3): 315, 1982.

** Chonecolea verae Potemkin, Proc. int. meeting 90th anniv. Abramova: 165, 2005.

** Grolleaceae Solari ex R.M.Schust.

** Grollea R.M.Schust., Nova Hedwigia 8 (3/4): 288, 1964.

** Grollea antheliopsis R.M.Schust., Nova Hedwigia 8 (3/4): 288, 1964.

*** Herbertaceae Müll.Frib. ex Fulford et Hatcher

by D. Bell

*** Herbertus Gray, Nat. Arr. Brit. Pl. 1: 705, 1821. 99

*** Herbertus aduncus (Dicks.) Gray, Nat. Arr. Brit. Pl. 1: 705, 1821. Bas.: Jungermannia adunca Dicks., Fasc. Pl. Crypt. Brit. 3: 12, 1793.

** Herbertus arcticus (Inoue et Steere) Schljakov, Novosti Sist. Nizš. Rast. 19: 209, 1982. Bas.: Herbertus sakuraii subsp. arcticus Inoue et Steere, J. Hattori Bot. Lab. 44: 266, 1978.

** Herbertus armitanus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 324, 1965. Bas.: Schisma armitanum Steph., Sp. Hepat. (Stephani) 4: 28, 1909.

** Herbertus asparus Tixier, Bryophyt. Biblioth. 18: 64, 1979.

*** Herbertus bivittatus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 343, 1885.

** Herbertus borealis Crundw., Trans. Brit. Bryol. Soc. 6 (1): 41, 1970.

** Herbertus buchii Juslén, Ann. Bot. Fenn. 43 (6): 416, 2006.

** Herbertus ceylanicus (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 43, 1959. Bas.: Schisma ceylanicum Steph., Sp. Hepat. (Stephani) 4: 22, 1909.

** Herbertus circinatus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 318, 1965. Bas.: Schisma circinatum Steph., Sp. Hepat. (Stephani) 4: 25, 1909.

*** Herbertus delavayi (Steph.) Steph., Hedwigia 34 (2): 43, 1895. Bas.: Schisma delavayi Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 228, 1894. 100

*** Herbertus dicranus (Gottsche, Lindenb. et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877. Bas.: Sendtnera dicrana Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 239, 1845.

** Herbertus durandii (Steph.) Herzog, Rev. Bryol. Lichénol. 11 (1): 25, 1938 [1939]. Bas.: Schisma durandii Steph., Trans. Linn. Soc. London, Bot. 6 (1): 99, 1901.

* Herbertus evittatus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 327, 1965. Bas.: Schisma evittatum Steph., Sp. Hepat. (Stephani) 6: 357, 1922.

** Herbertus gaochienii X.Fu, Fl. Bryoph. Sin. 9: 38, 2003.

** Herbertus gracilis (Mont.) Steph., Bull. Herb. Boissier 5 (10): 842, 1897. Bas.: Mastigophora gracilis Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 254, 1843.

** Herbertus guangdongii P.J.Lin ex Piippo, Bryobrothera 1: 206, 1992.

** Herbertus hawaiiensis H.A.Mill., J. Hattori Bot. Lab. 28: 317, 1965.

** Herbertus helleri (Steph.) W.E.Nicholson, Rev. Bryol. Lichénol. 13: 143, 1942. Bas.: Schisma helleri Steph., Sp. Hepat. (Stephani) 4: 29, 1909.

** Herbertus herpocladioides E.B.Scott et H.A.Mill., Bryologist 62 (2): 116, 1959.

*** Herbertus hutchinsiae (Gottsche et Rabenh.) A.Evans, Bull. Torrey Bot. Club 44 (4): 214, 1917. Bas.: Sendtnera adunca var. hutchinsiae Gottsche et Rabenh., Hepat. Eur., Leberm. 21-22: no. 210, 1862.

*** Herbertus juniperoideus (Sw.) Grolle, Rev. Bryol. Lichénol. 30 (1/2): 80, 1961. Bas.: Jungermannia juniperoidea Sw., Prodr. (Swartz): 144, 1788. 101

*** Herbertus juniperoideus subsp. acanthelius (Spruce) K.Feldberg et Heinrichs, Bot. J. Linn. Soc. 151: 326, 2006. Bas.: Herbertus acanthelius Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 341, 1885.

*** Herbertus juniperoideus subsp. pensilis (Taylor) K.Feldberg et Heinrichs, Bot. J. Linn. Soc. 151: 329, 2006. Bas.: Sendtnera pensilis Taylor, London J. Bot. 5: 372, 1846.

** Herbertus kurzii (Steph.) R.S.Chopra, J. Indian Bot. Soc. 22: 247, 1943. Bas.: Schisma kurzii Steph., Sp. Hepat. (Stephani) 4: 24, 1909.

** Herbertus leratii (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 327, 1965. Bas.: Schisma leratii Steph., Sp. Hepat. (Stephani) 6: 360, 1922.

** Herbertus lonchobasis H.A.Mill., J. Hattori Bot. Lab. 28: 306, 1965.

** Herbertus longifissus Steph., Hedwigia 34 (2): 44, 1895.

** Herbertus longispinus J.B.Jack et Steph., Hedwigia 31 (1): 15, 1892.

** Herbertus mauritianus N.G.Hodgetts, J. Bryol. 30 (4): 247, 2008.

*** Herbertus norenus D.G.Long, D.Bell et H.H.Blom, Molec. Ecol. Res. 12: 44, 2012.

** Herbertus oldfieldianus (Steph.) Rodway, Tasm. Bryoph.: 72, 1917. Bas.: Schisma oldfieldianum Steph., Sp. Hepat. (Stephani) 4: 20, 1909.

** Herbertus pilifer Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 252, 1893.

** Herbertus pocsii N.G.Hodgetts, J. Bryol. 30 (4): 249, 2008.

** Herbertus pumilus Steph., Hedwigia 34 (2): 44, 1895.

** Herbertus ramosus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 314, 1965. Bas.: Schisma ramosum Steph., Sp. Hepat. (Stephani) 4: 23, 1909.

** Herbertus runcinatus (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877. Bas.: Sendtnera runcinata Taylor, London J. Bot. 5: 372, 1846.

*** Herbertus sendtneri (Nees) Lindb., Hepat. Scand. Exsicc.: no. 4, 1874. Bas.: Schisma sendtneri Nees, Naturgesch. Eur. Leberm. 3: 575, 1838.

** Herbertus spicatus N.G.Hodgetts, J. Bryol. 30 (4): 244, 2008.

*** Herbertus stramineus (Dumort.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877. Bas.: Schisma stramineum Dumort., Syll. Jungerm. Europ.: 77, 1831.

** Herbertus streimannii M.L.So, Syst. Bot. 28 (1): 13, 2003.

*** Herbertus tenuis A.Evans, Bull. Torrey Bot. Club 44 (4): 219, 1917.

** Herbertus udarii D.Kumar et N.Manocha, Geophytology 29 (1/2): 105, 1999 [2000].

Excluded from the genus

* Herbertus subrotundatus X.Fu et Y.J.Yi, Acta Phytotax. Sin. 39 (1): 89, 2001. 102

*** Triandrophyllum Fulford et Hatcher, Bryologist 64 (4): 349, 1961 [1962]. Based on: Triandrophyllum Fulford et Hatcher, Bryologist 61 (4): 277, 1958 [1959].

*** Triandrophyllum eophyllum (R.M.Schust.) Gradst., Mem. New York Bot. Gard. 86: 104, 2001. Bas.: Olgantha eophylla R.M.Schust., Nova Hedwigia 63 (3/4): 535, 1996.

** Triandrophyllum fernandeziense (S.W.Arnell) Grolle ex Fulford et Hatcher, Bryologist 64 (4): 351, 1961 [1962]. Bas.: Acromastigum fernandeziense S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 10, 1957.

** Triandrophyllum heterophyllum (Steph.) Grolle, J. Jap. Bot. 39 (8): 238, 1964. Bas.: Mastigophora heterophylla Steph., Sp. Hepat. (Stephani) 6: 367, 1922.

*** Triandrophyllum subtrifidum (Hook.f. et Taylor) Fulford et Hatcher, Bryologist 64 (4): 350, 1961 [1962]. Bas.: Jungermannia subtrifida Hook.f. et Taylor, London J. Bot. 3: 579, 1844.

** Triandrophyllum subtrifidum var. trifidum (Gottsche) Solari, Bol. Soc. Argent. Bot. 15 (2/3): 201, 1973. Bas.: Sendtnera trifida Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 142, 1864.

** Triandrophyllum symmetricum J.J.Engel, Haussknechtia, Beih. 9: 115, 1999.

*** Lepicoleaceae R.M.Schust.

by M. von Konrat

Lepicoleaceae has conventionally been considered a monogeneric family only containing Lepicolea (e.g. ). However, we expand Lepicoleaceae here to include the monogeneric family Vetaformaceae. A number of molecular analyses have shown that Vetaforma dusenii is sister to Lepicolea (e.g., ) together forming a robust clade.

*** Lepicolea Dumort., Recueil Observ. Jungerm.: 20, 1835.

*** Lepicolea attenuata (Mitt.) Steph., J. Linn. Soc., Bot. 29 (201): 276, 1892. Bas.: Sendtnera attenuata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 153, 1854.

*** Lepicolea magellanica (Gola) Solari, Lindbergia 9 (2): 86, 1983. Bas.: Lepicolea scolopendra var. magellanica Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 29 (1/4): 170, 1922 [1923].

*** Lepicolea norrisii Piippo, Ann. Bot. Fenn. 25 (1): 55, 1988.

*** Lepicolea ochroleuca (Spreng.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 345, 1885. Bas.: Jungermannia ochroleuca Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827.

*** Lepicolea pruinosa (Taylor) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 345, 1885. Bas.: Sendtnera pruinosa Taylor, London J. Bot. 5: 373, 1846.

*** Lepicolea ramentifissa Herzog, Biblioth. Bot. 88: 30, 1920 [1921].

*** Lepicolea rara (Steph.) Grolle, Nova Hedwigia 16: 152, 1968. Bas.: Lepidozia rara Steph., Sp. Hepat. (Stephani) 3: 618, 1909. 103

*** Lepicolea rigida (De Not.) E.B.Scott, Nova Hedwigia 2: 148, 1960. Bas.: Sendtnera rigida De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 229, 1857.

*** Lepicolea scolopendra (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 398, 1877. Bas.: Jungermannia scolopendra Hook., Musci Exot. 1: tab. 40, 1818.

*** Lepicolea yakusimensis (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 10: 42, 1953. Bas.: Lepicolea scolopendra var. yakusimensis S.Hatt., J. Hattori Bot. Lab. 2: 9, 1947 [1948].

*** Vetaforma Fulford et J.Taylor, Nova Hedwigia 4 (1/2): 81, 1962.

*** Vetaforma dusenii (Steph.) Fulford et J.Taylor, Nova Hedwigia 4 (1/2): 82, 1962. Bas.: Lepidozia dusenii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 52, 1900.

*** Lepidoziaceae Limpr.

by E.D. Cooper

The classification within Lepidoziaceae is in a state of flux. The treatment provided here follows the interim classification proposed by based on recent molecular phylogenetic studies incorporating nomenclatural changes from. Several higher taxa are unlikely to represent monophyletic units, but the phylogenetic data currently available are insufficient to re-circumscribe the doubtful subfamilies and genera. Infrageneric taxa have been retained where phylogenetic data are absent or inconclusive.

** Meinungeria Frank Müll., Bryologist 110 (3): 494, 2007.

** Meinungeria mouensis Frank Müll., Bryologist 110 (3): 494, 2007.

*** Bazzanioideae Rodway

*** Acromastigum A.Evans, Bull. Torrey Bot. Club 27 (3): 103, 1900. 104

** subg. Acromastigum

*** Acromastigum caledonicum (Steph.) Grolle, Österr. Bot. Z. 111 (2/3): 243, 1964. Bas.: Acolea caledonica Steph., Nova Caledonia, Bot. 1: 19, 1914.

*** Acromastigum cavifolium R.M.Schust., J. Hattori Bot. Lab. 26: 257, 1963.

** Acromastigum herzogii Grolle, Österr. Bot. Z. 111 (2/3): 250, 1964.

*** Acromastigum homodictyon (Herzog) Grolle, Österr. Bot. Z. 111 (2/3): 245, 1964. Bas.: Acromastigum integrifolium var. homodictyon Herzog, Ark. Bot. (n.ser.) 3 (3): 44, 1953.

** Acromastigum integrifolium (Austin) A.Evans, Bull. Torrey Bot. Club 27 (3): 103, 1900. Bas.: Mastigobryum integrifolium Austin, Bot. Bull. (Hanover) 1 (7): 32, 1876.

*** Acromastigum stellare N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 109, 1985.

** Acromastigum stenophyllum R.M.Schust., Nova Hedwigia 15: 465, 1968.

*** Acromastigum verticale (Steph.) E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 18, 1954. Bas.: Bazzania verticalis Steph., Hedwigia 32 (4): 214, 1893.

** subg. Inaequilatera (Schiffn.) Grolle, J. Hattori Bot. Lab. 44: 2, 1978. Bas.: Bazzania sect. Inaequilaterae Schiffn., Hepat. (Engl.-Prantl): 101, 1893.

*** Acromastigum adaptatum Hürl., Bauhinia 7 (4): 263, 1983.

*** Acromastigum anisostomum (Lehm. et Lindenb.) A.Evans, Ann. Bryol., Suppl. 3: 48, 1934. Bas.: Jungermannia anisostoma Lehm. et Lindenb., Nov. Stirp. Pug. 6: 57, 1834.

** Acromastigum anisostomum var. minutum E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 71, 1965.

** Acromastigum aurescens A.Evans, Ann. Bryol., Suppl. 3: 45, 1934.

** Acromastigum bancanum (Sande Lac.) A.Evans, Ann. Bryol., Suppl. 3: 20, 1934. Bas.: Mastigobryum bancanum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 301, 1864.

** Acromastigum brotheri (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 70, 1934. Bas.: Mastigobryum brotheri Steph., Sp. Hepat. (Stephani) 3: 536, 1909.

** Acromastigum capillare (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 37, 1934. Bas.: Mastigobryum capillare Steph., Sp. Hepat. (Stephani) 6: 457, 1924.

*** Acromastigum colensoanum (Mitt.) A.Evans ex Reimers, Hedwigia 73 (3/4): 142, 1933. Bas.: Mastigobryum colensoanum Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 147, 1854.

*** Acromastigum cunninghamii (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 106, 1934. Bas.: Bazzania cunninghamii Steph., Hedwigia 32 (4): 205, 1893.

** Acromastigum curtilobum A.Evans, Ann. Bryol., Suppl. 3: 97, 1934.

*** Acromastigum divaricatum (Nees) A.Evans ex Reimers, Hedwigia 73 (3/4): 142, 1933. Bas.: Mastigobryum divaricatum Nees, Syn. Hepat. 2: 219, 1845.

** Acromastigum echinatiforme (De Not.) A.Evans, Ann. Bryol., Suppl. 3: 64, 1934. Bas.: Mastigobryum echinatiforme De Not., Epat. Borneo: 38, 1874.

* Acromastigum echinatum (Gottsche) A.Evans, Ann. Bryol., Suppl. 3: 147, 1934. Bas.: Mastigobryum echinatum Gottsche, Syn. Hepat. 2: 218, 1845. 105

*** Acromastigum exiguum (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 75, 1934. Bas.: Mastigobryum exiguum Steph., Hedwigia 25 (1): 6, 1886.

** Acromastigum exile (Lindenb.) A.Evans, Ann. Bryol., Suppl. 3: 24, 1934. Bas.: Mastigobryum exile Lindenb., Syn. Hepat. 2: 217, 1845.

*** Acromastigum filum (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 31, 1934. Bas.: Bazzania filum Steph., Hedwigia 32 (4): 206, 1893.

*** Acromastigum filum var. papillosum N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 112, 1985.

** Acromastigum fimbriatum (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 153, 1934. Bas.: Mastigobryum fimbriatum Steph., Sp. Hepat. (Stephani) 3: 538, 1909.

** Acromastigum furcatifolium (Steph.) E.A.Br., Phytotaxa 65: 58, 2012. Bas.: Lepidozia furcatifolia Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 112, 1914.

*** Acromastigum inaequilaterum (Lehm. et Lindenb.) A.Evans, Ann. Bryol., Suppl. 3: 129, 1934. Bas.: Jungermannia inaequilatera Lehm. et Lindenb., Nov. Stirp. Pug. 6: 56, 1834.

** Acromastigum interstisiale E.A.Br. et M.A.M.Renner, Telopea 17: 268, 2014.

** Acromastigum laetevirens (Sande Lac. ex Steph.) A.Evans, Ann. Bryol., Suppl. 3: 94, 1934. Bas.: Mastigobryum laetevirens Sande Lac. ex Steph., Hedwigia 25 (4): 133, 1886.

** Acromastigum laevigatum A.Evans, Ann. Bryol., Suppl. 3: 101, 1934.

** Acromastigum linganum (De Not.) A.Evans, Ann. Bryol., Suppl. 3: 118, 1934. Bas.: Mastigobryum linganum De Not., Epat. Borneo: 37, 1874.

** Acromastigum lobuliferum A.Evans, Ann. Bryol., Suppl. 3: 157, 1934.

** Acromastigum longirete Grolle, J. Hattori Bot. Lab. 44: 11, 1978.

*** Acromastigum mooreanum (Steph.) E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 19, 1954. Bas.: Bazzania mooreana Steph., Hedwigia 33 (1): 1, 1894.

*** Acromastigum moratii N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 119, 1985.

** Acromastigum obliquatum (Mitt.) A.Evans, Ann. Bryol., Suppl. 3: 110, 1934. Bas.: Bazzania obliquata Mitt., Hedwigia 32 (4): 211, 1893.

** Acromastigum tenax (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 41, 1934. Bas.: Mastigobryum tenax Steph., Sp. Hepat. (Stephani) 6: 483, 1924.

Incertae sedis

** Acromastigum fumosum E.A.Br. et M.A.M.Renner, Telopea 17: 274, 2014.

** Acromastigum leptophyllum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 309, 1950.

*** Acromastigum marginatum E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 22, 1954.

** Acromastigum microstictum A.Evans, Ann. Bryol., Suppl. 3: 115, 1934.

** Acromastigum prismaticale E.A.Br. et M.A.M.Renner, Telopea 17: 281, 2014.

** Acromastigum pusillum N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 116, 1985.

** Acromastigum rigidum R.M.Schust., Nova Hedwigia 64 (3/4): 617, 1997.

** Acromastigum subechinatiforme Hürl., Bauhinia 7 (4): 266, 1983.

*** Bazzania Gray, Nat. Arr. Brit. Pl. 1: 704, 1821 nom. conserv. 106

*** Bazzania acanthostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 381, 1885.

*** Bazzania accreta (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum accretum Lehm. et Lindenb., Syn. Hepat. 2: 222, 1845.

** Bazzania acinaciformis Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 290, 1907.

** Bazzania acuminata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum acuminatum Lindenb. et Gottsche, Syn. Hepat. 5: 719, 1847.

** Bazzania acutifolia (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 147, 1898. Bas.: Mastigobryum acutifolium Steph., Hedwigia 24 (5): 214, 1885.

*** Bazzania adnexa (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Jungermannia adnexa Lehm. et Lindenb., Nov. Stirp. Pug. 4: 58, 1832.

** Bazzania adnexa var. aucklandica (Lindenb. et Gottsche) J.J.Engel et G.L.Merr., Bryologist 97 (3): 319, 1994. Bas.: Mastigobryum novae-hollandiae f. aucklandicum Lindenb. et Gottsche, Sp. Hepat. (Lindenberg) 8-11: 33, 1851.

*** Bazzania affinis (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Mastigobryum affine Lindenb. et Gottsche, Syn. Hepat. 5: 720, 1847.

** Bazzania albifolia Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 195, 1934.

** Bazzania ambigua (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum ambiguum Lindenb., Syn. Hepat. 2: 217, 1845.

*** Bazzania amblyphylla Meagher, Nova Hedwigia 92 (3/4): 487, 2011.

** Bazzania aneityensis (Steph.) Tixier, Bull. Mus. Natl. Hist. Nat. (Sér. 3), Bot. 10 (190): 80, 1973. Bas.: Mastigobryum aneityense Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 121, 1914.

** Bazzania angusta (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 307, 1950. Bas.: Mastigobryum angustum Steph., Sp. Hepat. (Stephani) 6: 453, 1924.

** Bazzania angustifalcata Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 166, 1955.

** Bazzania angustifolia Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 198, 1934.

** Bazzania angustisedens (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 445, 1972 [1973]. Bas.: Mastigobryum angustisedens Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 745 (429), 1908.

** Bazzania angustistipula N.Kitag., J. Hattori Bot. Lab. 30: 268, 1967.

*** Bazzania appendiculata (Mitt.) S.Hatt., Fl. E. Himalaya: 505, 1966. Bas.: Mastigobryum appendiculatum Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 105, 1860 [1861].

** Bazzania approximata Onr., Bull. Jard. Bot. Natl. Belg. 47 (1/2): 139, 1977.

** Bazzania arcuata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum arcuatum Lindenb. et Gottsche, Syn. Hepat. 5: 718, 1847.

** Bazzania arcuata var. mamillosa Gradst. et A.R.Benitez, Nova Hedwigia 99 (1/2): 113, 2014.

** Bazzania armatistipula (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 114, 1946. Bas.: Mastigobryum armatistipulum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 866 (490), 1908.

** Bazzania asperrima Steph., Rev. Bryol. 34 (3): 48, 1907.

* Bazzania asymmetrica (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 28 (2): 77, 1979. Bas.: Mastigobryum asymmetricum Steph., Sp. Hepat. (Stephani) 6: 454, 1924. 107

** Bazzania aterrima (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 26 (2): 77, 1977. Bas.: Mastigobryum aterrimum Steph., Sp. Hepat. (Stephani) 6: 454, 1924.

*** Bazzania aurescens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 374, 1885.

** Bazzania avia Meagher, Nova Hedwigia 97 (3/4): 529, 2013.

** Bazzania azorica H.Buch et Perss., Bryophyt. Azoren Madeira: 3, 1941.

** Bazzania baldwinii Austin, Trans. Connecticut Acad. Arts 8 (15): 255, 1891.

** Bazzania bernieri (Steph.) Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 142, 1965. Bas.: Mastigobryum bernieri Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 852 (476), 1908.

*** Bazzania bescherellei Steph., Hedwigia 32 (4): 204, 1893.

** Bazzania bhutanica N.Kitag. et Grolle, J. Hattori Bot. Lab. 61: 269, 1986 [1987].

** Bazzania bicrenata N.Kitag., J. Hattori Bot. Lab. 47: 127, 1980.

*** Bazzania bidens (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Mastigobryum bidens Gottsche et Lindenb., Syn. Hepat. 2: 228, 1845.

** Bazzania bidens var. heterodonta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 372, 1885.

** Bazzania bidentula (Steph.) Yasuda, Shokubutsugaku Kakuron: 711, 1911. Bas.: Pleuroschisma bidentulum Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 222, 1894.

*** Bazzania bilobata N.Kitag., J. Hattori Bot. Lab. 30: 257, 1967.

** Bazzania borneensis N.Kitag., J. Hattori Bot. Lab. 37: 263, 1973. Nom. nov. pro Mastigobryum borneense Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 839 (463), 1908, nom. illeg.

** Bazzania brasiliensis (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Mastigobryum brasiliense Gottsche et Lindenb., Syn. Hepat. 2: 227, 1845.

** Bazzania brighamii (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 255, 1891. Bas.: Mastigobryum brighamii Austin, Bull. Torrey Bot. Club 5 (3): 16, 1874.

** Bazzania cadens N.Kitag., J. Hattori Bot. Lab. 47: 129, 1980.

*** Bazzania calcarata (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 149, 1898. Bas.: Mastigobryum calcaratum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 304, 1864.

** Bazzania callida (Sande Lac. ex Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959. Bas.: Mastigobryum callidum Sande Lac. ex Steph., Hedwigia 24 (6): 246, 1885.

*** Bazzania canelensis (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 152, 1946. Bas.: Mastigobryum canelense Steph., Sp. Hepat. (Stephani) 3: 518, 1909.

** Bazzania caudata (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 301, 1950. Bas.: Mastigobryum caudatum Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 947 (497), 1908.

*** Bazzania caudistipula (Steph.) Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 141, 1965. Bas.: Mastigobryum caudistipulum Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 945 (495), 1908.

** Bazzania ceylanica (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 306, 1896. Bas.: Mastigobryum ceylanicum Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 105, 1860 [1861].

** Bazzania chilensis (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 51, 1946. Bas.: Mastigobryum chilense Steph., Hedwigia 24 (6): 247, 1885.

*** Bazzania chimantensis Fulford, Bryologist 63 (2): 89, 1960.

** Bazzania cincinnata (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum cincinnatum De Not., Epat. Borneo: 34, 1874.

*** Bazzania citharodes Meagher, Nova Hedwigia 86 (3/4): 481, 2008.

** Bazzania combinata (J.B.Jack et Steph.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 306, 1896. Bas.: Mastigobryum combinatum J.B.Jack et Steph., Bot. Centralbl. 60 (4): 102, 1894.

** Bazzania commutata (Lindenb. et Gottsche) Schiffn., Consp. Hepat. Arch. Ind.: 149, 1898. Bas.: Mastigobryum commutatum Lindenb. et Gottsche, Sp. Hepat. (Lindenberg) 8-11: 97, 1851.

* Bazzania comorensis Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 197, 1891 [1892].

** Bazzania confertifolia (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 298, 1950. Bas.: Mastigobryum confertifolium Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 858 (482), 1908.

** Bazzania conistipula (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981. Bas.: Mastigobryum conistipulum Steph., Sp. Hepat. (Stephani) 6: 458, 1924.

** Bazzania conophylla (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 150, 1898. Bas.: Mastigobryum conophyllum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 304, 1864.

** Bazzania consanguinea (Hampe et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum consanguineum Hampe et Lindenb., Linnaea 20 (3): 327, 1847.

** Bazzania consociata (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981. Bas.: Mastigobryum consociatum Steph., Sp. Hepat. (Stephani) 6: 458, 1924.

* Bazzania corbieri (Steph.) Meagher, Nova Hedwigia 86 (3/4): 483, 2008. Bas.: Mastigobryum corbieri Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 122, 1914. 108

** Bazzania crassidentata Fulford, Bull. Torrey Bot. Club 86 (5): 338, 1959.

** Bazzania crassitexta Steph., Hedwigia 32 (4): 205, 1893.

* Bazzania crenata (Trevis.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Pleuroschisma crenatum Trevis., Herb. Crypt. Trev. 2: 30, 1853 (; non vidi). 109

** Bazzania cubensis (Gottsche ex Steph.) Pagán, Bryologist 42 (2): 38, 1939. Bas.: Mastigobryum cubense Gottsche ex Steph., Hedwigia 24 (6): 248, 1885.

** Bazzania cucullata Onr., Bull. Jard. Bot. Natl. Belg. 47 (1/2): 142, 1977.

*** Bazzania cuneistipula (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum cuneistipulum Gottsche et Lindenb., Syn. Hepat. 2: 225, 1845.

* Bazzania curvidens Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 197, 1891 [1892].

** Bazzania debilis N.Kitag., J. Hattori Bot. Lab. 30: 256, 1967.

** Bazzania deciduifolia Onr., Bull. Jard. Bot. Natl. Belg. 47 (1/2): 144, 1977.

*** Bazzania decrescens (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Jungermannia decrescens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 57, 1832. 110

** Bazzania decrescens var. dentistipula Kiaer et Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 6, 1893.

** Bazzania decrescens subsp. molleri (Steph.) E.W.Jones, J. Bryol. 8 (3): 303, 1975. Bas.: Mastigobryum molleri Steph., Bot. Jahrb. Syst. 8 (2): 84, 1886.

** Bazzania decrescens subsp. pumila (Mitt.) Pócs, Trop. Bryol. 9: 129, 1994. Bas.: Bazzania pumila Mitt., J. Linn. Soc., Bot. 22 (146): 322, 1886.

** Bazzania densa (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 151, 1898. Bas.: Mastigobryum densum Sande Lac., Ned. Kruidk. Arch. 3: 418, 1854 [1855].

** Bazzania densa var. connata (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 151, 1898. Bas.: Mastigobryum densum β connatum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 302, 1864.

*** Bazzania denticulata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum denticulatum Lindenb. et Gottsche, Syn. Hepat. 5: 718, 1847.

** Bazzania denticulifera Mägd., Nova Hedwigia 38: 53, 1983.

** Bazzania denudata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum denudatum Lindenb. et Gottsche, Syn. Hepat. 2: 216, 1845.

*** Bazzania deplanchei (Gottsche) Jovet-Ast, Rev. Bryol. Lichénol. 18 (1/2): 83, 1949. Bas.: Mastigobryum deplanchei Gottsche, Bull. Herb. Boissier (sér. 2) 8 (12): 955 (505), 1908.

** Bazzania deplanchei var. filamentosa Tixier, Cryptog. Bryol. Lichénol. 6 (2): 179, 1985.

** Bazzania desciscens (Steph.) Meijer, Blumea 10 (2): 382, 1960. Bas.: Mastigobryum desciscens Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 862 (486), 1908.

** Bazzania didericiana (Gottsche ex Steph.) Steph., Bull. Herb. Boissier 5 (10): 841, 1897. Bas.: Mastigobryum didericianum Gottsche ex Steph., Hedwigia 24 (6): 249, 1885.

** Bazzania diminuta Herzog, Trans. Brit. Bryol. Soc. 1 (4): 305, 1950.

* Bazzania distans (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum distans Nees, Syn. Hepat. 2: 216, 1845. 111

*** Bazzania diversicuspis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 373, 1885.

** Bazzania drepanophylla Herzog, Trans. Brit. Bryol. Soc. 1 (4): 301, 1950.

** Bazzania dulitensis Herzog, Trans. Brit. Bryol. Soc. 1 (4): 299, 1950.

** Bazzania dulongensis L.P.Zhou et Li Zhang, J. Bryol. 34 (1): 22, 2012.

** Bazzania eggersiana (Steph.) Pagán, Bryologist 42 (2): 39, 1939. Bas.: Mastigobryum eggersianum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 844 (468), 1908.

** Bazzania elmeri (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 446, 1972 [1973]. Bas.: Mastigobryum elmeri Steph., Leafl. Philipp. Bot. 6: 2289, 1914.

** Bazzania elongata Fulford, Bull. Torrey Bot. Club 86 (5): 337, 1959.

** Bazzania emarginata (Steph.) C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 17, 1904. Bas.: Mastigobryum didericianum var. emarginatum Steph., Hedwigia 24 (6): 249, 1885.

** Bazzania engelii Glenny, Fieldiana, Bot. (n.ser.) 47: 176, 2007.

*** Bazzania erosa (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Jungermannia erosa Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 230, 1824 [1825].

** Bazzania erosa var. pulopenangensis (Lindenb. et Gottsche) Schiffn., Consp. Hepat. Arch. Ind.: 157, 1898. Bas.: Mastigobryum erosum δ pulopenangense Lindenb. et Gottsche, Sp. Hepat. (Lindenberg) 8-11: 99, 1851.

*** Bazzania exempta J.J.Engel, J. Hattori Bot. Lab. 99: 197, 2006.

*** Bazzania falcata (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Mastigobryum falcatum Lindenb., Syn. Hepat. 2: 231, 1845.

** Bazzania falcifolia (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981. Bas.: Mastigobryum falcifolium Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 33, 1911.

* Bazzania fallax (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 158, 1898. Bas.: Mastigobryum fallax Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 304, 1864.

*** Bazzania fasciculata (Steph.) Meagher, Australas. Bryol. Newslett. 46: 6, 2002. Bas.: Mastigobryum fasciculatum Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 748 (432), 1908.

** Bazzania fauriana (Steph.) S.Hatt., Bot. Mag. (Tokyo) 59 (693/694): 27, 1946. Bas.: Mastigobryum faurianum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 843 (467), 1908.

** Bazzania filiformis Steph., Hedwigia 28 (2): 131, 1889.

*** Bazzania flaccida (Dumort.) Grolle, Lindbergia 1 (3/4): 200, 1972 [1973]. Bas.: Pleuroschisma flaccidum Dumort., Syll. Jungerm. Europ.: 71, 1831.

** Bazzania flavescens (Sande Lac. ex Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 158, 1898. Bas.: Mastigobryum flavescens Sande Lac. ex Steph., Hedwigia 25 (1): 6, 1886.

** Bazzania fleischeri (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959. Bas.: Mastigobryum fleischeri Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 773 (457), 1908.

* Bazzania francana (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 446, 1972 [1973]. Bas.: Mastigobryum francanum Steph., Sp. Hepat. (Stephani) 6: 463, 1924.

** Bazzania friabilis N.Kitag. et T.Kodama, J. Hattori Bot. Lab. 39: 67, 1975.

*** Bazzania fuhreri Meagher, Nova Hedwigia 92 (3/4): 488, 2011.

** Bazzania fuscescens A.Evans, Pap. Michigan Acad. Sci. 17: 85, 1932 [1933].

*** Bazzania gamscottii Meagher, Nova Hedwigia 92 (3/4): 492, 2011.

*** Bazzania gedeana (Steph.) Meijer, Blumea 10 (2): 378, 1960. Bas.: Mastigobryum gedeanum Steph., Sp. Hepat. (Stephani) 3: 540, 1909.

*** Bazzania gracilis (Hampe et Gottsche) Steph., Hedwigia 27 (11/12): 279, 1888. Bas.: Mastigobryum gracile Hampe et Gottsche, Linnaea 25 (3): 346, 1852 [1853].

** Bazzania grandiretis (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 298, 1950. Bas.: Mastigobryum grandirete Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 747 (431), 1908.

** Bazzania griffithiana (Steph.) Mizut., J. Hattori Bot. Lab. 30: 82, 1967. Bas.: Mastigobryum griffithianum Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 959 (509), 1908.

** Bazzania gunniana (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981. Bas.: Mastigobryum gunnianum Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 123, 1914.

** Bazzania hainanensis L.P.Zhou et Li Zhang, J. Bryol. 34 (1): 25, 2012.

** Bazzania halconiensis (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 447, 1972 [1973]. Bas.: Mastigobryum halconiense Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 759 (443), 1908.

** Bazzania hamatifolia (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981. Bas.: Mastigobryum hamatifolium Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 862 (486), 1908.

** Bazzania harpago (De Not.) Schiffn., Consp. Hepat. Arch. Ind.: 159, 1898. Bas.: Mastigobryum harpago De Not., Epat. Borneo: 29, 1874.

** Bazzania hebridensis (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981. Bas.: Mastigobryum hebridense Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 124, 1914.

** Bazzania herminieri (Gottsche ex Steph.) Pagán, Bryologist 45 (4): 90, 1942. Bas.: Mastigobryum herminieri Gottsche ex Steph., Hedwigia 25 (1): 8, 1886.

** Bazzania herzogiana Meijer, Blumea 10 (2): 371, 1960. Nom. nov. pro Bazzania remotifolia Herzog, Trans. Brit. Bryol. Soc. 1 (4): 304, 1950, nom. illeg.

** Bazzania heterostipa (Steph.) Fulford, Bull. Torrey Bot. Club 86 (6): 410, 1959. Bas.: Mastigobryum heterostipum Steph., Sp. Hepat. (Stephani) 3: 532, 1909.

*** Bazzania himalayana (Mitt.) Schiffn., Österr. Bot. Z. 49 (4): 132, 1899. Bas.: Mastigobryum himalayanum Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 105, 1860 [1861].

*** Bazzania hochstetteri (Reichardt) E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 11, 1954. Bas.: Mastigobryum hochstetteri Reichardt, Verh. K.K. Zool.-Bot. Ges. Wien 16: 959, 1866.

*** Bazzania hookeri (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum hookeri Lindenb., Syn. Hepat. 2: 226, 1845.

** Bazzania horridula Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 258, 1893.

** Bazzania inaequabilis Steph., Trans. Connecticut Acad. Arts 12 (1): 21, 1904.

** Bazzania inaequitexta Steph., Hedwigia 32 (4): 208, 1893.

** Bazzania incrassata (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 448, 1972 [1973]. Bas.: Mastigobryum incrassatum Steph., Sp. Hepat. (Stephani) 6: 469, 1924.

** Bazzania indica (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Mastigobryum indicum Gottsche et Lindenb., Syn. Hepat. 2: 230, 1845.

** Bazzania indigenarum (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 26 (2): 82, 1977. Bas.: Mastigobryum indigenarum Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 124, 1914.

** Bazzania insignis (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum insigne De Not., Epat. Borneo: 26, 1874.

* Bazzania intermedia (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Mastigobryum intermedium Gottsche et Lindenb., Sp. Hepat. (Lindenberg) 8-11: 82, 1851.

** Bazzania intermedia var. sarawakiana (De Not.) Schiffn., Consp. Hepat. Arch. Ind.: 162, 1898. Bas.: Mastigobryum intermedium var. sarawakianum De Not., Epat. Borneo: 32, 1874.

** Bazzania involuta (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Herpetium involutum Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 253, 1843.

** Bazzania involuta var. submutica (Lindenb. et Gottsche) J.J.Engel et G.L.Merr., Bryologist 97 (3): 314, 1994. Bas.: Mastigobryum novae-hollandiae γ3 submuticum Lindenb. et Gottsche, Sp. Hepat. (Lindenberg) 8-11: 33, 1851.

** Bazzania involutiformis (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum involutiforme De Not., Epat. Borneo: 28, 1874.

** Bazzania irregularis (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 163, 1898. Bas.: Mastigobryum irregulare Steph., Hedwigia 25 (4): 133, 1886.

*** Bazzania jamaicensis (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Herpetium jamaicense Lehm. et Lindenb., Nov. Stirp. Pug. 7: 7, 1838.

** Bazzania japonica (Sande Lac.) Lindb., Acta Soc. Sci. Fenn. 10: 224, 1872 [1873]. Bas.: Mastigobryum japonicum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 303, 1864.

** Bazzania japonica var. sumatrana Herzog, Ann. Naturhist. Mus. Wien 53 (1): 366, 1942 [1943].

** Bazzania javanica (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 163, 1898. Bas.: Mastigobryum javanicum Sande Lac., Ned. Kruidk. Arch. 3: 418, 1854 [1855].

** Bazzania kernii Steph., Hedwigia 32 (4): 208, 1893.

** Bazzania kokawana N.Kitag. et T.Kodama, J. Jap. Bot. 50 (1): 11, 1975.

** Bazzania latifolia Steph., Hedwigia 32 (4): 209, 1893.

* Bazzania lehmanniana (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum lehmannianum Lindenb., Syn. Hepat. 2: 223, 1845. 112

** Bazzania leratii (Beauverd) H.A.Mill., Phytologia 47 (4): 320, 1981. Bas.: Mastigobryum leratii Beauverd, Sp. Hepat. (Stephani) 6: 477, 1924.

** Bazzania lessonii (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981. Bas.: Mastigobryum lessonii Steph., Sp. Hepat. (Stephani) 3: 531, 1909.

** Bazzania levieri (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 448, 1972 [1973]. Bas.: Mastigobryum levieri Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 944 (494), 1908.

** Bazzania liebmaniana (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum liebmanianum Lindenb. et Gottsche, Syn. Hepat. 5: 719, 1847.

** Bazzania linearis Herzog, Trans. Brit. Bryol. Soc. 1 (4): 303, 1950.

** Bazzania linguiformis (Sande Lac.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum linguiforme Sande Lac., Plagiochila Sandei: 8, 1856.

*** Bazzania longa (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Jungermannia longa Nees, Linnaea 6 (4): 623, 1831.

** Bazzania longa var. papillata (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 94, 1946. Bas.: Mastigobryum papillatum Steph., Sp. Hepat. (Stephani) 3: 526, 1909.

*** Bazzania longicaulis (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 165, 1898. Bas.: Mastigobryum longicaule Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 303, 1864.

** Bazzania longicaulis var. latiareata Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 46, 1950 [1951].

*** Bazzania longistipula (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Mastigobryum longistipulum Lindenb., Syn. Hepat. 2: 228, 1845.

*** Bazzania loricata (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Jungermannia loricata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 233, 1824 [1825].

** Bazzania lowii (Sande Lac. ex Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 166, 1898. Bas.: Mastigobryum lowii Sande Lac. ex Steph., Hedwigia 25 (5): 204, 1886.

** Bazzania luzonensis (Steph.) Del Ros., Philipp. J. Sci. 100 (3/4): 231, 1971. Bas.: Mastigobryum luzonense Steph., Sp. Hepat. (Stephani) 6: 472, 1924.

** Bazzania macgregorii Steph., Hedwigia 32 (4): 210, 1893.

** Bazzania macrostipula Fulford, Bull. Torrey Bot. Club 86 (6): 407, 1959.

** Bazzania magna Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 197, 1934.

** Bazzania magnistipula N.Kitag., J. Hattori Bot. Lab. 47: 132, 1980.

** Bazzania malaccensis (Steph.) Tixier, Gard. Bull. Singapore 25 (3): 342, 1971. Bas.: Mastigobryum malaccense Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 944 (494), 1908.

** Bazzania manillana (Gottsche ex Steph.) S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 113, 1951. Bas.: Mastigobryum manillanum Gottsche ex Steph., Hedwigia 25 (5): 204, 1886.

* Bazzania marginata (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 449, 1972 [1973]. Bas.: Mastigobryum marginatum Steph., Rev. Bryol. 35 (2): 31, 1908.

** Bazzania marginella (Herzog) N.Kitag. et T.Kodama, Bull. Osaka Mus. Nat. Hist. 27: 17, 1973. Bas.: Mastigobryum marginellum Herzog, Ann. Bryol. 5: 91, 1932.

* Bazzania mascarena (Steph.) Herzog, Bot. Not. 100 (4): 334, 1947. Bas.: Mastigobryum mascarenum Steph., Hedwigia 25 (5): 205, 1886.

** Bazzania mayebarae S.Hatt., J. Hattori Bot. Lab. 19: 91, 1958.

* Bazzania menzelii E.D.Cooper, Phytotaxa 97 (2): 52, 2013. Nom. nov. pro Acromastigum emarginatum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 309, 1950.

** Bazzania merrillana (Steph.) Inoue ex Bonner, Index Hepat. 3: 359, 1963. Bas.: Mastigobryum merrillanum Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 944 (494), 1908.

** Bazzania minuta (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 255, 1891. Bas.: Mastigobryum minutum Austin, Bull. Torrey Bot. Club 5 (3): 17, 1874.

** Bazzania minutidens (Steph.) Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 142, 1965. Bas.: Mastigobryum minutidens Steph., Sp. Hepat. (Stephani) 6: 474, 1924.

** Bazzania minutiserra (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 26 (2): 77, 1977. Bas.: Mastigobryum minutiserrum Steph., Sp. Hepat. (Stephani) 6: 474, 1924.

** Bazzania missionum (Herzog) Jovet-Ast, Rev. Bryol. Lichénol. 20 (1/2): 96, 1951. Bas.: Mastigobryum missionum Herzog, Ann. Bryol. 5: 93, 1932.

** Bazzania mittenii (Steph.) Steph., Hedwigia 28 (2): 132, 1889. Bas.: Mastigobryum mittenii Steph., Hedwigia 25 (6): 245, 1886.

*** Bazzania monilinervis (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Jungermannia monilinervis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 56, 1832.

** Bazzania morokensis (Steph.) Grolle, J. Hattori Bot. Lab. 31: 1, 1968. Bas.: Mastigobryum morokense Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 765 (449), 1908.

*** Bazzania nitida (F.Weber) Grolle, Rev. Bryol. Lichénol. 29 (3/4): 210, 1960 [1961]. Bas.: Jungermannia nitida F.Weber, Hist. Musc. Hepat. Prodr.: 43, 1815.

*** Bazzania novae-zelandiae (Mitt.) Besch. et C.Massal., Miss. sci. Cape Horn, Bot. 5: 233, 1889. Bas.: Mastigobryum novae-zelandiae Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 148, 1854.

** Bazzania nudicaulis A.Evans, Bryologist 26 (6): 62, 1923 [1924].

** Bazzania nuuanuensis C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 15, 1904.

** Bazzania obcuneata (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981. Bas.: Mastigobryum obcuneatum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 863 (487), 1908.

** Bazzania obtusata (Mitt.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959. Bas.: Mastigobryum obtusatum Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 106, 1860 [1861].

** Bazzania okaritana Meagher et Glenny, J. Bryol. 29 (1): 60, 2007.

** Bazzania orbanii Pócs, Acta Biol. Pl. Agr. 1: 16, 2010 [2011].

** Bazzania ovistipula (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959. Bas.: Mastigobryum ovistipulum Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 760 (444), 1908.

** Bazzania pallidevirens (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 42, 1946. Bas.: Mastigobryum pallidevirens Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 849 (473), 1908.

** Bazzania papillosa S.W.Arnell, Svensk Bot. Tidskr. 59 (1): 67, 1965.

** Bazzania paradoxa (Sande Lac.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 307, 1896. Bas.: Mastigobryum paradoxum Sande Lac., Ned. Kruidk. Arch. 3: 419, 1854 [1855].

*** Bazzania parisii (Steph.) N.Kitag., J. Hattori Bot. Lab. 47: 135, 1980. Bas.: Mastigobryum parisii Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 769 (453), 1908.

** Bazzania parvitexta Steph., Hedwigia 32 (4): 211, 1893.

** Bazzania patens (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Herpetium patens Mont., Ann. Sci. Nat. Bot. (sér. 2) 20: 295, 1844.

** Bazzania patentistipa (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 168, 1898. Bas.: Mastigobryum patentistipum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 302, 1864.

** Bazzania paucidens (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981. Bas.: Mastigobryum paucidens Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 860 (484), 1908.

** Bazzania pearsonii Steph., Hedwigia 32 (4): 212, 1893.

*** Bazzania pectinata (Lindenb. et Gottsche) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 259, 1893. Bas.: Mastigobryum pectinatum Lindenb. et Gottsche, Sp. Hepat. (Lindenberg) 8-11: 84, 1851.

** Bazzania perfalcata N.Kitag., J. Hattori Bot. Lab. 47: 135, 1980.

** Bazzania perrotana E.W.Jones, J. Bryol. 8 (3): 310, 1975. Nom. nov. pro Mastigobryum perrotanum Steph., Sp. Hepat. (Stephani) 6: 476, 1924, nom. illeg.

** Bazzania peruviana (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum peruvianum Nees, Syn. Hepat. 2: 220, 1845.

*** Bazzania phyllobola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 372, 1885.

*** Bazzania placophylla (Taylor) Grolle, Rev. Bryol. Lichénol. 27 (1/2): 54, 1958. Bas.: Jungermannia placophylla Taylor, London J. Bot. 5: 276, 1846.

** Bazzania platycnema (Schwägr. ex Steph.) H.A.Mill., Bryologist 63 (2): 121, 1960. Bas.: Mastigobryum platycnemum Schwägr. ex Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 776 (460), 1908.

** Bazzania pompeana (Sande Lac.) Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 200, 1891. Bas.: Mastigobryum pompeanum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 304, 1864.

*** Bazzania praerupta (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Jungermannia praerupta Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 229, 1824 [1825].

* Bazzania praerupta var. obliquata (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 170, 1898. Bas.: Jungermannia obliquata Nees, Enum. Pl. Crypt. Javae: 62, 1830.

** Bazzania praerupta var. thermalis Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 396, 1955.

** Bazzania pseudovittata N.Kitag. et T.Kodama, J. Hattori Bot. Lab. 39: 69, 1975.

** Bazzania pusilla (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 307, 1896. Bas.: Mastigobryum pusillum Mitt., Fl. vit.: 406, 1871 [1873].

** Bazzania pycnophylla (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum pycnophyllum Taylor, London J. Bot. 5: 371, 1846.

** Bazzania quadratistipula H.A.Mill., Phytologia 47 (4): 321, 1981. Nom. nov. pro Mastigobryum quadratum Steph., Sp. Hepat. (Stephani) 6: 477, 1924, nom. illeg.

** Bazzania rabenhorstii (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959. Bas.: Mastigobryum rabenhorstii Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 774 (458), 1908.

** Bazzania recurva (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Herpetium recurvum Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 253, 1843.

* Bazzania recurva var. major (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 172, 1898. Bas.: Mastigobryum recurvum var. majus Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 304, 1864.

** Bazzania reflexa (Gottsche) Steph., Rev. Bryol. 18 (4): 56, 1891. Bas.: Mastigobryum reflexum Gottsche, Abh. Naturwiss. Vereins Bremen 7: 347, 1882.

** Bazzania reinwardtii (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 172, 1898. Bas.: Mastigobryum reinwardtii Sande Lac., Pl. Ind. Batav. Orient.: 22, 1856.

** Bazzania renistipula Steph., Hedwigia 32 (4): 212, 1893.

** Bazzania revoluta (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 450, 1972 [1973]. Bas.: Mastigobryum revolutum Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 961 (511), 1908.

*** Bazzania rimosa Meagher, Nova Hedwigia 86 (3/4): 489, 2008.

** Bazzania roccatii Gola, Ann. Bot. (Rome) 6 (2): 273, 1907.

*** Bazzania roraimensis (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 27, 1946. Bas.: Mastigobryum roraimense Steph., Trans. Linn. Soc. London, Bot. 6 (1): 97, 1901.

** Bazzania sandvicensis (Gottsche ex Steph.) Steph., Bull. Herb. Boissier 5 (10): 841, 1897. Bas.: Mastigobryum sandvicense Gottsche ex Steph., Hedwigia 25 (5): 207, 1886.

** Bazzania sauropoda Meagher, Austrobaileya 7 (1): 129, 2005.

*** Bazzania scalaris Meagher, Telopea 11 (3): 247, 2006.

*** Bazzania schlimiana (Gottsche) Fulford, Bull. Torrey Bot. Club 86 (6): 401, 1959. Bas.: Mastigobryum schlimianum Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 140, 1864.

** Bazzania schultze-motelii N.Kitag., J. Hattori Bot. Lab. 47: 138, 1980.

** Bazzania schusterana N.Kitag., J. Hattori Bot. Lab. 47: 139, 1980.

*** Bazzania schwaneckiana (Hampe et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum schwaneckianum Hampe et Gottsche, Linnaea 25 (3): 345, 1852 [1853].

* Bazzania scutigera (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Herpetium scutigerum Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 44, 1838.

** Bazzania semicordata (Lindenb. et Gottsche) Kuntze, Revis. Gen. Pl. 2: 832, 1891. Bas.: Mastigobryum semicordatum Lindenb. et Gottsche, Syn. Hepat. 5: 720, 1847.

* Bazzania serpentina (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Jungermannia serpentina Nees, Enum. Pl. Crypt. Javae: 62, 1830. 113

** Bazzania serrapiculata Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 141, 1965.

** Bazzania serrata Fulford, Bull. Torrey Bot. Club 86 (5): 321, 1959.

** Bazzania serrulatoides Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 200, 1934.

* Bazzania sikkimensis (Steph.) Herzog, Ann. Bryol. 12: 78, 1939. Bas.: Mastigobryum sikkimense Steph., Hedwigia 44 (2): 73, 1905. 114

** Bazzania spinosa S.Okamura, J. Coll. Sci. Imp. Univ. Tokyo 38 (4): 2, 1916.

** Bazzania spiralis (Reinw., Blume et Nees) Meijer, Blumea 10 (2): 381, 1960. Bas.: Jungermannia spiralis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 231, 1824 [1825].

** Bazzania spruceana Steph., Hedwigia 32 (4): 213, 1893.

** Bazzania squarrosa (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981. Bas.: Mastigobryum squarrosum Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 776 (460), 1908.

*** Bazzania stolonifera (Sw.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Jungermannia stolonifera Sw., Prodr. (Swartz): 144, 1788.

* Bazzania stolonifera var. granatensis (Gottsche) Fulford, Ann. Cryptog. Phytopathol. 3: 51, 1946. Bas.: Mastigobryum stoloniferum var. granatense Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 141, 1864.

** Bazzania stresemannii (Herzog) N.Kitag., Bull. Nara Univ. Educ., B 28 (2): 77, 1979. Bas.: Mastigobryum stresemannii Herzog, Beih. Bot. Centralbl. 38 (2): 324, 1921.

** Bazzania subacuta (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 307, 1896. Bas.: Mastigobryum subacutum Mitt., Fl. vit.: 406, 1871 [1873].

** Bazzania subaequitexta (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 28 (2): 81, 1979. Bas.: Mastigobryum subaequitextum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 859 (483), 1908.

** Bazzania subintegra (Steph.) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 69, 2015. Bas.: Mastigobryum subintegrum Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 775 (459), 1908.

** Bazzania sublonga Fulford, Bull. Torrey Bot. Club 86 (5): 334, 1959.

** Bazzania subserrifolia (Beauverd) H.A.Mill., Phytologia 47 (4): 321, 1981. Bas.: Mastigobryum subserrifolium Beauverd, Sp. Hepat. (Stephani) 6: 480, 1924.

** Bazzania subserrulata A.Evans, Pap. Michigan Acad. Sci. 17: 97, 1932 [1933].

*** Bazzania subtilis (Sande Lac.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum subtile Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 302, 1864.

** Bazzania succulenta N.Kitag., J. Hattori Bot. Lab. 47: 141, 1980.

** Bazzania sumatrana (Sande Lac. ex Steph.) Steph., Hedwigia 32 (4): 209, 1893. Bas.: Mastigobryum sumatranum Sande Lac. ex Steph., Hedwigia 25 (6): 234, 1886.

* Bazzania sumbavensis (Gottsche ex Steph.) Steph., Hedwigia 32 (4): 204, 1893. Bas.: Mastigobryum sumbavense Gottsche ex Steph., Hedwigia 25 (6): 236, 1886. 115

*** Bazzania taleana (Gottsche) Fulford, Ann. Cryptog. Phytopathol. 3: 54, 1946. Bas.: Mastigobryum taleanum Gottsche, Mexik. Leverm.: 131, 1863.

*** Bazzania tayloriana (Mitt.) Kuntze, Revis. Gen. Pl. 2: 832, 1891. Bas.: Mastigobryum taylorianum Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 147, 1854.

** Bazzania temariana (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981. Bas.: Mastigobryum temarianum Steph., Sp. Hepat. (Stephani) 3: 532, 1909.

*** Bazzania tessellata Meagher, Nova Hedwigia 92 (3/4): 492, 2011.

** Bazzania tiaoloensis Mizut. et K.C.Chang, J. Hattori Bot. Lab. 60: 432, 1986.

*** Bazzania tricrenata (Wahlenb.) Lindb., Musci Fenn. Exsic., fasc. 2: [2 (adnot.)], 1872. Bas.: Jungermannia tricrenata Wahlenb., Fl. Carpat. Princ.: 364, 1814.

** Bazzania tricrenata var. fulfordiae W.S.Hong, Bryologist 91 (4): 331, 1988.

*** Bazzania tridens (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Jungermannia tridens Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 228, 1824 [1825].

*** Bazzania tridens var. assamica (Steph.) Pócs, J. Hattori Bot. Lab. 32: 86, 1969. Bas.: Mastigobryum assamicum Steph., Hedwigia 24 (5): 216, 1885.

** Bazzania tridens var. cornutistipula (Steph.) Pócs, J. Hattori Bot. Lab. 32: 83, 1969. Bas.: Mastigobryum cornutistipulum Steph., Rev. Bryol. 35 (2): 35, 1908.

*** Bazzania trilobata (L.) Gray, Nat. Arr. Brit. Pl. 1: 704, 1821. Bas.: Jungermannia trilobata L., Sp. Pl. 1: 1133, 1753.

** Bazzania trilobata var. depauperata (Müll.Frib.) Grolle, Lindbergia 1 (3/4): 197, 1972 [1973]. Bas.: Pleuroschisma trilobatum var. depauperatum Müll.Frib., Lebermoose 2 (18): 266, 1913.

*** Bazzania uncigera (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Jungermannia uncigera Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 230, 1824 [1825].

** Bazzania uncigera var. brevifolia Herzog, Trans. Brit. Bryol. Soc. 1 (4): 301, 1950.

** Bazzania uncigera var. gibba (Sande Lac.) Meijer, Blumea 10 (2): 378, 1960. Bas.: Mastigobryum gibbum Sande Lac., Plagiochila Sandei: 8, 1856.

** Bazzania undulata Herzog, Trans. Brit. Bryol. Soc. 1 (4): 306, 1950.

** Bazzania vietnamica Pócs, J. Hattori Bot. Lab. 32: 90, 1969.

** Bazzania vitiana Mitt., Hedwigia 32 (4): 214, 1893.

*** Bazzania vittata (Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum vittatum Gottsche, Syn. Hepat. 2: 216, 1845.

** Bazzania vittata var. luxurians (De Not.) Schiffn., Consp. Hepat. Arch. Ind.: 178, 1898. Bas.: Mastigobryum vittatum var. luxurians De Not., Epat. Borneo: 29, 1874.

** Bazzania wallichiana (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877. Bas.: Mastigobryum wallichianum Lindenb., Syn. Hepat. 2: 229, 1845. 116

** Bazzania watanabei Inoue, J. Jap. Bot. 34 (9): 269, 1959.

* Bazzania wattsiana (Steph.) Meagher, Australas. Bryol. Newslett. 50: 8, 2005. Bas.: Mastigobryum wattsianum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 850 (474), 1908. 117

** Bazzania wiltensii (Sande Lac. ex Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 180, 1898. Bas.: Mastigobryum wiltensii Sande Lac. ex Steph., Hedwigia 25 (6): 237, 1886.

** Bazzania wooroonooran Meagher, Nova Hedwigia 100 (3/4): 549, 2015.

** Bazzania wrightii (Gottsche ex Steph.) Steph., Hedwigia 27 (11/12): 279, 1888. Bas.: Mastigobryum wrightii Gottsche ex Steph., Hedwigia 25 (6): 237, 1886.

** Bazzania yoshinagana (Steph.) Yasuda, Shokubutsugaku Kakuron: 711, 1911. Bas.: Mastigobryum yoshinaganum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 866 (490), 1908.

** Bazzania zollingeri (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877. Bas.: Mastigobryum zollingeri Lindenb., Bot. Zeitung (Berlin) 6 (25): 462, 1848.

*** Bazzania zonulata Meagher, Nova Hedwigia 86 (3/4): 491, 2008.

* Mastigopelma Mitt., Fl. vit.: 406, 1871 [1873]. 118

** Mastigopelma fragile (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 454, 1972 [1973]. Bas.: Mastigobryum fragile Steph., Sp. Hepat. (Stephani) 6: 463, 1924.

** Mastigopelma pulvinulatum (De Not.) Grolle, J. Hattori Bot. Lab. 33: 39, 1970. Bas.: Mastigobryum pulvinulatum De Not., Epat. Borneo: 40, 1874.

** Mastigopelma simplex Mitt., Fl. vit.: 406, 1871 [1873].

** Mastigopelma subfissum Grolle, J. Hattori Bot. Lab. 33: 36, 1970.

*** Drucelloideae R.M.Schust.

*** Drucella E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (3): 45, 1962.

*** Drucella integristipula (Steph.) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (3): 45, 1962. Bas.: Lepidozia integristipula Steph., Sp. Hepat. (Stephani) 6: 331, 1922.

*** Lembidioideae R.M.Schust.

** Dendrolembidium Herzog, Ark. Bot. (n.ser.) 1 (13): 497, 1951.

*** Dendrolembidium dendroides (Carrington et Pearson) Herzog, Ark. Bot. (n.ser.) 1 (13): 500, 1951. Bas.: Lembidium dendroides Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1047, 1888.

*** Dendrolembidium tenax (Grev.) Herzog, Ark. Bot. (n.ser.) 1 (13): 499, 1951. Bas.: Jungermannia tenax Grev., Ann. Lyceum Nat. Hist. New York 1 (2): 277, 1825.

** Hygrolembidium R.M.Schust., J. Hattori Bot. Lab. 26: 277, 1963.

*** Hygrolembidium acrocladum (Berggr.) R.M.Schust., J. Hattori Bot. Lab. 26: 277, 1963. Bas.: Aplozia acroclada Berggr., New Zealand Hepat.: 9, 1898.

** Hygrolembidium andinum (Herzog) R.M.Schust., Nova Hedwigia 10 (1/2): 23, 1965. Bas.: Lembidium andinum Herzog, Rev. Bryol. Lichénol. 23 (1/2): 48, 1954.

*** Hygrolembidium australe (Steph.) Grolle, J. Jap. Bot. 41 (8): 229, 1966. Bas.: Hygrobiella australis Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 574 (358), 1908.

*** Hygrolembidium boschianum (Sande Lac.) R.M.Schust., J. Hattori Bot. Lab. 26: 277, 1963. Bas.: Jungermannia boschiana Sande Lac., Ned. Kruidk. Arch. 3: 521, 1855.

** Hygrolembidium isophyllum R.M.Schust., Nova Hedwigia 15: 467, 1968.

*** Hygrolembidium rigidum R.M.Schust. et J.J.Engel, Phytologia 62 (1): 9, 1987.

*** Hygrolembidium triquetrum J.J.Engel et R.M.Schust., Phytologia 62 (1): 11, 1987.

** Hygrolembidium ventrosum (Mitt.) Grolle, Marion Prince Edw. Is: 233, 1971. Bas.: Lembidium ventrosum Mitt., J. Linn. Soc., Bot. 15 (82): 69, 1876.

** Isolembidium R.M.Schust., Nova Hedwigia 15: 466, 1968.

*** Isolembidium anomalum (Rodway) Grolle, J. Bryol. 10 (3): 264, 1979. Bas.: Lembidium anomalum Rodway, Tasm. Bryoph.: 70, 1917.

** Isolembidium anomalum var. cucullatum (E.A.Hodgs.) J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 63: 268, 1987. Bas.: Lembidium cucullatum E.A.Hodgs., Rec. Domin. Mus. 4 (11): 110, 1962.

*** Kurzia G.Martens, Flora 53 (27): 417, 1870 (0).

** subg. Kurzia

*** Kurzia capillaris (Sw.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 173, 1963 [1964]. Bas.: Jungermannia capillaris Sw., Prodr. (Swartz): 144, 1788.

*** Kurzia capillaris subsp. capillaris var. verrucosa (Steph.) Pócs, Proc. Third Meeting Bryol. C. & E. Europe: 111, 1984. Bas.: Lepidozia verrucosa Steph., Hedwigia 24 (4): 167, 1885.

*** Kurzia capillaris subsp. paramicola Pócs, Acta Biol. Pl. Agr. 2: 102, 2012.

*** Kurzia capillaris subsp. stephanii (Renauld ex Steph.) Pócs, Proc. Third Meeting Bryol. C. & E. Europe: 111, 1984. Bas.: Lepidozia stephanii Renauld ex Steph., Bot. Gaz. 15 (11): 287, 1890.

*** Kurzia gonyotricha (Sande Lac.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 167, 1963 [1964]. Bas.: Lepidozia gonyotricha Sande Lac., Ned. Kruidk. Arch. 3: 521, 1855.

** Kurzia nemoides (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 173, 1963 [1964]. Bas.: Jungermannia nemoides Hook.f. et Taylor, London J. Bot. 4: 84, 1845.

** subg. Micrisophylla (Fulford) J.J.Engel ex R.M.Schust., J. Hattori Bot. Lab. 48: 347, 1980. Bas.: Micrisophylla Fulford, Brittonia 14 (1): 124, 1962.

** Kurzia saddlensis (Besch. et C.Massal.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 174, 1963 [1964]. Bas.: Lepidozia saddlensis Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 637, 1886.

** subg. Microlepidozia (Spruce) R.M.Schust., J. Hattori Bot. Lab. 48: 355, 1980. Bas.: Lepidozia subg. Microlepidozia Spruce, J. Bot. 14: 165, 1876.

** Kurzia calcarata (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964]. Bas.: Lepidozia calcarata Steph., Sp. Hepat. (Stephani) 3: 592, 1909.

** Kurzia compacta (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964]. Bas.: Lepidozia compacta Steph., Sp. Hepat. (Stephani) 3: 592, 1909.

** Kurzia helophila R.M.Schust., J. Hattori Bot. Lab. 48: 368, 1980.

** Kurzia helophila var. flaccida R.M.Schust. ex J.J.Engel, Novon 17 (3): 310, 2007.

*** Kurzia hippurioides (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964]. Bas.: Jungermannia hippurioides Hook.f. et Taylor, London J. Bot. 3: 287 [387], 1844.

** Kurzia hippurioides var. ornata J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 80: 228, 1996.

** Kurzia irregularis (Steph.) Grolle, J. Hattori Bot. Lab. 36: 548, 1972 [1973]. Bas.: Lepidozia irregularis Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 120, 1911.

** Kurzia moniliformis J.J.Engel, Cryptog. Bryol. 26 (1): 73, 2005.

*** Kurzia pauciflora (Dicks.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964]. Bas.: Jungermannia pauciflora Dicks., Fasc. Pl. Crypt. Brit. 2: 15, 1790.

Incertae sedis

** Kurzia abbreviata Mizut., J. Hattori Bot. Lab. 38: 379, 1974.

** Kurzia abietinella (Herzog) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964]. Bas.: Lepidozia abietinella Herzog, Trans. Brit. Bryol. Soc. 1 (4): 311, 1950.

*** Kurzia bisetula (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964]. Bas.: Lepidozia bisetula Steph., Sp. Hepat. (Stephani) 6: 323, 1922.

** Kurzia borneensis Mizut., J. Hattori Bot. Lab. 38: 377, 1974.

** Kurzia brasiliensis (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 174, 1963 [1964]. Bas.: Psiloclada brasiliensis Steph., Sp. Hepat. (Stephani) 3: 550, 1909.

** Kurzia brevicalycina (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 175, 1963 [1964]. Bas.: Lepidozia brevicalycina Steph., Sp. Hepat. (Stephani) 3: 580, 1909.

* Kurzia caduciloba R.M.Schust., Beih. Nova Hedwigia 118: 273, 2000.

* Kurzia cucullifolia (Steph.) R.M.Schust., J. Hattori Bot. Lab. 48: 350, 1980. Bas.: Lepidozia cucullifolia Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 51, 1900. 119

*** Kurzia flagellifera (Steph.) Grolle, J. Jap. Bot. 39 (3): 80, 1964. Bas.: Lepidozia flagellifera Steph., Sp. Hepat. (Stephani) 3: 571, 1909.

*** Kurzia fragilifolia R.M.Schust., J. Hattori Bot. Lab. 48: 364, 1980.

** Kurzia fragillima (Herzog) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 174, 1963 [1964]. Bas.: Lepidozia fragillima Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 726, 1942, nom. illeg.

** Kurzia geniculata Mizut., J. Hattori Bot. Lab. 38: 383, 1974.

** Kurzia hawaica (C.M.Cooke) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964]. Bas.: Lepidozia hawaica C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 8, 1904.

** Kurzia hispida (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964]. Bas.: Lepidozia hispida Steph., Sp. Hepat. (Stephani) 3: 607, 1909.

** Kurzia lateconica (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 175, 1963 [1964]. Bas.: Lepidozia lateconica Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 113, 1914.

** Kurzia lineariloba Mizut., J. Hattori Bot. Lab. 38: 382, 1974.

*** Kurzia longicaulis Piippo, Acta Bot. Fenn. 131: 174, 1985.

** Kurzia makinoana (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964]. Bas.: Lepidozia makinoana Steph., Bull. Herb. Boissier 5 (2): 94, 1897.

** Kurzia mauiensis (H.A.Mill.) H.A.Mill., J. Hattori Bot. Lab. 30: 274, 1967. Bas.: Microlepidozia mauiensis H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 496, 1963.

** Kurzia mollis (Steph.) J.J.Engel et R.M.Schust., Bryologist 79 (4): 514, 1976 [1977]. Bas.: Lepidozia mollis Steph., Sp. Hepat. (Stephani) 3: 601, 1909.

*** Kurzia nivicola (R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 52, 2013. Bas.: Telaranea nivicola R.M.Schust., Nova Hedwigia 15: 460, 1968.

*** Kurzia pallescens Grolle, Rev. Bryol. Lichénol. 32 (3/4): 177, 1963 [1964].

** Kurzia pallida Piippo, Acta Bot. Fenn. 131: 178, 1985.

*** Kurzia quinquespina J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 80: 217, 1996.

** Kurzia reversa (Carrington et Pearson) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 175, 1963 [1964]. Bas.: Lepidozia reversa Carrington et Pearson, J. Bot. 27: 225, 1889.

** Kurzia setiformis (De Not.) J.J.Engel et R.M.Schust., Bryologist 79 (4): 514, 1976 [1977]. Bas.: Lepidozia setiformis De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 225, 1857.

** Kurzia sexfida (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964]. Bas.: Lepidozia sexfida Steph., Sp. Hepat. (Stephani) 3: 582, 1909.

** Kurzia sinensis K.C.Chang, Bull. Bot. Res., Harbin 4 (3): 83, 1984.

** Kurzia sylvatica (A.Evans) Grolle, Herzogia 3: 77, 1973. Bas.: Lepidozia sylvatica A.Evans, Rhodora 6 (69): 186, 1904.

*** Kurzia tasmanica (Steph.) E.D.Cooper, Phytotaxa 97 (2): 52, 2013. Bas.: Lepidozia tasmanica Steph., Sp. Hepat. (Stephani) 3: 580, 1909.

** Kurzia tayloriana (H.A.Mill.) H.A.Mill., J. Hattori Bot. Lab. 30: 274, 1967. Bas.: Microlepidozia tayloriana H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 497, 1963.

** Kurzia tenerrima (Mitt.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964]. Bas.: Lepidozia tenerrima Mitt., Sp. Hepat. (Stephani) 3: 607, 1909.

** Kurzia touwii N.Kitag., Acta Phytotax. Geobot. 29 (1/5): 56, 1978.

** Kurzia trichoclados (Müll.Frib.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964]. Bas.: Lepidozia trichoclados Müll.Frib., Hedwigia 38 (4): 197, 1899.

*** Kurzia trilobata (R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 118: 270, 2000. Bas.: Kurzia quadriseta var. trilobata R.M.Schust., J. Hattori Bot. Lab. 48: 363, 1980.

** Kurzia verticellata (Carrington) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964]. Bas.: Lepidozia verticellata Carrington, Pap. & Proc. Roy. Soc. Tasmania 1887: 3, 1888.

** Lembidium Mitt., Handb. N. Zeal. fl. 2: 754, 1867 nom. conserv.

*** Lembidium berggrenii Herzog, Ark. Bot. (n.ser.) 1 (13): 485, 1951.

*** Lembidium longifolium R.M.Schust., Phytologia 45 (5): 420, 1980.

*** Lembidium nutans (Hook.f. et Taylor) Mitt., Handb. N. Zeal. Fl. 2: 754, 1867. Bas.: Jungermannia nutans Hook.f. et Taylor, London J. Bot. 3: 289 [389], 1844.

** Lembidium nutans var. flagelliferum E.A.Hodgs., Rec. Domin. Mus. 4 (11): 110, 1962.

** Megalembidium R.M.Schust., J. Hattori Bot. Lab. 26: 258, 1963.

*** Megalembidium insulanum (W.Martin et E.A.Hodgs.) R.M.Schust., J. Hattori Bot. Lab. 26: 258, 1963. Bas.: Lembidium insulanum W.Martin et E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 78 (4): 497, 1950.

** Pseudocephalozia R.M.Schust., Nova Hedwigia 10 (1/2): 21, 1965.

** sect. Lobulatae R.M.Schust., J. Hattori Bot. Lab. 36: 371, 1972.

** Pseudocephalozia cucullata J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 38: 694, 1974.

*** Pseudocephalozia lobulata (Herzog) R.M.Schust., J. Hattori Bot. Lab. 36: 371, 1972 [1973]. Bas.: Lembidium lobulatum Herzog, Arch. Esc. Fárm. Fac. Ci. Méd. Córdoba 7: 24, 1938.

*** Pseudocephalozia quadriloba (Steph.) R.M.Schust., J. Hattori Bot. Lab. 36: 371, 1972 [1973]. Bas.: Isotachis quadriloba Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 54, 1900.

** sect. Pseudocephalozia

** Pseudocephalozia lepidozioides R.M.Schust., Nova Hedwigia 10 (1/2): 22, 1965.

** Pseudocephalozia leptodictyon R.M.Schust., J. Hattori Bot. Lab. 36: 369, 1972 [1973].

*** Pseudocephalozia paludicola R.M.Schust., Nova Hedwigia 10 (1/2): 21, 1965.

*** Lepidozioideae Müll.Frib.

*** Ceramanus E.D.Cooper, Phytotaxa 97 (2): 53, 2013.

*** Ceramanus centipes (Lindenb. et Gottsche) E.D.Cooper, Phytotaxa 97 (2): 54, 2013. Bas.: Lepidozia centipes Lindenb. et Gottsche, Syn. Hepat. 2: 204, 1845.

*** Ceramanus clatritexta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013. Bas.: Lepidozia clatritexta Steph., Sp. Hepat. (Stephani) 3: 583, 1909.

*** Ceramanus elegans (Colenso) E.D.Cooper, Phytotaxa 97 (2): 54, 2013. Bas.: Lepidozia elegans Colenso, Trans. & Proc. New Zealand Inst. 21: 65, 1889.

*** Ceramanus grossiseta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013. Bas.: Lepidozia grossiseta Steph., Sp. Hepat. (Stephani) 3: 584, 1909.

*** Ceramanus perfragilis (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013. Bas.: Telaranea perfragilis J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 72, 2004.

*** Ceramanus pruinosa (Herzog) E.D.Cooper, Phytotaxa 97 (2): 54, 2013. Bas.: Lepidozia pruinosa Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 93, 1952.

*** Ceramanus tuberifera (J.J.Engel et R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013. Bas.: Telaranea tuberifera J.J.Engel et R.M.Schust., Fieldiana, Bot. (n.ser.) 14: 2, 1983.

*** Lepidozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 19, 1835 nom. conserv. Bas.: Pleuroschisma sect. Lepidozia Dumort., Syll. Jungerm. Europ.: 69, 1831. 120

** Lepidozia acantha J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 71, 2001.

** Lepidozia aequiloba Steph., Sp. Hepat. (Stephani) 6: 319, 1922.

** Lepidozia africana Steph., Sp. Hepat. (Stephani) 6: 320, 1922.

*** Lepidozia alstonii Fulford, Mem. New York Bot. Gard. 11 (2): 211, 1966.

** Lepidozia ambigua De Not., Epat. Borneo: 25, 1874.

** Lepidozia andicola Beauverd, Sp. Hepat. (Stephani) 6: 572, 1924. Nom. nov. pro Lepidozia appendiculata Steph., Biblioth. Bot. 87 (2): 225, 1916, nom. illeg.

** Lepidozia appressifolia Steph., Sp. Hepat. (Stephani) 3: 583, 1909.

*** Lepidozia armata Steph., Sp. Hepat. (Stephani) 3: 567, 1909.

** Lepidozia asymmetrica Steph., Sp. Hepat. (Stephani) 3: 586, 1909.

** Lepidozia auriculata Mitt., Sp. Hepat. (Stephani) 3: 579, 1909.

** Lepidozia australis (Lehm. et Lindenb.) Mitt., Fl. vit.: 406, 1871 [1873]. Bas.: Jungermannia australis Lehm. et Lindenb., Nov. Stirp. Pug. 6: 28, 1834.

** Lepidozia bidens J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 79, 2001.

** Lepidozia biloba Herzog, Ann. Bryol. 4: 83, 1931.

*** Lepidozia bisbifida Steph., Sp. Hepat. (Stephani) 3: 593, 1909.

** Lepidozia borneensis Steph., Sp. Hepat. (Stephani) 3: 625, 1909.

** Lepidozia bragginsiana E.D.Cooper et M.A.M.Renner, Phytotaxa 173 (2): 118, 2014.

* Lepidozia brasiliensis Steph., Sp. Hepat. (Stephani) 3: 571, 1909. 121

** Lepidozia brevidentata Mitt., Fl. vit.: 406, 1871 [1873].

** Lepidozia brevifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 104, 1860 [1861].

** Lepidozia brevifolia var. planifolia Schiffn., Ann. Bryol. 8: 155, 1935.

** Lepidozia brotheri Steph., Sp. Hepat. (Stephani) 3: 623, 1909.

** Lepidozia buffalona Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 111, 1914.

** Lepidozia bursifera S.Hatt. et Grolle, J. Hattori Bot. Lab. 30: 115, 1967.

** Lepidozia caespitosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 362, 1885.

** Lepidozia caledonica Steph., Rev. Bryol. 35 (2): 31, 1908.

** Lepidozia caledonica var. tenuisecta Hürl., Bauhinia 8 (2): 109, 1985.

* Lepidozia ceramensis Herzog, Hedwigia 66 (6): 340, 1926.

** Lepidozia cherydrion Hürl., Bauhinia 8 (2): 109, 1985.

** Lepidozia chiloensis Steph., Sp. Hepat. (Stephani) 6: 322, 1922.

** Lepidozia chordulifera Taylor, London J. Bot. 5: 371, 1846.

*** Lepidozia cladorhiza (Reinw., Blume et Nees) Nees, Syn. Hepat. 2: 210, 1845. Bas.: Jungermannia cladorhiza Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 203, 1824 [1825].

*** Lepidozia coilophylla Taylor, London J. Bot. 5: 370, 1846.

** Lepidozia coilophylla var. apiculiloba (Steph.) Fulford, Mem. New York Bot. Gard. 11 (2): 194, 1966. Bas.: Lepidozia apiculiloba Steph., Sp. Hepat. (Stephani) 6: 321, 1922.

** Lepidozia communis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 111, 1914.

*** Lepidozia concinna Colenso, Trans. & Proc. New Zealand Inst. 18: 244, 1886.

** Lepidozia cordata Lindenb., Syn. Hepat. 2: 207, 1845.

* Lepidozia cordistipula Steph., Sp. Hepat. (Stephani) 6: 345, 1922.

** Lepidozia crassitexta Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 111, 1914.

*** Lepidozia cupressina (Sw.) Lindenb., Syn. Hepat. 2: 207, 1845. Bas.: Jungermannia cupressina Sw., Prodr. (Swartz): 144, 1788.

** Lepidozia cupressina subsp. natalensis (Steph.) Pócs, Proc. Third Meeting Bryol. C. & E. Europe: 109, 1984. Bas.: Lepidozia natalensis Steph., Sp. Hepat. (Stephani) 3: 562, 1909.

* Lepidozia cupressina subsp. pinnata (Hook.) Pócs, Proc. Third Meeting Bryol. C. & E. Europe: 109, 1984. Bas.: Jungermannia reptans var. pinnata Hook., Brit. Jungermann.: tab. 75, 1815.

* Lepidozia cupressina subsp. quinquefida (Steph.) Pócs, Proc. Third Meeting Bryol. C. & E. Europe: 109, 1984. Bas.: Lepidozia quinquefida Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 123, 1911.

** Lepidozia decaisnei Steph., Sp. Hepat. (Stephani) 3: 588, 1909.

** Lepidozia dendritica Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 362, 1885.

** Lepidozia densa Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 26, 1925.

*** Lepidozia digitata Herzog, Trans. & Proc. Roy. Soc. New Zealand 68 (1): 45, 1938.

*** Lepidozia eenii S.W.Arnell, Svensk Bot. Tidskr. 57 (2): 190, 1963.

** Lepidozia elobata R.M.Schust., Fieldiana, Bot. (n.ser.) 42: 74, 2001.

** Lepidozia erosa Steph., Sp. Hepat. (Stephani) 3: 621, 1909.

** Lepidozia erronea Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 725, 1942. Nom. nov. pro Lepidozia fernandeziensis Steph., Sp. Hepat. (Stephani) 6: 326, 1922, nom. illeg.

** Lepidozia everettii Steph., Sp. Hepat. (Stephani) 3: 622, 1909.

* Lepidozia everettii var. javensis Herzog, Ann. Bryol. 5: 78, 1932.

** Lepidozia fauriana Steph., Sp. Hepat. (Stephani) 3: 631, 1909.

*** Lepidozia ferdinandi-muelleri Steph., Sp. Hepat. (Stephani) 3: 614, 1909.

** Lepidozia filamentosa (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 2: 206, 1845. Bas.: Jungermannia filamentosa Lehm. et Lindenb., Nov. Stirp. Pug. 6: 29, 1834.

** Lepidozia fistulosa Mitt., Fl. vit.: 406, 1871 [1873].

** Lepidozia flexuosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 103, 1860 [1861].

** Lepidozia fuegiensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 63, 1911.

** Lepidozia fugax J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 63, 2001.

* Lepidozia gedena Steph., Sp. Hepat. (Stephani) 6: 327, 1922.

*** Lepidozia glaucescens J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 101, 2001.

*** Lepidozia glaucophylla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 207, 1845. Bas.: Jungermannia glaucophylla Hook.f. et Taylor, London J. Bot. 3: 580, 1844.

** Lepidozia grandifolia Steph., Sp. Hepat. (Stephani) 3: 625, 1909.

* Lepidozia griseola Herzog, Hedwigia 66 (6): 340, 1926.

** Lepidozia groenlandica Lehm., Nov. Stirp. Pug. 10: 7, 1857.

** Lepidozia gwamii Piippo, Ann. Bot. Fenn. 21 (4): 311, 1984.

** Lepidozia hampeana Lindenb., Syn. Hepat. 2: 208, 1845.

*** Lepidozia haskarliana (Gottsche, Lindenb. et Nees) Steph., Sp. Hepat. (Stephani) 3: 614, 1909. Bas.: Lepidozia supradecomposita β haskarliana Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 202, 1845.

** Lepidozia hastatistipula Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 113, 1914.

*** Lepidozia hirta Steph., Sp. Hepat. (Stephani) 3: 599, 1909.

*** Lepidozia holorhiza (Reinw., Blume et Nees) Nees, Syn. Hepat. 2: 210, 1845. Bas.: Jungermannia holorhiza Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 204, 1824 [1825].

** Lepidozia holorhiza var. laxa (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 186, 1898. Bas.: Jungermannia holorhiza β laxa Nees, Enum. Pl. Crypt. Javae: 14, 1830.

** Lepidozia inaequalis (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 2: 209, 1845. Bas.: Jungermannia inaequalis Lehm. et Lindenb., Nov. Stirp. Pug. 5: 1, 1833.

*** Lepidozia incurvata Lindenb., Syn. Hepat. 2: 203, 1845.

** Lepidozia infuscata Mitt., Fl. vit.: 406, 1871 [1873].

** Lepidozia integrifolia Doei, J. Hattori Bot. Lab. 63: 421, 1987.

*** Lepidozia jamaicensis Steph., Sp. Hepat. (Stephani) 3: 568, 1909.

** Lepidozia kashyapii D.Singh et D.K.Singh, Nova Hedwigia 94 (1/2): 222, 2012.

** Lepidozia kinabaluensis Mizut., J. Hattori Bot. Lab. 38: 372, 1974.

*** Lepidozia kirkii Steph., Sp. Hepat. (Stephani) 3: 598, 1909.

** Lepidozia lacerifolia Steph., Sp. Hepat. (Stephani) 6: 332, 1922.

*** Lepidozia laevifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 208, 1845. Bas.: Jungermannia laevifolia Hook.f. et Taylor, London J. Bot. 3: 285 [385], 1844.

** Lepidozia laevifolia var. acutiloba J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 61, 2001.

** Lepidozia laevifolia var. alpina R.M.Schust. et J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 62, 2001.

** Lepidozia lindigiana Steph., Sp. Hepat. (Stephani) 3: 573, 1909.

** Lepidozia loheri Steph., Sp. Hepat. (Stephani) 3: 621, 1909.

** Lepidozia longifolia Steph., Sp. Hepat. (Stephani) 3: 606, 1909.

** Lepidozia loriana Steph., Sp. Hepat. (Stephani) 6: 333, 1922.

*** Lepidozia macrocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 363, 1885.

* Lepidozia massartiana Schiffn. ex Steph., Sp. Hepat. (Stephani) 3: 611, 1909. Based on: Lepidozia massartiana Schiffn., Hedwigia 39 (4): 196, 1900, nom. inval. 122

*** Lepidozia microphylla (Hook.) Lindenb., Syn. Hepat. 2: 202, 1845. Bas.: Jungermannia microphylla Hook., Musci Exot. 1: tab. 80, 1818.

** Lepidozia microstipula Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 114, 1914.

** Lepidozia minima Steph., Sp. Hepat. (Stephani) 6: 335, 1922.

* Lepidozia minor (Gottsche, Lindenb. et Nees) Solari, Boll. Mus. Civico Storia Nat. Verona 10: 203, 1983 [1985]. Bas.: Lepidozia truncatella β minor Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 209, 1845. 123

** Lepidozia miqueliana Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 301, 1864.

** Lepidozia montana Steph., Sp. Hepat. (Stephani) 3: 587, 1909.

* Lepidozia newtonii Steph., Sp. Hepat. (Stephani) 3: 623, 1909.

* Lepidozia nova Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 115, 1914.

*** Lepidozia novae-zelandiae Steph., Sp. Hepat. (Stephani) 3: 595, 1909.

** Lepidozia novae-zelandiae var. heterostipa R.M.Schust., Fieldiana, Bot. (n.ser.) 42: 70, 2001.

** Lepidozia novae-zelandiae var. minima R.M.Schust., Fieldiana, Bot. (n.ser.) 42: 71, 2001.

*** Lepidozia obtusiloba Steph., Sp. Hepat. (Stephani) 3: 598, 1909.

*** Lepidozia obtusiloba var. parvula J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 48, 2001.

** Lepidozia omeiensis P.C.Chen ex Mizut. et K.C.Chang, J. Hattori Bot. Lab. 60: 421, 1986.

*** Lepidozia ornata J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 49, 2001.

* Lepidozia pallida Steph., Sp. Hepat. (Stephani) 3: 604, 1909. 124

* Lepidozia palmicola Steph., Sp. Hepat. (Stephani) 6: 346, 1922.

** Lepidozia paschalis Steph., Sp. Hepat. (Stephani) 6: 336, 1922.

*** Lepidozia patens Lindenb., Syn. Hepat. 2: 202, 1845.

* Lepidozia paucifolia Steph., Sp. Hepat. (Stephani) 3: 610, 1909. 125

** Lepidozia paupercula Steph., Sp. Hepat. (Stephani) 6: 337, 1922.

** Lepidozia pearsonii Spruce, J. Bot. 19: 34, 1881.

*** Lepidozia pendulina (Hook.) Lindenb., Syn. Hepat. 2: 208, 1845. Bas.: Jungermannia pendulina Hook., Musci Exot. 1: tab. 60, 1818.

** Lepidozia peruviensis Steph., Sp. Hepat. (Stephani) 3: 575, 1909.

*** Lepidozia pinnaticruris Spruce ex Steph., Sp. Hepat. (Stephani) 3: 579, 1909.

* Lepidozia plumula Herzog, Beih. Bot. Centralbl. 38 (2): 331, 1921.

** Lepidozia portoricensis Fulford, Mem. New York Bot. Gard. 11 (2): 187, 1966.

*** Lepidozia procera Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 231, 1860.

* Lepidozia pseudocupressina Schiffn., Krit. Bemerk. Eur. Lebermoose 14: 9, 1919.

*** Lepidozia pumila J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 76, 2001.

*** Lepidozia quadridens (Nees) Nees, Syn. Hepat. 2: 209, 1845. Bas.: Jungermannia quadridens Nees, Enum. Pl. Crypt. Javae: 18, 1830.

** Lepidozia quadrifida Lindenb., Syn. Hepat. 2: 203, 1845.

*** Lepidozia reptans (L.) Dumort., Recueil Observ. Jungerm.: 19, 1835. Bas.: Jungermannia reptans L., Sp. Pl. 1: 1133, 1753.

** Lepidozia richardsii Herzog, Trans. Brit. Bryol. Soc. 1 (4): 313, 1950.

** Lepidozia rigida Steph., Sp. Hepat. (Stephani) 6: 340, 1922.

** Lepidozia robusta Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 217, 1894.

* Lepidozia rufescens Steph., Biblioth. Bot. 87 (2): 226, 1916.

** Lepidozia sandvicensis Lindenb., Syn. Hepat. 2: 201, 1845.

** Lepidozia schwabei Herzog, Rev. Bryol. Lichénol. 23 (1/2): 49, 1954.

** Lepidozia sellingiana H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 494, 1963.

*** Lepidozia septemfida Steph., Sp. Hepat. (Stephani) 3: 588, 1909.

** Lepidozia serpens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 364, 1885.

*** Lepidozia serrulata J.J.Engel, J. Hattori Bot. Lab. 96: 273, 2004.

*** Lepidozia setigera Steph., Sp. Hepat. (Stephani) 3: 599, 1909.

** Lepidozia sikkimensis Steph., Sp. Hepat. (Stephani) 6: 341, 1922.

*** Lepidozia spinosissima (Hook.f. et Taylor) Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 146, 1854. Bas.: Sendtnera spinosissima Hook.f. et Taylor, London J. Bot. 5: 373, 1846.

* Lepidozia squamifolia Steph., Sp. Hepat. (Stephani) 6: 341, 1922.

*** Lepidozia squarrosa Steph., Sp. Hepat. (Stephani) 3: 573, 1909.

** Lepidozia stahlii Steph., Sp. Hepat. (Stephani) 3: 616, 1909.

** Lepidozia stuhlmannii Steph., Bot. Jahrb. Syst. 20 (3): 308, 1895.

** Lepidozia stuhlmannii var. carnosa (Steph.) Pócs, Lidia 4 (1): 22, 1997. Bas.: Lepidozia carnosa Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 122, 1911.

** Lepidozia stuhlmannii subsp. pulvinata (Steph.) Pócs, Trop. Bryol. 9: 127, 1994. Bas.: Lepidozia pulvinata Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 121, 1911.

** Lepidozia subdichotoma Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 361, 1885.

** Lepidozia subintegra Lindenb., Syn. Hepat. 2: 201, 1845.

** Lepidozia subtransversa Steph., Bull. Herb. Boissier 5 (2): 95, 1897.

** Lepidozia subtrichodes Steph., Sp. Hepat. (Stephani) 3: 615, 1909.

** Lepidozia succida Mitt., Trans. Linn. Soc. London 23 (1): 57, 1860.

** Lepidozia supradecomposita Lindenb., Syn. Hepat. 2: 202, 1845.

* Lepidozia supradecomposita var. falcifolia Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 191, 1931.

** Lepidozia suyungii C.Gao et X.L.Bai, J. Hattori Bot. Lab. 92: 192, 2002.

** Lepidozia terricola Steph., Sp. Hepat. (Stephani) 3: 585, 1909.

** Lepidozia triangulifolia Steph., Sp. Hepat. (Stephani) 6: 344, 1922.

*** Lepidozia trichodes (Reinw., Blume et Nees) Nees, Syn. Hepat. 2: 203, 1845. Bas.: Jungermannia trichodes Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 199, 1824 [1825].

* Lepidozia tricuspidata Steph., Sp. Hepat. (Stephani) 6: 344, 1922.

** Lepidozia tunguraguae Steph., Sp. Hepat. (Stephani) 6: 345, 1922.

** Lepidozia ubangiensis Steph., Sp. Hepat. (Stephani) 3: 561, 1909.

*** Lepidozia udarii S.C.Srivast., D.Kumar et D.Sharma, Cryptog. Bryol. Lichénol. 9 (3): 237, 1988.

*** Lepidozia ulothrix (Schwägr.) Lindenb., Syn. Hepat. 2: 210, 1845. Bas.: Jungermannia ulothrix Schwägr., Hist. Musc. Hepat. Prodr.: 21, 1814.

** Lepidozia vitrea Steph., Bull. Herb. Boissier 5 (2): 96, 1897.

** Lepidozia wattsiana Steph., Sp. Hepat. (Stephani) 3: 586, 1909.

** Lepidozia weymouthiana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 117, 1914.

Excluded from the genus

* Lepidozia hexiloba Pearson, Ann. Cryptog. Exot. 4 (2): 67, 1931. 126

* Lepidozia parvistipa Taylor, London J. Bot. 5: 370, 1846. 127

*** Neolepidozia Fulford et J.Taylor, Brittonia 11 (2): 81, 1959.

*** Neolepidozia aubertii (Jovet-Ast) E.D.Cooper, Phytotaxa 97 (2): 55, 2013. Bas.: Lepidozia aubertii Jovet-Ast, Candollea 11: 35, 1947.

*** Neolepidozia autoica (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013. Bas.: Telaranea autoica J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 124, 2004.

*** Neolepidozia capilligera (Schwägr.) Fulford et J.Taylor, Brittonia 11 (2): 84, 1959. Bas.: Jungermannia capilligera Schwägr., Hist. Musc. Hepat. Prodr.: 21, 1814.

*** Neolepidozia consobrina (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013. Bas.: Telaranea consobrina J.J.Engel et G.L.Merr., Novon 9 (3): 339, 1999.

*** Neolepidozia cuneifolia (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959. Bas.: Lepidozia cuneifolia Steph., Sp. Hepat. (Stephani) 3: 618, 1909.

*** Neolepidozia disparata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013. Bas.: Telaranea disparata J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 147, 2004.

*** Neolepidozia disticha (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959. Bas.: Lepidozia disticha Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 62, 1911.

*** Neolepidozia gibbsiana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013. Bas.: Lepidozia gibbsiana Steph., Sp. Hepat. (Stephani) 6: 328, 1922.

*** Neolepidozia heterotexta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013. Bas.: Lepidozia heterotexta Steph., Sp. Hepat. (Stephani) 6: 329, 1922.

*** Neolepidozia hodgsoniae (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013. Bas.: Telaranea hodgsoniae J.J.Engel et G.L.Merr., Phytologia 79 (3): 251, 1995 [1996].

*** Neolepidozia longitudinalis (Herzog) E.D.Cooper, Phytotaxa 97 (2): 56, 2013. Bas.: Lepidozia longitudinalis Herzog, Trans. Brit. Bryol. Soc. 1 (4): 312, 1950.

*** Neolepidozia mamillosa (Schiffn.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013. Bas.: Lepidozia mamillosa Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 254, 1893.

*** Neolepidozia meridiana (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013. Bas.: Lepidozia meridiana E.A.Hodgs., Trans. Roy. Soc. New Zealand 83 (4): 611, 1956.

*** Neolepidozia oligophylla (Lehm. et Lindenb.) Fulford et J.Taylor, Brittonia 11 (2): 84, 1959. Bas.: Jungermannia oligophylla Lehm. et Lindenb., Nov. Stirp. Pug. 6: 26, 1834.

*** Neolepidozia ophiria (Gottsche) E.D.Cooper, Phytotaxa 97 (2): 56, 2013. Bas.: Lepidozia ophiria Gottsche, Sp. Hepat. (Stephani) 3: 611, 1909.

*** Neolepidozia palmata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013. Bas.: Telaranea palmata J.J.Engel et G.L.Merr., Novon 9 (3): 344, 1999.

*** Neolepidozia paludicola (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013. Bas.: Lepidozia meridiana var. paludicola E.A.Hodgs., Trans. Roy. Soc. New Zealand 83 (4): 611, 1956.

*** Neolepidozia papulosa (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959. Bas.: Lepidozia papulosa Steph., Sp. Hepat. (Stephani) 3: 609, 1909.

*** Neolepidozia parvifolia (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959. Bas.: Lepidozia parvifolia Steph., Sp. Hepat. (Stephani) 6: 337, 1922.

*** Neolepidozia patentissima (Hook.f. et Taylor) E.D.Cooper, Phytotaxa 97 (2): 56, 2013. Bas.: Jungermannia patentissima Hook.f. et Taylor, London J. Bot. 3: 286 [386], 1844.

*** Neolepidozia patentissima var. ampliata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013. Bas.: Telaranea patentissima var. ampliata J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 48, 2004.

*** Neolepidozia patentissima var. zebrina (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013. Bas.: Telaranea patentissima var. zebrina J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 50, 2004.

*** Neolepidozia pennata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013. Bas.: Telaranea pennata J.J.Engel et G.L.Merr., Phytologia 79 (3): 252, 1995 [1996].

*** Neolepidozia planifolia (Steph.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013. Bas.: Lepidozia planifolia Steph., Sp. Hepat. (Stephani) 3: 629, 1909.

*** Neolepidozia praenitens (Lehm. et Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013. Bas.: Jungermannia praenitens Lehm. et Lindenb., Nov. Stirp. Pug. 6: 27, 1834.

*** Neolepidozia praenitens var. dentifolia (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013. Bas.: Telaranea praenitens var. dentifolia J.J.Engel et G.L.Merr., Phytologia 79 (3): 253, 1995 [1996].

*** Neolepidozia quadristipula (Steph.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013. Bas.: Lepidozia quadristipula Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 115, 1914.

*** Neolepidozia rectangularis (R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013. Bas.: Telaranea rectangularis R.M.Schust., Phytologia 39 (4): 241, 1978.

*** Neolepidozia seriatitexta (Steph.) Fulford, Mem. New York Bot. Gard. 11 (2): 215, 1966. Bas.: Lepidozia seriatitexta Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 53, 1900.

*** Neolepidozia tetrapila (Hook.f. et Taylor) E.D.Cooper, Phytotaxa 97 (2): 57, 2013. Bas.: Lepidozia tetrapila Hook.f. et Taylor, London J. Bot. 5: 370, 1846.

*** Neolepidozia tetrapila var. cancellata (Colenso) E.D.Cooper, Phytotaxa 97 (2): 57, 2013. Bas.: Lepidozia cancellata Colenso, Trans. & Proc. New Zealand Inst. 18: 244, 1886.

*** Neolepidozia tetrapila var. roseana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013. Bas.: Lepidozia roseana Steph., Sp. Hepat. (Stephani) 3: 590, 1909.

*** Neolepidozia tridactylis (Lehm. et Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013. Bas.: Jungermannia tridactylis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 41, 1832.

*** Neolepidozia trifida (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013. Bas.: Lepidozia trifida Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 120, 1911.

*** Neolepidozia verruculosa (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013. Bas.: Telaranea verruculosa J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 197, 2004.

*** Neolepidozia wallichiana (Gottsche) Fulford et J.Taylor, Brittonia 11 (2): 84, 1959. Bas.: Lepidozia wallichiana Gottsche, Syn. Hepat. 2: 204, 1845.

*** Tricholepidozia (R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013. Bas.: Telaranea subg. Tricholepidozia R.M.Schust., J. Hattori Bot. Lab. 26: 256, 1963.

*** Tricholepidozia chaetocarpa (Pearson) E.D.Cooper, Phytotaxa 97 (2): 58, 2013. Bas.: Lepidozia chaetocarpa Pearson, J. Linn. Soc., Bot. 46 (305): 27, 1922.

*** Tricholepidozia fernandeziensis (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013. Bas.: Lepidozia fernandeziensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 63, 1911.

*** Tricholepidozia ferruginea (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013. Bas.: Telaranea ferruginea J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 159, 2004.

*** Tricholepidozia fissifolia (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013. Bas.: Lepidozia fissifolia Steph., Sp. Hepat. (Stephani) 3: 610, 1909.

*** Tricholepidozia jowettiana (H.A.Mill.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013. Bas.: Telaranea jowettiana H.A.Mill., J. Bryol. 14 (2): 235, 1986.

*** Tricholepidozia kaindina (Grolle) E.D.Cooper, Phytotaxa 97 (2): 59, 2013. Bas.: Telaranea kaindina Grolle, J. Hattori Bot. Lab. 31: 9, 1968.

*** Tricholepidozia kogiana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013. Bas.: Lepidozia kogiana Steph., Sp. Hepat. (Stephani) 6: 332, 1922.

*** Tricholepidozia lawesii (Steph.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013. Bas.: Lepidozia lawesii Steph., Hedwigia 28 (4): 264, 1889.

*** Tricholepidozia leratii (Steph.) E.D.Cooper, Phytotaxa 167 (2): 218, 2014. Bas.: Lepidozia leratii Steph., Sp. Hepat. (Stephani) 6: 333, 1922.

*** Tricholepidozia lindenbergii (Gottsche) E.D.Cooper, Phytotaxa 97 (2): 59, 2013. Bas.: Lepidozia lindenbergii Gottsche, Syn. Hepat. 2: 213, 1845.

*** Tricholepidozia lindenbergii var. complanata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013. Bas.: Telaranea lindenbergii var. complanata J.J.Engel et G.L.Merr., Phytologia 79 (3): 252, 1995 [1996].

*** Tricholepidozia lindenbergii var. mellea (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013. Bas.: Telaranea lindenbergii var. mellea J.J.Engel et G.L.Merr., Phytologia 79 (3): 252, 1995 [1996].

*** Tricholepidozia lindenbergii var. papillata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013. Bas.: Telaranea lindenbergii var. papillata J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 83, 2004.

*** Tricholepidozia marginata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013. Bas.: Telaranea marginata J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 166, 2004.

*** Tricholepidozia martinii (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013. Bas.: Lepidozia martinii E.A.Hodgs., Trans. Roy. Soc. New Zealand 83 (4): 602, 1956.

*** Tricholepidozia melanesica (H.A.Mill.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013. Bas.: Telaranea melanesica H.A.Mill., J. Bryol. 14 (2): 237, 1986.

*** Tricholepidozia neesii (Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013. Bas.: Lepidozia neesii Lindenb., Syn. Hepat. 2: 212, 1845.

*** Tricholepidozia octoloba (Del Ros.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013. Bas.: Telaranea octoloba Del Ros., Philipp. J. Sci. 100 (3/4): 239, 1971.

*** Tricholepidozia plumulosa (Lehm. et Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013. Bas.: Jungermannia plumulosa Lehm. et Lindenb., Nov. Stirp. Pug. 6: 30, 1834.

*** Tricholepidozia pulcherrima (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013. Bas.: Lepidozia pulcherrima Steph., Sp. Hepat. (Stephani) 3: 600, 1909.

*** Tricholepidozia pulcherrima var. mooreana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013. Bas.: Lepidozia mooreana Steph., Sp. Hepat. (Stephani) 3: 585, 1909.

*** Tricholepidozia quadriseta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013. Bas.: Lepidozia quadriseta Steph., Sp. Hepat. (Stephani) 3: 582, 1909.

*** Tricholepidozia remotifolia (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013. Bas.: Telaranea remotifolia E.A.Hodgs., Rec. Domin. Mus. 4 (11): 107, 1962.

*** Tricholepidozia semperiana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013. Bas.: Lepidozia semperiana Steph., Sp. Hepat. (Stephani) 3: 612, 1909.

*** Tricholepidozia tetradactyla (Hook.f. et Taylor) E.D.Cooper, Phytotaxa 97 (2): 61, 2013. Bas.: Jungermannia tetradactyla Hook.f. et Taylor, London J. Bot. 3: 286 [386], 1844.

*** Tricholepidozia trichocoleoides (Herzog) E.D.Cooper, Phytotaxa 97 (2): 61, 2013. Bas.: Lepidozia trichocoleoides Herzog, Trans. Brit. Bryol. Soc. 1 (4): 314, 1950.

** Micropterygioideae Grolle

** Micropterygium Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 233, 1845.

** Micropterygium angustistipulum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 385, 1885.

*** Micropterygium bialatum Fulford, Mem. New York Bot. Gard. 11 (2): 268, 1966.

*** Micropterygium bolivarense Fulford, Mem. New York Bot. Gard. 11 (2): 263, 1966.

*** Micropterygium campanense Spruce, Hedwigia 73 (3/4): 157, 1933.

*** Micropterygium carinatum (Grev.) Reimers, Hedwigia 76 (3): 166, 1936. Bas.: Jungermannia carinata Grev., Ann. Lyceum Nat. Hist. New York 1 (2): 276, 1825.

*** Micropterygium conchifolium Reimers, Hedwigia 73 (3/4): 155, 1933.

*** Micropterygium duidae Reimers, Hedwigia 73 (3/4): 147, 1933.

** Micropterygium exalatum Steph., Sp. Hepat. (Stephani) 3: 547, 1909.

** Micropterygium grandistipulum Steph., Trans. Linn. Soc. London, Bot. 6 (1): 98, 1901.

** Micropterygium laeve H.Rob., Bol. Soc. Venez. Ci. Nat. 32 (132/133): 252, 1976.

** Micropterygium lechleri Reimers, Hedwigia 73 (3/4): 184, 1933.

*** Micropterygium leiophyllum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 386, 1885.

** Micropterygium longicellulatum Uribe et E.L.Linares, Phytotaxa 213 (3): 297, 2015.

** Micropterygium parvistipulum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 383, 1885.

*** Micropterygium pterygophyllum (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 413, 1877. Bas.: Jungermannia pterygophylla Nees, Fl. Bras. (Martius) 1 (1): 377, 1833.

* Micropterygium pterygophyllum var. lancifolium Reimers, Hedwigia 73 (3/4): 176, 1933.

** Micropterygium reimersianum Herzog, Hedwigia 81 (5/6): 226, 1943.

*** Micropterygium steyermarkii Fulford, Mem. New York Bot. Gard. 11 (2): 270, 1966.

*** Micropterygium tatei Reimers, Hedwigia 73 (3/4): 150, 1933.

*** Micropterygium tenax (Steph.) Grolle, J. Bryol. 10 (3): 265, 1979. Bas.: Harpalejeunea tenax Steph., Trans. Linn. Soc. London, Bot. 6 (1): 100, 1901.

*** Micropterygium trachyphyllum Reimers, Hedwigia 73 (3/4): 186, 1933. 128

* Micropterygium trachyphyllum var. brasiliense Reimers, Hedwigia 73 (3/4): 195, 1933.

* Micropterygium trachyphyllum var. cubense Reimers, Hedwigia 73 (3/4): 188, 1933.

* Micropterygium trachyphyllum var. guadeloupense Reimers, Hedwigia 73 (3/4): 190, 1933.

* Micropterygium trachyphyllum var. jamaicense Reimers, Hedwigia 73 (3/4): 190, 1933.

*** Micropterygium tumidulum Fulford, Mem. New York Bot. Gard. 11 (2): 272, 1966.

** Mytilopsis Spruce, Cephalozia: 90, 1882.

*** Mytilopsis albifrons Spruce, Cephalozia: 91, 1882.

** Protocephalozioideae R.M.Schust.

** Protocephalozia (Spruce) K.I.Goebel, Flora 77 (2): 83, 1893. Bas.: Cephalozia subg. Protocephalozia Spruce, Cephalozia: 24, 1882.

*** Protocephalozia ephemeroides (Spruce) K.I.Goebel, Flora 77 (2): 83, 1893. Bas.: Cephalozia ephemeroides Spruce, Cephalozia: 24, 1882.

*** Zoopsidoideae R.M.Schust.

** Amazoopsis J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 242, 2004.

*** Amazoopsis diplopoda (Pócs) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 245, 2004. Bas.: Arachniopsis diplopoda Pócs, Proc. Third Meeting Bryol. C. & E. Europe: 114, 1984.

*** Amazoopsis dissotricha (Spruce) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 247, 2004. Bas.: Arachniopsis dissotricha Spruce, Cephalozia: 86, 1882.

*** Amazoopsis gracilis J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 246, 2004.

* Hyalolepidozia S.W.Arnell ex Grolle, Rev. Bryol. Lichénol. 32 (3/4): 179, 1963 [1964]. Based on: Hyalolepidozia S.W.Arnell, Bot. Not. 115: 203, 1962.

** Hyalolepidozia bicuspidata (C.Massal.) S.W.Arnell ex Grolle, Rev. Bryol. Lichénol. 32 (3/4): 179, 1963 [1964]. Bas.: Lepidozia bicuspidata C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 239, 1885.

* Monodactylopsis (R.M.Schust.) R.M.Schust., Nova Hedwigia 56 (1/2): 45, 1993. Bas.: Arachniopsis subg. Monodactylopsis R.M.Schust., Nova Hedwigia 10 (1/2): 24, 1965.

** Monodactylopsis monodactyla (Spruce) R.M.Schust., Nova Hedwigia 69 (3/4): 523, 1999. Bas.: Cephalozia monodactyla Spruce, Cephalozia: 28, 1882.

** Neogrollea E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 70, 1965.

*** Neogrollea notabilis E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 70, 1965.

* Odontoseries Fulford, Mem. New York Bot. Gard. 11 (3): 364, 1968.

* Odontoseries chimantana Fulford, Mem. New York Bot. Gard. 11 (3): 366, 1968.

*** Paracromastigum Fulford et J.Taylor, Brittonia 13 (4): 336, 1961.

*** Paracromastigum denticulatum (Steph.) E.D.Cooper, Phytotaxa 97 (2): 61, 2013. Bas.: Lembidium denticulatum Steph., Sp. Hepat. (Stephani) 6: 444, 1924.

*** Paracromastigum drucei (R.M.Schust.) R.M.Schust., J. Hattori Bot. Lab. 38: 700, 1974. Bas.: Pseudocephalozia drucei R.M.Schust., Nova Hedwigia 10 (1/2): 22, 1965.

*** Paracromastigum dusenii (Steph.) R.M.Schust., J. Hattori Bot. Lab. 38: 700, 1974. Bas.: Alobiella dusenii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 48, 1900.

** Paracromastigum fiordlandiae R.M.Schust. et J.J.Engel, Brittonia 48 (2): 167, 1996.

*** Paracromastigum furcifolium (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 276, 1963. Bas.: Cephalozia furcifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 485 (315), 1908.

** Paracromastigum granatense (Gottsche) R.M.Schust., J. Hattori Bot. Lab. 48: 341, 1980. Bas.: Lepidozia granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 139, 1864.

** Paracromastigum longiscyphum (Taylor) R.M.Schust. et J.J.Engel, Brittonia 48 (2): 167, 1996. Bas.: Jungermannia longiscypha Taylor, London J. Bot. 5: 280, 1846.

** Paracromastigum macrostipum (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 276, 1963. Bas.: Cephalozia macrostipa Steph., Hedwigia 32 (5): 315, 1893.

** Paracromastigum micromera (Spruce) R.M.Schust., J. Hattori Bot. Lab. 26: 276, 1963. Bas.: Cephalozia micromera Spruce, Cephalozia: 32, 1882.

** Paracromastigum microphyllum (R.M.Schust. ex J.J.Engel) E.D.Cooper, Phytotaxa 97 (2): 61, 2013. Bas.: Hyalolepidozia microphylla R.M.Schust. ex J.J.Engel, Novon 17 (3): 310, 2007.

*** Paracromastigum pachyrhizum (Nees) Fulford, Mem. New York Bot. Gard. 11 (3): 390, 1968. Bas.: Jungermannia pachyrhiza Nees, Fl. Bras. (Martius) 1 (1): 339, 1833.

** Paracromastigum ryszardii Váňa, Bedn.-Ochyra et Cykowska, Nova Hedwigia 89 (1/2): 122, 2009.

** Paracromastigum stipulatum (Herzog) Fulford, Mem. New York Bot. Gard. 11 (3): 390, 1968. Bas.: Cephalozia stipulata Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 105, 1952.

** Paracromastigum subsimplex (Steph.) Fulford et J.Taylor, Brittonia 13 (4): 336, 1961. Bas.: Lepidozia subsimplex Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 66, 1911.

** Paracromastigum succulentum (Sim) J.J.Engel et G.L.Merr., Bryologist 104 (1): 151, 2001. Bas.: Lepidozia succulenta Sim, Trans. Roy. Soc. South Africa 15 (1): 90, 1926.

** Paracromastigum tristanianum (R.M.Schust.) J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 38: 700, 1974. Bas.: Pseudocephalozia tristaniana R.M.Schust., Nova Hedwigia 10 (1/2): 23, 1965.

** Paracromastigum vastilobum (Steph.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 255, 2004. Bas.: Lepidozia vastiloba Steph., Sp. Hepat. (Stephani) 3: 581, 1909.

** Psiloclada Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 143, 1854.

*** Psiloclada clandestina Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 143, 1854.

** Psiloclada clandestina subsp. melanesica R.M.Schust., J. Hattori Bot. Lab. 48: 411, 1980.

** Psiloclada clandestina subsp. spinosa (S.W.Arnell) R.M.Schust., J. Hattori Bot. Lab. 48: 410, 1980. Bas.: Lepidozia spinosa S.W.Arnell, Bot. Not. 107: 427, 1954.

** Pteropsiella Spruce, J. Bot. 14: 161, 1876.

*** Pteropsiella frondiformis Spruce, J. Bot. 14: 161, 1876.

** Pteropsiella metzgeriiformis R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965.

** Telaranea Spruce ex Schiffn., Hepat. (Engl.-Prantl): 103, 1893 nom. conserv. Based on: Telaranea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 365, 1885. 129

* sect. Telaranea

*** Telaranea apiahyna (Steph.) Fulford, Brittonia 15 (1): 71, 1963. Bas.: Lepidozia apiahyna Steph., Sp. Hepat. (Stephani) 3: 572, 1909.

*** Telaranea bicruris (Steph.) M.Howe, Bull. Torrey Bot. Club 29 (5): 287, 1902. Bas.: Lepidozia bicruris Steph., Hedwigia 24 (4): 166, 1885.

*** Telaranea blepharostoma (Steph.) Fulford, Brittonia 15 (1): 73, 1963. Bas.: Lepidozia blepharostoma Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 22, 1901.

*** Telaranea breviseta (Herzog) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 131, 2004. Bas.: Lepidozia sejuncta var. breviseta Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 723, 1942.

*** Telaranea chaetophylla (Spruce) Schiffn., Hepat. (Engl.-Prantl): 103, 1893. Bas.: Lepidozia chaetophylla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 365, 1885.

*** Telaranea europaea J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 150, 2004.

*** Telaranea fragilis Mizut., J. Hattori Bot. Lab. 40: 449, 1976.

*** Telaranea granulata J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 103, 2004.

*** Telaranea longifolia (M.Howe) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 163, 2004. Bas.: Telaranea nematodes var. longifolia M.Howe, Bull. Torrey Bot. Club 29 (5): 286, 1902.

*** Telaranea nematodes (Gottsche ex Austin) M.Howe, Bull. Torrey Bot. Club 29 (5): 284, 1902. Bas.: Cephalozia nematodes Gottsche ex Austin, Bull. Torrey Bot. Club 6 (52): 302, 1879.

*** Telaranea panchoi Del Ros., Philipp. J. Sci. 100 (3/4): 238, 1971.

*** Telaranea pellucida J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 179, 2004.

*** Telaranea pseudozoopsis (Herzog) Fulford, Brittonia 15 (1): 71, 1963. Bas.: Lepidozia pseudozoopsis Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 723, 1942.

*** Telaranea redacta (Steph.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 186, 2004. Bas.: Lepidozia redacta Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 119, 1911.

*** Telaranea rosarioana H.A.Mill., J. Bryol. 14 (2): 240, 1986.

*** Telaranea setosa J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 192, 2004.

*** Telaranea trisetosa (Steph.) Grolle, J. Hattori Bot. Lab. 29: 280, 1966. Bas.: Lepidozia trisetosa Steph., Sp. Hepat. (Stephani) 3: 607, 1909.

* sect. Tenuifoliae (R.M.Schust.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 112, 2004. Bas.: Arachniopsis sect. Tenuifoliae R.M.Schust., Beih. Nova Hedwigia 118: 461, 2000.

*** Telaranea anomala R.M.Schust. ex J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 121, 2004.

** Telaranea bischleriana Pócs, Acta Bot. Hung. 48 (1/2): 120, 2006.

*** Telaranea coactilis (Spruce) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 140, 2004. Bas.: Arachniopsis coactilis Spruce, Cephalozia: 85, 1882.

*** Telaranea confervoides J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 143, 2004. Based on: Arachniopsis pecten var. confervoides R.M.Schust., Beih. Nova Hedwigia 118: 455, 2000, nom. inval.

*** Telaranea diacantha (Mont.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 145, 2004. Bas.: Jungermannia diacantha Mont., Ann. Sci. Nat. Bot. (sér. 4) 5: 349, 1856.

*** Telaranea herzogii (E.A.Hodgs.) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 106, 1962. Bas.: Lepidozia herzogii E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 78 (4): 500, 1950.

*** Telaranea inaequalis R.M.Schust. ex J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 117, 2004.

** Telaranea maorensis Pócs, Acta Bot. Hung. 48 (1/2): 124, 2006.

*** Telaranea microstipulata R.M.Schust., Phytologia 39 (4): 241, 1978.

*** Telaranea monocera (Mitt. ex R.M.Schust. et Grolle) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 168, 2004. Bas.: Arachniopsis monocera Mitt. ex R.M.Schust. et Grolle, Nova Hedwigia 10 (1/2): 25, 1965.

*** Telaranea pecten (Spruce) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 178, 2004. Bas.: Arachniopsis pecten Spruce, Cephalozia: 85, 1882.

*** Telaranea sejuncta (Ångstr.) S.W.Arnell, Bot. Not. 110 (1): 18, 1957. Bas.: Blepharostoma sejunctum Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 78, 1876 [1877].

*** Telaranea tenuifolia J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 193, 2004. Based on: Arachniopsis tenuifolia R.M.Schust., Beih. Nova Hedwigia 118: 461, 2000, nom. inval.

Incertae sedis

** Telaranea azorica (H.Buch et Perss.) Pócs in Schumacker et Váňa, Identif. keys liverw. hornw. Europe: 160, 2005. Bas.: Lepidozia azorica H.Buch et Perss., Bryophyt. Azoren Madeira: 4, 1941.

** Telaranea indica (S.C.Srivast. et P.K.Verma) A.E.D.Daniels et P.Daniel, Bull. Bot. Surv. India 49 (1/4): 231, 2007. Bas.: Arachniopsis indica S.C.Srivast. et P.K.Verma, Natl. Acad. Sci. Lett. 27 (7/8): 270, 2004 [2006].

** Telaranea major (Herzog) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 165, 2004. Bas.: Arachniopsis major Herzog, Trans. Brit. Bryol. Soc. 1 (4): 294, 1950.

** Zoopsidella R.M.Schust., Nova Hedwigia 10 (1/2): 24, 1965.

** Zoopsidella antillana (Steph.) H.Rob., Bol. Soc. Venez. Ci. Nat. 32 (132/133): 254, 1976. Bas.: Zoopsis antillana Steph., Bull. Herb. Boissier (sér. 2) 8 (4): 268 (282), 1908.

** Zoopsidella antillana subsp. jamaicensis R.M.Schust., Nova Hedwigia 69 (1/2): 132, 1999.

*** Zoopsidella caledonica (Steph.) R.M.Schust., Taxon 18 (1): 57, 1969. Bas.: Zoopsis caledonica Steph., Sp. Hepat. (Stephani) 6: 318, 1922.

** Zoopsidella cynosurandra (Steph.) R.M.Schust., Nova Hedwigia 10 (1/2): 24, 1965. Bas.: Zoopsis cynosurandra Steph., Bull. Herb. Boissier (sér. 2) 8 (4): 269 (283), 1908.

*** Zoopsidella integrifolia (Spruce) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969. Bas.: Cephalozia integrifolia Spruce, Cephalozia: 29, 1882.

** Zoopsidella macella (Spruce) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969. Bas.: Cephalozia macella Spruce, Cephalozia: 29, 1882.

*** Zoopsidella serra (Spruce) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969. Bas.: Cephalozia serra Spruce, Cephalozia: 32, 1882.

** Zoopsis Hook.f. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 473, 1846.

** subg. Eozoopsis R.M.Schust., J. Hattori Bot. Lab. 36: 373, 1972.

*** Zoopsis leitgebiana (Carrington et Pearson) Bastow, Pap. & Proc. Roy. Soc. Tasmania 1887: 269, 1888. Bas.: Cephalozia leitgebiana Carrington et Pearson, Pap. & Proc. Roy. Soc. Tasmania 1887: 3, 1888.

*** Zoopsis macrophylla R.M.Schust., Nova Hedwigia 68 (1/2): 14, 1999.

** subg. Zoopsis

*** Zoopsis argentea (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 473, 1846. Bas.: Jungermannia argentea Hook.f. et Taylor, London J. Bot. 3: 400, 1844.

** Zoopsis argentea var. flagelliformis (Colenso) R.M.Schust., Nova Hedwigia 68 (1/2): 38, 1999. Bas.: Zoopsis flagelliformis Colenso, Trans. & Proc. New Zealand Inst. 18: 250, 1886.

*** Zoopsis bicruris Glenny et E.A.Br., J. Bryol. 28 (4): 332, 2006.

** Zoopsis liukiuensis Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 65, 1931.

*** Zoopsis matawaia M.A.M.Renner, J. Bryol. 28 (4): 334, 2006.

*** Zoopsis nitida Glenny, Braggins et R.M.Schust., J. Bryol. 19 (4): 776, 1997.

*** Zoopsis setulosa Leitg., Mitt. Naturwiss. Vereins Steiermark 13: 24, 1876.

Incertae sedis

*** Zoopsis ceratophylla (Spruce) Hamlin, Rec. Domin. Mus. 7: 311, 1972. Bas.: Cephalozia ceratophylla Spruce, Cephalozia: 32, 1882.

* Zoopsis ciliata Colenso, Trans. & Proc. New Zealand Inst. 20: 253, 1888. 130

* Zoopsis martinicensis Steph., Bull. Herb. Boissier (sér. 2) 8 (4): 268 (282), 1908.

** Zoopsis setigera K.I.Goebel, Flora 77 (2): 93, 1893.

** Zoopsis uleana Steph., Hedwigia 44 (4): 225, 1905.

*** Lophocoleaceae Vanden Berghen

by B.J. Crandall-Stotler, R. Stotler, J. Váňa, J.J.Engel and L. Söderström

outlined the current status of Lophocoleaceae noting that several taxonomic problems in delimitating genera remain. The placement of several species is also unclear. Nomenclatural and taxonomic notes can be found in, ).

** Bragginsella R.M.Schust., Bryologist 100 (3): 363, 1997.

*** Bragginsella anomala R.M.Schust., Bryologist 100 (3): 363, 1997.

*** Chiloscyphus Corda, Gen. hepat.: 651, 1829 nom. conserv.

** Chiloscyphus kashyapii A.Srivast. et S.C.Srivast., Indian Geocalyc.: 34, 2002.

*** Chiloscyphus pallescens (Ehrh.) Dumort., Syll. Jungerm. Europ.: 67, 1831. Bas.: Jungermannia pallescens Ehrh., Deutschl. Fl., Theil 2 (Hoffm.): 87, 1795 [1796].

* Chiloscyphus pallescens var. fragilis (Roth) Müll.Frib., Ber. Deutsch. Bot. Ges. 59 (10): 429, 1942. Bas.: Jungermannia fragilis Roth, Tent. Fl. Germ. 3: 370, 1800. 131

*** Chiloscyphus polyanthos (L.) Corda, Gen. hepat.: 651, 1829. Bas.: Jungermannia polyanthos L., Sp. Pl. 1: 1131, 1753.

* Chiloscyphus polyanthos var. rivularis (Schrad.) Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 24, 1889. Bas.: Jungermannia pallescens var. rivularis Schrad., Syst. Samml. Crypt. Gew. 2: 7, 1797. 132

Incertae sedis 133

** Chiloscyphus acutus Steph., Sp. Hepat. (Stephani) 6: 302, 1922.

** Chiloscyphus alpicola J.J.Engel, Phytotaxa 207 (2): 181, 2015.

** Chiloscyphus beesleyanus Pearson, J. Linn. Soc., Bot. 46 (305): 22, 1922.

* Chiloscyphus bifidus Schiffn., Hep. Fl. Buitenzorg: 200, 1900. 134

** Chiloscyphus breviculus B.Y.Yang et W.C.Lee, Bot. Bull. Acad. Sin. (n.ser.) 5 (2): 190, 1964.

* Chiloscyphus brevistipulus Steph., Sp. Hepat. (Stephani) 6: 303, 1922.

** Chiloscyphus chinnarensis Manju, K.P.Rajesh et Madhus., Acta Bot. Hung. 53 (1/2): 152, 2011.

** Chiloscyphus confertifolius Steph., Sp. Hepat. (Stephani) 6: 304, 1922.

** Chiloscyphus confertus Steph., Sp. Hepat. (Stephani) 6: 305, 1922.

** Chiloscyphus cornutistipulus Steph., Sp. Hepat. (Stephani) 6: 303, 1922.

* Chiloscyphus durus (Steph.) Hässel, Revista Mus. Argent. Ci. Nat. (n.ser.) 1 (2): 122, 1999. Bas.: Lophocolea dura Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 43, 1911. 135

* Chiloscyphus ernstianus Steph., Sp. Hepat. (Stephani) 6: 306, 1922.

** Chiloscyphus etesseanus Steph., Bull. Herb. Boissier (sér. 2) 7 (10): 845 (217), 1907.

** Chiloscyphus francanus Steph., Sp. Hepat. (Stephani) 6: 306, 1922.

** Chiloscyphus graeffeanus Steph., Sp. Hepat. (Stephani) 6: 307, 1922.

** Chiloscyphus greenwelliae (H.A.Mill.) H.A.Mill., J. Hattori Bot. Lab. 30: 275, 1967. Bas.: Lophocolea greenwelliae H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 504, 1963.

** Chiloscyphus hookeri J.J.Engel, J. Hattori Bot. Lab. 36: 150, 1972 [1973].

* Chiloscyphus hookeri var. constantifolius J.J.Engel, J. Hattori Bot. Lab. 36: 155, 1972 [1973].

** Chiloscyphus integerrimus Schiffn., Hep. Fl. Buitenzorg: 197, 1900.

** Chiloscyphus kehdingianus (Steph.) N.Kitag., Hikobia, Suppl. 1: 68, 1981. Bas.: Lophocolea kehdingiana Steph., Sp. Hepat. (Stephani) 6: 278, 1922.

** Chiloscyphus kilauensis Steph., Sp. Hepat. (Stephani) 6: 309, 1922.

** Chiloscyphus koeppensis (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 139 (255), 1908. Bas.: Jungermannia koeppensis Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 452, 1890.

** Chiloscyphus laceratus Steph., Sp. Hepat. (Stephani) 6: 310, 1922.

** Chiloscyphus lambertonii H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 506, 1963.

** Chiloscyphus latistipus Steph., Sp. Hepat. (Stephani) 6: 309, 1922.

** Chiloscyphus lepervanchei (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 418, 1984 [1985]. Bas.: Lophocolea lepervanchei Steph., Bull. Herb. Boissier (sér. 2) 7 (4): 310 (174), 1907.

** Chiloscyphus longifissus Steph., Sp. Hepat. (Stephani) 6: 310, 1922.

** Chiloscyphus propagulifer Schiffn., Hep. Fl. Buitenzorg: 208, 1900.

** Chiloscyphus purpureus Steph., Sp. Hepat. (Stephani) 6: 312, 1922.

** Chiloscyphus quadricilius Steph., Sp. Hepat. (Stephani) 6: 312, 1922.

*** Chiloscyphus quadridentatus (Spruce) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 422, 1984 [1985]. Bas.: Lophocolea quadridentata Spruce, Mem. Torrey Bot. Club 1 (3): 137, 1890.

** Chiloscyphus rotundifolius Mitt., Rep. Challenger, Bot. 1 (3, 1): 85, 1884.

** Chiloscyphus rotundiphyllus H.A.Mill., Phytologia 47 (4): 321, 1981. Nom. nov. pro Chiloscyphus rotundifolius Steph., Sp. Hepat. (Stephani) 6: 313, 1922, nom. illeg.

** Chiloscyphus scaberulus Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cc, 1889 [1890].

** Chiloscyphus septatus J.J.Engel, Fieldiana, Bot. (n.ser.) 48: 125, 2010.

** Chiloscyphus similis Steph., Rev. Bryol. 35 (2): 28, 1908.

** Chiloscyphus skottsbergianus H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 508, 1963.

** Chiloscyphus subacuminatus Herzog, Ark. Bot. (n.ser.) 3 (3): 49, 1953.

** Chiloscyphus subsimilis Steph., Sp. Hepat. (Stephani) 6: 314, 1922.

* Chiloscyphus tridens Steph., Sp. Hepat. (Stephani) 6: 315, 1922.

** Chiloscyphus trigonifolius Steph., Sp. Hepat. (Stephani) 6: 316, 1922.

* Chiloscyphus venustulus Colenso, Trans. & Proc. New Zealand Inst. 21: 60, 1889.

** Clasmatocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 440, 1885. 136

*** Clasmatocolea bisexualis Glenny et J.J.Engel, New Zealand J. Bot. 51 (1): 23, 2013.

*** Clasmatocolea crassiretis (Herzog) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 69, 1962 [1963]. Bas.: Lophocolea crassiretis Herzog, Trans. & Proc. Roy. Soc. New Zealand 65 (3): 354, 1936.

*** Clasmatocolea ctenophylla (Schiffn.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 71, 1960. Bas.: Lophocolea ctenophylla Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 12, 1890.

*** Clasmatocolea cucullistipula (Steph.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 71, 1960. Bas.: Lophocolea cucullistipula Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 37, 1900.

*** Clasmatocolea fasciculata (Nees) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960. Bas.: Jungermannia fasciculata Nees, Horae Phys. Berol.: 46, 1820.

*** Clasmatocolea fulvella (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960. Bas.: Jungermannia fulvella Hook.f. et Taylor, London J. Bot. 3: 464, 1844.

*** Clasmatocolea gayana (Mont.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960. Bas.: Jungermannia gayana Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 349, 1845.

*** Clasmatocolea humilis (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960. Bas.: Jungermannia humilis Hook.f. et Taylor, London J. Bot. 3: 468, 1844.

*** Clasmatocolea humilis var. polymorpha J.J.Engel, Phytologia 41 (5): 309, 1979.

*** Clasmatocolea humilis var. suspecta (C.Massal.) J.J.Engel, Phytologia 41 (5): 309, 1979. Bas.: Lophocolea puccioana β suspecta C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 228, 1885.

*** Clasmatocolea inflexispina (Hook.f. et Taylor) J.J.Engel, Bryologist 94 (4): 436, 1991. Bas.: Jungermannia inflexispina Hook.f. et Taylor, London J. Bot. 4: 82, 1845.

*** Clasmatocolea marginata (Steph.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 73, 1962 [1963]. Bas.: Leioscyphus marginatus Steph., Bull. Herb. Boissier (sér. 2) 6 (3): 223 (23), 1906.

*** Clasmatocolea minutiretis J.J.Engel et Grolle, Phytologia 41 (5): 309, 1979.

*** Clasmatocolea moniliformis J.J.Engel, Phytologia 41 (5): 310, 1979.

*** Clasmatocolea navistipula (Steph.) Grolle, Feddes Repert. 82 (1): 88, 1971. Bas.: Lophocolea navistipula Steph., Bull. Herb. Boissier (sér. 2) 6 (7): 543 (57), 1906.

*** Clasmatocolea navistipula var. parceramosa J.J.Engel, Phytologia 41 (5): 311, 1979.

*** Clasmatocolea notophylla (Hook.f. et Taylor) Grolle, J. Jap. Bot. 41 (8): 228, 1966. Bas.: Jungermannia notophylla Hook.f. et Taylor, London J. Bot. 3: 376, 1844.

*** Clasmatocolea obvoluta (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960. Bas.: Jungermannia obvoluta Hook.f. et Taylor, London J. Bot. 4: 80, 1845.

*** Clasmatocolea obvoluta var. cookiana (C.Massal.) J.J.Engel, Phytologia 41 (5): 311, 1979. Bas.: Lophocolea cookiana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 224, 1885.

*** Clasmatocolea puccioana (De Not.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960. Bas.: Jungermannia puccioana De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 221, 1857.

*** Clasmatocolea rigens (Hook.f. et Taylor) J.J.Engel, J. Hattori Bot. Lab. 36: 156, 1972 [1973]. Bas.: Jungermannia rigens Hook.f. et Taylor, London J. Bot. 3: 461, 1844.

*** Clasmatocolea strongylophylla (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 73, 1960. Bas.: Jungermannia strongylophylla Hook.f. et Taylor, London J. Bot. 3: 370, 1844.

*** Clasmatocolea trachyopa (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 73, 1960. Bas.: Jungermannia trachyopa Hook.f. et Taylor, London J. Bot. 3: 471, 1844.

*** Clasmatocolea vermicularis (Lehm.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 78, 1960. Bas.: Jungermannia vermicularis Lehm., Linnaea 4: 361, 1829.

*** Clasmatocolea verrucosa J.J.Engel, Bryologist 83 (2): 220, 1980.

*** Conoscyphus Mitt., Fl. vit.: 404, 1871 [1873].

** Conoscyphus koponenii Piippo, Mamontov et Potemkin, Acta Bryolichenol. Asiat. 5: 20, 2014.

*** Conoscyphus trapezioides (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 125, 1898. Bas.: Chiloscyphus trapezioides Sande Lac., Ned. Kruidk. Arch. 3: 417, 1854 [1855].

*** Cryptolophocolea L.Söderstr., Crand.-Stotl., Stotler et Váňa, Phytotaxa 97 (2): 39, 2013. Based on: Plagiochila sect. Connatae Lindenb., Monogr. hep. gen. Plagiochilae: xxix, 1844 [1843].

*** Cryptolophocolea aculeata (Mitt.) L.Söderstr., Phytotaxa 112 (1): 21, 2013. Bas.: Chiloscyphus aculeatus Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 140, 1854.

** Cryptolophocolea chiloscyphoidea (Lindenb.) L.Söderstr. et Crand.-Stotl., Phytotaxa 112 (1): 18, 2013. Bas.: Plagiochila chiloscyphoidea Lindenb., Nov. Stirp. Pug. 8: 4, 1844.

*** Cryptolophocolea ciliolata (Nees) L.Söderstr., Crand.-Stotl., Stotler et Váňa, Phytotaxa 97 (2): 39, 2013. Bas.: Jungermannia ciliolata Nees, Enum. Pl. Crypt. Javae: 68, 1830.

** Cryptolophocolea compacta (Mitt.) L.Söderstr., Phytotaxa 112 (1): 19, 2013. Bas.: Lophocolea compacta Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 198, 1891.

*** Cryptolophocolea connata (Sw.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 19, 2013. Bas.: Jungermannia connata Sw., Prodr. (Swartz): 143, 1788.

*** Cryptolophocolea connatifolia (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 21, 2013. Bas.: Chiloscyphus connatifolius J.J.Engel, Phytologia 83 (1): 42, 1997 [1998].

*** Cryptolophocolea costata (Nees) L.Söderstr., Phytotaxa 112 (1): 19, 2013. Bas.: Jungermannia costata Nees, Enum. Pl. Crypt. Javae: 69, 1830.

*** Cryptolophocolea edentata (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 21, 2013. Bas.: Chiloscyphus edentatus J.J.Engel, Phytologia 83 (1): 43, 1997 [1998].

** Cryptolophocolea explanata (Mitt.) Váňa et Crand.-Stotl., Phytotaxa 202 (1): 69, 2015. Bas.: Lophocolea explanata Mitt., Fl. vit.: 404, 1871 [1873].

** Cryptolophocolea fleischeri (Steph.) L.Söderstr., Phytotaxa 112 (1): 21, 2013. Bas.: Lophocolea fleischeri Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 952 (132), 1906.

*** Cryptolophocolea guadalupensis (Steph.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 19, 2013. Bas.: Lophocolea guadalupensis Steph., Bull. Herb. Boissier (sér. 2) 7 (1): 65 (153), 1907.

*** Cryptolophocolea helmsiana (Steph.) L.Söderstr., Phytotaxa 112 (1): 19, 2013. Bas.: Lophocolea helmsiana Steph., Bull. Herb. Boissier (sér. 2) 6 (9): 794 (94), 1906.

*** Cryptolophocolea leucophylla (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 19, 2013. Bas.: Jungermannia leucophylla Hook.f. et Taylor, London J. Bot. 3: 384, 1844.

* Cryptolophocolea levieri (Schiffn.) L.Söderstr., Phytotaxa 112 (1): 21, 2013. Bas.: Lophocolea levieri Schiffn., Hep. Fl. Buitenzorg: 182, 1900. 137

* Cryptolophocolea lilliena (Steph.) L.Söderstr., Phytotaxa 112 (1): 21, 2013. Bas.: Lophocolea lilliena Steph., Sp. Hepat. (Stephani) 6: 282, 1922. 138

*** Cryptolophocolea martiana (Nees) L.Söderstr., Crand.-Stotl. et Stotler, Phytotaxa 112 (1): 20, 2013. Bas.: Lophocolea martiana Nees, Syn. Hepat. 2: 152, 1845.

** Cryptolophocolea martiana subsp. bidentula (Nees) L.Söderstr., Crand.-Stotl. et Stotler, Phytotaxa 112 (1): 20, 2013. Bas.: Chiloscyphus bidentulus Nees, Syn. Hepat. 2: 181, 1845.

** Cryptolophocolea martiana var. perissodonta (Spruce) Gradst., Phytoneuron 2015 (22): 1, 2015. Bas.: Lophocolea martiana var. perissodonta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 432, 1885.

* Cryptolophocolea massalongoana (Schiffn.) L.Söderstr., Phytotaxa 112 (1): 20, 2013. Bas.: Lophocolea massalongoana Schiffn., Hep. Fl. Buitenzorg: 183, 1900. 139

*** Cryptolophocolea mitteniana (Colenso) L.Söderstr., Phytotaxa 112 (1): 21, 2013. Bas.: Isotachis mitteniana Colenso, Trans. & Proc. New Zealand Inst. 21: 69, 1889.

*** Cryptolophocolea mitteniana var. obtusa (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 21, 2013. Bas.: Chiloscyphus mittenianus var. obtusus J.J.Engel, Phytologia 83 (1): 44, 1997 [1998].

*** Cryptolophocolea mitteniana var. symmetrica (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 22, 2013. Bas.: Chiloscyphus mittenianus var. symmetricus J.J.Engel, Phytologia 83 (1): 44, 1997 [1998].

*** Cryptolophocolea pallida (Mitt.) L.Söderstr., Phytotaxa 112 (1): 22, 2013. Bas.: Lophocolea pallida Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 135, 1854.

** Cryptolophocolea pallidovirens (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 20, 2013. Bas.: Jungermannia pallidovirens Hook.f. et Taylor, London J. Bot. 3: 473, 1844.

* Cryptolophocolea proteus (Herzog) L.Söderstr., Phytotaxa 112 (1): 22, 2013. Bas.: Lophocolea proteus Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 164, 1955.

* Cryptolophocolea pycnophylla (Spruce) L.Söderstr., Phytotaxa 112 (1): 22, 2013. Bas.: Lophocolea pycnophylla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 434, 1885.

*** Cryptolophocolea regularis (Steph.) L.Söderstr., Phytotaxa 112 (1): 23, 2013. Bas.: Chiloscyphus regularis Steph., Hedwigia 32 (5): 325, 1893.

*** Cryptolophocolea spinifera (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 20, 2013. Bas.: Jungermannia spinifera Hook.f. et Taylor, London J. Bot. 3: 381, 1844.

* Cryptolophocolea stephanii (Schiffn.) L.Söderstr., Phytotaxa 112 (1): 23, 2013. Bas.: Lophocolea stephanii Schiffn., Hep. Fl. Buitenzorg: 181, 1900. 140

*** Cryptolophocolea subopposita (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 23, 2013. Bas.: Chiloscyphus suboppositus J.J.Engel, Phytologia 83 (1): 45, 1997 [1998].

* Cryptolophocolea thermarum (Schiffn.) L.Söderstr., Phytotaxa 112 (1): 23, 2013. Bas.: Lophocolea thermarum Schiffn., Hep. Fl. Buitenzorg: 180, 1900. 141

*** Cryptolophocolea trialata (Gottsche) L.Söderstr., Phytotaxa 112 (1): 23, 2013. Bas.: Lophocolea trialata Gottsche, Linnaea 28 (5): 552, 1856 [1857].

** Cryptolophocolea tricorata (Hässel) Crand.-Stotl. et Stotler, Phytotaxa 112 (1): 23, 2013. Bas.: Chiloscyphus tricoratus Hässel, Nova Hedwigia 70 (3/4): 456, 2000.

*** Cryptolophocolea tuberculata (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 23, 2013. Bas.: Chiloscyphus tuberculatus J.J.Engel, Phytologia 83 (1): 45, 1997 [1998].

* Cryptolophocolea whittieriana (Inoue et H.A.Mill.) L.Söderstr., Phytotaxa 112 (1): 24, 2013. Bas.: Lophocolea whittieriana Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 143, 1965. 142

*** Deceptifrons J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 54, 2013.

*** Deceptifrons plagiochiloides J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 54, 2013.

*** Evansianthus R.M.Schust. et J.J.Engel, Bryologist 76 (4): 516, 1973.

*** Evansianthus georgiensis (Gottsche) R.M.Schust. et J.J.Engel, Bryologist 76 (4): 518, 1973. Bas.: Lophocolea georgiensis Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 453, 1890.

*** Hepatostolonophora J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 46: 91, 1979. Nom. nov. pro Stolonophora J.J.Engel et R.M.Schust., Fieldiana, Bot. 36 (11): 111, 1975.

*** Hepatostolonophora abnormis (Besch. et C.Massal.) J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 46: 94, 1979. Bas.: Leioscyphus abnormis Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 629, 1886.

** Hepatostolonophora conica (Steph.) Hässel, Bol. Soc. Argent. Bot. 22 (1/4): 123, 1983. Bas.: Plagiochila conica Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 28, 1911.

*** Hepatostolonophora paucistipula (Rodway) J.J.Engel, J. Hattori Bot. Lab. 46: 103, 1979. Bas.: Lophocolea paucistipula Rodway, Pap. & Proc. Roy. Soc. Tasmania 1916: 46, 1917.

*** Hepatostolonophora rotata (Hook.f. et Taylor) J.J.Engel, J. Hattori Bot. Lab. 46: 98, 1979. Bas.: Jungermannia rotata Hook.f. et Taylor, London J. Bot. 3: 560, 1844.

*** Hepatostolonophora rotata var. perssonii (R.M.Schust.) J.J.Engel, J. Hattori Bot. Lab. 46: 103, 1979. Bas.: Calyptrocolea perssonii R.M.Schust., Rev. Bryol. Lichénol. 34 (3/4): 699, 1966 [1967].

*** Heteroscyphus Schiffn., Österr. Bot. Z. 60 (5): 171, 1910 nom. conserv.

** Heteroscyphus acutangulus (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus acutangulus Schiffn., Hep. Fl. Buitenzorg: 200, 1900.

** Heteroscyphus allodontus (Hook.f. et Taylor) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 399, 1984 [1985]. Bas.: Jungermannia allodonta Hook.f. et Taylor, London J. Bot. 3: 382, 1844.

** Heteroscyphus amboinensis (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus endlicherianus var. amboinensis Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 15, 1890.

** Heteroscyphus ammophilus (Colenso) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980. Bas.: Chiloscyphus ammophilus Colenso, Trans. & Proc. New Zealand Inst. 21: 59, 1889.

** Heteroscyphus ammophilus var. obtusifolius J.J.Engel et G.L.Merr., Nova Hedwigia 99 (1/2): 158, 2014.

*** Heteroscyphus argutus (Reinw., Blume et Nees) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Jungermannia arguta Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 206, 1824 [1825].

** Heteroscyphus argutus var. brevidens (Schiffn.) Herzog et Nog., J. Hattori Bot. Lab. 14: 40, 1955. Bas.: Chiloscyphus argutus var. brevidens Schiffn., Hep. Fl. Buitenzorg: 195, 1900.

*** Heteroscyphus aselliformis (Reinw., Blume et Nees) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Jungermannia aselliformis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 412, 1824 [1825].

** Heteroscyphus assurgentifolius J.J.Engel, Nova Hedwigia 99 (1/2): 158, 2014.

** Heteroscyphus baduinus (Nees) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910. Bas.: Jungermannia baduina Nees, Enum. Pl. Crypt. Javae: 26, 1830.

** Heteroscyphus balnetii (Herzog) Grolle, Acta Bot. Fenn. 125: 68, 1984. Bas.: Chiloscyphus balnetii Herzog, Ann. Bryol. 5: 89, 1932.

*** Heteroscyphus billardierei (Schwägr.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Jungermannia billardierei Schwägr., Hist. Musc. Hepat. Prodr.: 19, 1814.

** Heteroscyphus billardierei var. clasmatocoleoides (J.J.Engel et G.L.Merr.) J.J.Engel, Nova Hedwigia 100 (3/4): 565, 2015. Bas.: Heteroscyphus circumdentatus var. clasmatocoleoides J.J.Engel et G.L.Merr., Polish Bot. J. 58 (1): 95, 2013.

** Heteroscyphus brassii (Grolle) Grolle, Acta Bot. Fenn. 125: 68, 1984. Bas.: Chiloscyphus brassii Grolle, J. Hattori Bot. Lab. 31: 2, 1968.

* Heteroscyphus caesius (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus caesius Schiffn., Hep. Fl. Buitenzorg: 207, 1900. 143

* Heteroscyphus caledonicus (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus caledonicus Steph., Bull. Herb. Boissier (sér. 2) 7 (10): 844 (216), 1907. 144

** Heteroscyphus chlorophyllus (Hook.f. et Taylor) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Jungermannia chlorophylla Hook.f. et Taylor, London J. Bot. 3: 562, 1844.

** Heteroscyphus ciliatus (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus ciliatus Steph., Hedwigia 32 (5): 320, 1893.

*** Heteroscyphus coalitus (Hook.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Jungermannia coalita Hook., Musci Exot. 2: tab. 123, 1820.

** Heteroscyphus coalitus var. simplicifolius J.J.Engel, Nova Hedwigia 100 (3/4): 572, 2015.

** Heteroscyphus combinatus (Nees) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Jungermannia combinata Nees, Enum. Pl. Crypt. Javae: 22, 1830.

*** Heteroscyphus conjugatus (Mitt.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 399, 1984 [1985]. Bas.: Chiloscyphus conjugatus Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 227, 1860.

*** Heteroscyphus contortuplicatus (Nees et Mont.) Grolle, J. Hattori Bot. Lab. 55: 504, 1984. Bas.: Jungermannia contortuplicata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 54, 1836.

*** Heteroscyphus cuneistipulus (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus cuneistipulus Steph., Hedwigia 32 (5): 322, 1893.

** Heteroscyphus darjeelingensis A.Srivast. et S.C.Srivast., Indian Geocalyc.: 115, 2002.

** Heteroscyphus deceptifrons J.J.Engel, Nova Hedwigia 99 (1/2): 162, 2014.

*** Heteroscyphus dentammophilus J.J.Engel et G.L.Merr., Polish Bot. J. 58 (1): 96, 2013.

** Heteroscyphus denticulatus (Mitt.) Schiffn., Beih. Bot. Centralbl. 29: 106, 1912. Bas.: Chiloscyphus denticulatus Mitt., Nat. hist. Azores: 320, 1870.

** Heteroscyphus deplanchei (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus deplanchei Steph., Bull. Herb. Boissier (sér. 2) 7 (8): 693 (203), 1907.

** Heteroscyphus diestianus (Sande Lac.) Piippo, Acta Bot. Fenn. 131: 165, 1985. Bas.: Chiloscyphus diestianus Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 296, 1864.

** Heteroscyphus divergenticiliatus (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 501, 1976. Bas.: Lophocolea divergenticiliata Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 38, 1900.

** Heteroscyphus dubius (Gottsche) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus dubius Gottsche, Abh. Naturwiss. Vereins Bremen 7: 346, 1882.

*** Heteroscyphus echinellus (Lindenb. et Gottsche) J.J.Engel et Xiao L.He, Bryologist 113 (1): 155, 2010. Bas.: Lophocolea echinella Lindenb. et Gottsche, Syn. Hepat. 5: 703, 1847.

** Heteroscyphus echinellus var. hyalinus J.J.Engel, Bryologist 113 (1): 161, 2010.

** Heteroscyphus elliottii (Steph.) Pagán, Bryologist 42 (1): 7, 1939. Bas.: Chiloscyphus elliottii Steph., Bull. Herb. Boissier (sér. 2) 8 (1): 55 (231), 1908.

* Heteroscyphus elliottii var. portoricensis Fulford, Mem. New York Bot. Gard. 11 (4): 489, 1976.

* Heteroscyphus falcifolius (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus falcifolius Steph., Bull. Herb. Boissier (sér. 2) 7 (8): 698 (208), 1907.

*** Heteroscyphus fissistipus (Hook.f. et Taylor) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Jungermannia fissistipa Hook.f. et Taylor, London J. Bot. 3: 384, 1844.

** Heteroscyphus fissistipus var. multispinus (E.A.Hodgs. et Allison) J.J.Engel, Nova Hedwigia 99 (1/2): 165, 2014. Bas.: Chiloscyphus multispinus E.A.Hodgs. et Allison, Trans. & Proc. Roy. Soc. New Zealand 73 (1): 37, 1943.

** Heteroscyphus fissistipus var. repandus J.J.Engel, Nova Hedwigia 99 (1/2): 165, 2014.

** Heteroscyphus flaccidus (Mitt.) A.Srivast. et S.C.Srivast., Indian Geocalyc.: 85, 2002. Bas.: Lophocolea flaccida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 99, 1860 [1861].

** Heteroscyphus fleischeri (Steph.) D.G.Long et Rubas., Ceylon J. Sci., Biol. Sci. 43 (1): 26, 2014. Bas.: Chiloscyphus fleischeri Steph., Sp. Hepat. (Stephani) 6: 306, 1922.

* Heteroscyphus fragilicilius (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus fragilicilius Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 210, 1900 [1901].

** Heteroscyphus furcistipulus (E.A.Hodgs.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985]. Bas.: Chiloscyphus furcistipulus E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 79, 1965.

** Heteroscyphus giganteus (Steph.) Hürl., Bauhinia 12 (1/2): 114, 1998. Bas.: Chiloscyphus giganteus Steph., Sp. Hepat. (Stephani) 6: 307, 1922.

** Heteroscyphus graeffei (J.B.Jack et Steph.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 29 (4): 645, 1980. Bas.: Lophocolea graeffei J.B.Jack et Steph., Bot. Centralbl. 60 (4): 101, 1894.

** Heteroscyphus grandiflorus (Steph.) Hürl., Bauhinia 12 (1/2): 114, 1998. Bas.: Chiloscyphus grandiflorus Steph., Sp. Hepat. (Stephani) 6: 307, 1922.

** Heteroscyphus grandistipus (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus grandistipus Steph., Bot. Gaz. 15 (11): 283, 1890.

** Heteroscyphus gunnianus (Mitt.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985]. Bas.: Chiloscyphus gunnianus Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 228, 1860.

** Heteroscyphus hastatus (E.A.Hodgs.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985]. Bas.: Chiloscyphus hastatus E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (11): 181, 1967.

** Heteroscyphus hebridensis (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus hebridensis Steph., Hedwigia 32 (5): 323, 1893.

** Heteroscyphus heterophyllus (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985]. Bas.: Chiloscyphus heterophyllus Steph., Sp. Hepat. (Stephani) 6: 308, 1922.

** Heteroscyphus hyalinus (Steph.) A.Srivast. et S.C.Srivast., Indian Geocalyc.: 118, 2002. Bas.: Lophocolea hyalina Steph., Bull. Soc. Roy. Bot. Belgique 38 (1): 46, 1899.

*** Heteroscyphus integrifolius (Lehm. et Lindenb.) Fulford, Mem. New York Bot. Gard. 11 (4): 495, 1976. Bas.: Jungermannia integrifolia Lehm. et Lindenb., Nov. Stirp. Pug. 6: 32, 1834.

** Heteroscyphus iwatsukii (S.Hatt.) Piippo, Willdenowia 18 (2): 522, 1989. Bas.: Saccogynidium iwatsukii S.Hatt., J. Jap. Bot. 39 (7): 206, 1964.

** Heteroscyphus jackii (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus jackii Steph., Bot. Centralbl. 60 (4): 102, 1894.

*** Heteroscyphus knightii (Steph.) Grolle, J. Hattori Bot. Lab. 61: 251, 1986 [1987]. Bas.: Chiloscyphus knightii Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 129 (245), 1908.

** Heteroscyphus levieri (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus levieri Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 132 (248), 1908.

** Heteroscyphus limosus (Carrington et Pearson) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus limosus Carrington et Pearson, Pap. & Proc. Roy. Soc. Tasmania 1887: 6, 1888.

** Heteroscyphus lingulatus (Colenso) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985]. Bas.: Chiloscyphus lingulatus Colenso, Trans. & Proc. New Zealand Inst. 21: 61, 1889.

** Heteroscyphus lophocoleoides S.Hatt., Bull. Tokyo Sci. Mus. 11: 45, 1944.

*** Heteroscyphus lyallii (Mitt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980. Bas.: Chiloscyphus lyallii Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 140, 1854.

*** Heteroscyphus magellanicus (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985]. Bas.: Chiloscyphus magellanicus Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 140 (256), 1908.

** Heteroscyphus mamillatus Piippo, Nova Hedwigia 56 (3/4): 360, 1993.

** Heteroscyphus marginatus (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 491, 1976. Bas.: Lophocolea marginata Steph., Sp. Hepat. (Stephani) 6: 282, 1922.

*** Heteroscyphus menziesii (Mitt.) J.J.Engel, Polish Bot. J. 58 (1): 99, 2013. Bas.: Chiloscyphus menziesii Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 139, 1854.

** Heteroscyphus merapiensis (Steph.) Piippo, Ann. Bot. Fenn. 30 (3): 200, 1993. Bas.: Chiloscyphus merapiensis Steph., Sp. Hepat. (Stephani) 6: 311, 1922.

** Heteroscyphus miradorensis (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus miradorensis Steph., Bull. Herb. Boissier (sér. 2) 8 (1): 56 (232), 1908.

*** Heteroscyphus mononuculus J.J.Engel, J. Hattori Bot. Lab. 90: 241, 2001.

** Heteroscyphus mononuculus var. ammophilopsis J.J.Engel, Nova Hedwigia 99 (1/2): 166, 2014.

** Heteroscyphus mononuculus var. bilobus J.J.Engel, Nova Hedwigia 99 (1/2): 167, 2014.

** Heteroscyphus montagnei (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 497, 1976. Bas.: Chiloscyphus montagnei Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 141 (257), 1908.

** Heteroscyphus multifidus (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985]. Bas.: Chiloscyphus multifidus Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 132 (248), 1908.

** Heteroscyphus multifidus var. subintegerrimus J.J.Engel, Nova Hedwigia 100 (3/4): 562, 2015.

** Heteroscyphus nadeaudii (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus nadeaudii Steph., Bull. Herb. Boissier (sér. 2) 7 (10): 848 (220), 1907.

** Heteroscyphus orbiculatus A.Srivast. et S.C.Srivast., Indian Geocalyc.: 140, 2002.

** Heteroscyphus palniensis A.Srivast. et S.C.Srivast., Indian Geocalyc.: 130, 2002.

** Heteroscyphus pandei S.C.Srivast. et A.Srivast., Lindbergia 15 (6): 197, 1989 [1991].

*** Heteroscyphus parallelifolius J.J.Engel, Polish Bot. J. 58 (1): 99, 2013.

** Heteroscyphus parvulus (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910. Bas.: Chiloscyphus parvulus Schiffn., Hep. Fl. Buitenzorg: 206, 1900.

** Heteroscyphus parvus A.Srivast. et S.C.Srivast., Indian Geocalyc.: 112, 2002.

** Heteroscyphus perfoliatus (Mont.) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910. Bas.: Lophocolea perfoliata Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 12, 1842.

** Heteroscyphus pertusus (Lehm.) Fulford, Mem. New York Bot. Gard. 11 (4): 505, 1976. Bas.: Chiloscyphus pertusus Lehm., Nov. Stirp. Pug. 10: 7, 1857.

*** Heteroscyphus planiusculus (Hook.f. et Taylor) J.J.Engel, J. Hattori Bot. Lab. 68: 315, 1990. Bas.: Jungermannia planiuscula Hook.f. et Taylor, London J. Bot. 3: 382, 1844.

** Heteroscyphus planus (Mitt.) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910. Bas.: Chiloscyphus planus Mitt., J. Linn. Soc., Bot. 8 (31): 157, 1864 [1865].

*** Heteroscyphus polyblepharis (Spruce) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus polyblepharis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 442, 1885.

*** Heteroscyphus polychaetus (Spruce) Hentschel et Heinrichs, Taxon 56 (4): 1139, 2007. Bas.: Lophocolea polychaeta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 436, 1885.

** Heteroscyphus polycladus (Hook.f. et Lév.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980. Bas.: Jungermannia polyclada Hook.f. et Lév., Choix Pl. Nouv.-Zel.: 8, 1846.

** Heteroscyphus rectangulatus (Herzog) Piippo, Ann. Bot. Fenn. 30 (3): 200, 1993. Bas.: Chiloscyphus rectangulatus Herzog, Ann. Naturhist. Mus. Wien 53 (1): 364, 1942 [1943].

** Heteroscyphus saccogynoides Herzog, J. Hattori Bot. Lab. 14: 40, 1955.

** Heteroscyphus sarawaketanus Piippo, Acta Bot. Fenn. 131: 143, 1985.

*** Heteroscyphus sinuosus (Hook.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Jungermannia sinuosa Hook., Musci Exot. 2: tab. 113, 1820.

** Heteroscyphus spectabilis (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus spectabilis Steph., Hedwigia 30 (5): 205, 1891.

** Heteroscyphus spinifer C.Gao, T.Cao et Y.H.Wu, J. Bryol. 26 (2): 97, 2004.

*** Heteroscyphus splendens (Lehm. et Lindenb.) Grolle, Acta Bot. Fenn. 125: 68, 1984. Bas.: Jungermannia splendens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 22, 1832.

** Heteroscyphus splendidus (E.A.Hodgs.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985]. Bas.: Chiloscyphus splendidus E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (11): 180, 1967.

*** Heteroscyphus stolonifer J.J.Engel, Polish Bot. J. 58 (1): 102, 2013.

*** Heteroscyphus succulentus (Gottsche) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910. Bas.: Chiloscyphus succulentus Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 576, 1853.

** Heteroscyphus supinus (Hook.f. et Taylor) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980. Bas.: Chiloscyphus supinus Hook.f. et Taylor, London J. Bot. 5: 284, 1846.

** Heteroscyphus tener (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus tener Steph., Bull. Herb. Boissier (sér. 2) 7 (8): 695 (205), 1907.

*** Heteroscyphus thraustus (Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 495, 1976. Bas.: Lophocolea thrausta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 437, 1885.

** Heteroscyphus timppae Piippo, Ann. Bot. Fenn. 29 (3): 246, 1992.

*** Heteroscyphus triacanthus (Hook.f. et Lév.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Jungermannia triacantha Hook.f. et Lév., Choix Pl. Nouv.-Zel.: 8, 1846.

** Heteroscyphus triacanthus var. magnistipulatus J.J.Engel, Nova Hedwigia 99 (1/2): 167, 2014.

** Heteroscyphus tridentatus (Sande Lac.) Grolle, Acta Bot. Fenn. 125: 68, 1984. Bas.: Lophocolea tridentata Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 296, 1864.

** Heteroscyphus turgidus (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910. Bas.: Chiloscyphus turgidus Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 212, 1900 [1901].

*** Heteroscyphus valdiviensis (Mont.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus valdiviensis Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 351, 1845.

** Heteroscyphus varians (Steph.) J.J.Engel, J. Hattori Bot. Lab. 68: 315, 1990. Bas.: Lophocolea varians Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 119, 1914.

** Heteroscyphus wettsteinii (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910. Bas.: Chiloscyphus wettsteinii Schiffn., Hep. Fl. Buitenzorg: 202, 1900.

** Heteroscyphus zollingeri (Gottsche) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910. Bas.: Chiloscyphus zollingeri Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 576, 1853.

*** Lamellocolea J.J.Engel, J. Hattori Bot. Lab. 70: 65, 1991.

*** Lamellocolea granditexta (Steph.) J.J.Engel, J. Hattori Bot. Lab. 70: 66, 1991. Bas.: Lophocolea granditexta Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 881 (106), 1906.

*** Lamellocolea integrostia J.J.Engel et Glenny, Bryologist 114 (1): 23, 2011.

*** Leptophyllopsis R.M.Schust., J. Hattori Bot. Lab. 26: 269, 1963.

*** Leptophyllopsis laxa (Mitt.) R.M.Schust. ex Hamlin, Rec. Domin. Mus. 7: 284, 1972. Bas.: Chiloscyphus laxus Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 142, 1854.

*** Leptoscyphopsis R.M.Schust., Phytologia 39 (4): 246, 1978.

*** Leptoscyphopsis paradoxa R.M.Schust., Phytologia 39 (4): 246, 1978.

*** Leptoscyphus Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851. 145

** subg. Anomylia (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 272, 1980. Bas.: Anomylia R.M.Schust., Amer. Midl. Naturalist 62 (1): 51, 1959.

*** Leptoscyphus cuneifolius (Hook.) Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851. Bas.: Jungermannia cuneifolia Hook., Brit. Jungermann.: tab. 64, 1814.

** Leptoscyphus cuneifolius subsp. fragilis (J.B.Jack et Steph.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 28, 1962 [1963]. Bas.: Leioscyphus fragilis J.B.Jack et Steph., Hedwigia 31 (1): 20, 1892.

** subg. Austroleptoscyphus Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 183, 2010.

*** Leptoscyphus antarcticus (C.Massal.) Solari, Cryptog. Bryol. Lichénol. 7 (3): 219, 1986. Bas.: Leioscyphus antarcticus C.Massal., Atti Reale Ist. Veneto Sci. Lett. Arti 87 (2): 229, 1928.

*** Leptoscyphus australis (Gottsche, Lindenb. et Nees) R.M.Schust., J. Hattori Bot. Lab. 26: 270, 1963. Bas.: Chiloscyphus australis Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 189, 1845.

*** Leptoscyphus belmoranus (Steph.) J.J.Engel, J. Hattori Bot. Lab. 74: 33, 1993. Bas.: Lophocolea belmorana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 117, 1914.

*** Leptoscyphus excipulatus (Steph.) J.J.Engel, J. Hattori Bot. Lab. 74: 33, 1993. Bas.: Lophocolea excipulata Steph., Bull. Herb. Boissier (sér. 2) 6 (9): 790 (90), 1906.

*** Leptoscyphus innovatus (E.A.Hodgs.) J.J.Engel, J. Hattori Bot. Lab. 74: 33, 1993. Bas.: Lophocolea innovata E.A.Hodgs., Trans. Roy. Soc. New Zealand 80 (3/4): 347, 1952 [1953].

*** Leptoscyphus longistipulus (Steph.) J.J.Engel, Bryologist 94 (4): 436, 1991. Bas.: Lophocolea longistipula Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 884 (109), 1906.

** subg. Leptoscyphus

** sect. Hexagonistipa Grolle, Nova Acta Leop. (n.ser.) 25 (161): 46, 1962.

*** Leptoscyphus gibbosus (Taylor) Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851. Bas.: Chiloscyphus gibbosus Taylor, London J. Bot. 5: 283, 1846.

*** Leptoscyphus gradsteinii Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 182, 2010.

*** Leptoscyphus hexagonus (Nees) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 47, 1962 [1963]. Bas.: Chiloscyphus hexagonus Nees, Syn. Hepat. 2: 177, 1845.

*** Leptoscyphus jackii (Steph.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 48, 1962 [1963]. Bas.: Leioscyphus jackii Steph., Hedwigia 31 (1): 21, 1892.

*** Leptoscyphus physocalyx (Hampe et Gottsche) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 33, 1858. Bas.: Jungermannia physocalyx Hampe et Gottsche, Linnaea 20 (3): 326, 1847.

*** Leptoscyphus sotiauxii Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 183, 2010.

** sect. Leptoscyphus

*** Leptoscyphus aequatus (Hook.f. et Taylor) Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851. Bas.: Jungermannia aequata Hook.f. et Taylor, London J. Bot. 3: 465, 1844.

*** Leptoscyphus intermedius Grolle, Nova Acta Leop. (n.ser.) 25 (161): 32, 1962 [1963].

*** Leptoscyphus lambinonii Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 179, 2010.

*** Leptoscyphus obcordatus (Spruce) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 33, 1962 [1963]. Bas.: Leioscyphus obcordatus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 446, 1885.

*** Leptoscyphus ovatus (Spruce) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 45, 1962 [1963]. Bas.: Leioscyphus ovatus Spruce, J. Linn. Soc., Bot. 30 (210): 357, 1895.

*** Leptoscyphus porphyrius (Nees) Grolle, Österr. Bot. Z. 117 (1): 3, 1969. Bas.: Chiloscyphus porphyrius Nees, Syn. Hepat. 2: 185, 1845.

*** Leptoscyphus porphyrius subsp. azoricus (H.Buch et Perss.) Vanderp. et Heinrichs, Taxon 59 (1): 181, 2010. Bas.: Mylia azorica H.Buch et Perss., Bryophyt. Azoren Madeira: 7, 1941.

** subg. Physoscyphus Grolle, Nova Acta Leop. (n.ser.) 25 (161): 51, 1962.

** sect. Homaloscyphus Grolle, Nova Acta Leop. (n.ser.) 25 (161): 58, 1962.

*** Leptoscyphus chilensis (De Not.) Hässel, J. Hattori Bot. Lab. 91: 207, 2001. Bas.: Lophocolea chilensis De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 222, 1857.

* Leptoscyphus difficilis (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 534, 1976. Bas.: Chiloscyphus difficilis Steph., Biblioth. Bot. 87 (2): 222, 1916. 146

*** Leptoscyphus diversifolius (Gottsche) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 58, 1962 [1963]. Bas.: Lophocolea diversifolia Gottsche, Syn. Hepat. 2: 166, 1845.

*** Leptoscyphus expansus (Lehm.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 60, 1962 [1963]. Bas.: Jungermannia expansa Lehm., Linnaea 4: 361, 1829.

*** Leptoscyphus hedbergii (S.W.Arnell) R.M.Schust., Amer. Midl. Naturalist 62 (1): 13, 1959. Bas.: Mylia hedbergii S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 547, 1956.

* Leptoscyphus huidobroanus (Mont.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 33, 1858. Bas.: Chiloscyphus huidobroanus Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 352, 1845. 147

*** Leptoscyphus huonicus Piippo, Acta Bot. Fenn. 131: 152, 1985.

*** Leptoscyphus magellanicus (Gola) Hässel, J. Hattori Bot. Lab. 91: 214, 2001. Bas.: Lophozia magellanica Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 29 (1/4): 165, 1922 [1923].

** sect. Physoscyphus Grolle, Nova Acta Leop. (n.ser.) 25 (161): 51, 1962.

*** Leptoscyphus amphibolius (Nees) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 54, 1962 [1963]. Bas.: Jungermannia amphibolia Nees, Fl. Bras. (Martius) 1 (1): 334, 1833.

*** Leptoscyphus infuscatus (Mitt.) E.W.Jones ex Grolle, Nova Acta Leop. (n.ser.) 25 (161): 52, 1962 [1963]. Bas.: Leioscyphus infuscatus Mitt., J. Linn. Soc., Bot. 22 (146): 321, 1886.

** subg. Spinoscyphus Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 183, 2010.

*** Leptoscyphus cleefii Fulford, Mem. New York Bot. Gard. 11 (4): 534, 1976.

*** Leptoscyphus spectabilis (Steph.) Grolle, J. Bryol. 11 (2): 328, 1980 [1981]. Bas.: Lophocolea spectabilis Steph., Bull. Herb. Boissier (sér. 2) 7 (4): 302 (166), 1907.

Incertae sedis

*** Leptoscyphus autoicus (J.J.Engel et Gradst.) Vanderp. et Gradst., J. Bryol. 34 (4): 252, 2012. Bas.: Physotheca autoica J.J.Engel et Gradst., Taxon 52 (4): 764, 2003.

** Leptoscyphus beckettianus (Steph.) R.M.Schust. ex J.J.Engel, Nova Hedwigia 93 (3/4): 402, 2011. Bas.: Chiloscyphus beckettianus Steph., Bull. Herb. Boissier (sér. 2) 8 (1): 59 (235), 1908.

** Leptoscyphus compactus (Colenso) J.J.Engel, Nova Hedwigia 100 (3/4): 579, 2015. Bas.: Chiloscyphus compactus Colenso, Trans. & Proc. New Zealand Inst. 21: 63, 1889.

** Leptoscyphus erraticus (W.Martin et E.A.Hodgs.) J.J.Engel, Nova Hedwigia 99 (1/2): 168, 2014. Bas.: Chiloscyphus erraticus W.Martin et E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 78 (4): 497, 1950.

*** Leptoscyphus horizontalis (Hook.) Kühnem., Revista Centro Estud. Doct. Ci. Nat. 1: 176, 1937. Bas.: Jungermannia horizontalis Hook., Musci Exot. 1: tab. 96, 1818.

** Leptoscyphus normalis (Steph.) J.J.Engel, Nova Hedwigia 100 (3/4): 579, 2015. Bas.: Lophocolea normalis Steph., Sp. Hepat. (Stephani) 6: 285, 1922.

** Leptoscyphus physanthus (Hook.f. et Taylor) J.J.Engel, Nova Hedwigia 99 (1/2): 168, 2014. Bas.: Jungermannia physantha Hook.f. et Taylor, London J. Bot. 3: 561, 1844.

** Leptoscyphus subemarginatus (Hook.f. et Taylor) J.J.Engel, Bryologist 94 (4): 436, 1991. Bas.: Lophocolea subemarginata Hook.f. et Taylor, London J. Bot. 5: 367, 1846.

*** Leptoscyphus trapezoïdes (Mont.) L.Söderstr., Phytotaxa 112 (1): 27, 2013. Bas.: Lophocolea trapezoïdes Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 251, 1843.

*** Lophocolea (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835. Bas.: Jungermannia sect. Lophocolea Dumort., Syll. Jungerm. Europ.: 59, 1831.

*** Lophocolea aberrans Lindenb. et Gottsche, Syn. Hepat. 5: 696, 1847.

*** Lophocolea aequifolia Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 55, 1836.

* Lophocolea angustistipula Steph., Sp. Hepat. (Stephani) 6: 260, 1922.

*** Lophocolea anisoloba (J.J.Engel et Glenny) L.Söderstr., Phytotaxa 112 (1): 25, 2013. Bas.: Chiloscyphus anisolobus J.J.Engel et Glenny, Bryologist 111 (1): 118, 2008.

** Lophocolea anomala Steph., Sp. Hepat. (Stephani) 6: 300, 1922.

** Lophocolea anomoda (Mont.) Steph., Hedwigia 32 (5): 327, 1893. Bas.: Chiloscyphus anomodus Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 352, 1845.

** Lophocolea apalachicola R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 195, 1980.

*** Lophocolea aperticaulis (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 25, 2013. Bas.: Chiloscyphus aperticaulis J.J.Engel, J. Hattori Bot. Lab. 95: 229, 2004.

** Lophocolea aphelophylla (Hässel) Váňa, Phytotaxa 112 (1): 25, 2013. Bas.: Chiloscyphus aphelophyllus Hässel, J. Hattori Bot. Lab. 98: 123, 2005.

** Lophocolea apophylla (Hässel) Váňa, Phytotaxa 112 (1): 25, 2013. Bas.: Chiloscyphus apophyllus Hässel, J. Hattori Bot. Lab. 98: 126, 2005.

*** Lophocolea appalachiana R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 208, 1980.

** Lophocolea ascensionis Steph., Sp. Hepat. (Stephani) 6: 261, 1922.

** Lophocolea asperrima Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 949 (129), 1906.

** Lophocolea atra Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 29 (1/4): 167, 1922 [1923].

*** Lophocolea attenuata Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 34, 1900.

*** Lophocolea australis Gottsche, Linnaea 28 (5): 553, 1856 [1857].

** Lophocolea autoica Steph., Sp. Hepat. (Stephani) 6: 262, 1922.

** Lophocolea baldwinii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 950 (130), 1906.

** Lophocolea bartlettii H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 506, 1963.

** Lophocolea bewsii (Sim) Grolle, Trans. Brit. Bryol. Soc. 3 (4): 588, 1959. Bas.: Leptoscyphus bewsii Sim, Trans. Roy. Soc. South Africa 15 (1): 103, 1926.

** Lophocolea bicuspidata Steph., Sp. Hepat. (Stephani) 6: 263, 1922.

*** Lophocolea bidentata (L.) Dumort., Recueil Observ. Jungerm.: 17, 1835. Bas.: Jungermannia bidentata L., Sp. Pl. 1: 1132, 1753. 148

** Lophocolea bifidistipula Steph., Sp. Hepat. (Stephani) 6: 264, 1922.

*** Lophocolea bispinosa (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 162, 1845. Bas.: Jungermannia bispinosa Hook.f. et Taylor, London J. Bot. 3: 378, 1844.

** Lophocolea bootanensis Steph., Sp. Hepat. (Stephani) 6: 265, 1922.

** Lophocolea boulyana Steph., Sp. Hepat. (Stephani) 6: 264, 1922.

** Lophocolea bowiena Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 117, 1914.

*** Lophocolea brookwoodiana Paton et Sheahan, J. Bryol. 28 (3): 163, 2006.

** Lophocolea caespitans Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 949 (129), 1906.

*** Lophocolea calcarea Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 884 (109), 1906.

** Lophocolea caledonica Steph., Sp. Hepat. (Stephani) 6: 267, 1922.

*** Lophocolea canaliculata (Gottsche, Lindenb. et Nees) Steph., Bull. Herb. Boissier (sér. 2) 6 (9): 786 (86), 1906. Bas.: Chiloscyphus canaliculatus Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 710, 1847.

*** Lophocolea canaliculata var. concava (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013. Bas.: Chiloscyphus canaliculatus var. concavus J.J.Engel, Fieldiana, Bot. (n.ser.) 48: 107, 2010.

** Lophocolea cervicornis Steph., Biblioth. Bot. 87 (2): 219, 1916.

** Lophocolea ciliifera Steph., Bull. Herb. Boissier (sér. 2) 6 (8): 660 (76), 1906.

* Lophocolea coadunata (Sw.) Mont., Voy. Amér. Mérid., Bot. 7 (1): 76, 1839. Bas.: Jungermannia coadunata Sw., Fl. Ind. Occid. 3: 1850, 1806. 149

** Lophocolea concreta Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 350, 1845.

** Lophocolea convexula Mitt., Fl. vit.: 405, 1871 [1873].

** Lophocolea corticola Steph., Sp. Hepat. (Stephani) 6: 268, 1922.

*** Lophocolea decurrens Herzog, Trans. & Proc. Roy. Soc. New Zealand 65 (3): 352, 1936.

* Lophocolea deningeri Herzog, Beih. Bot. Centralbl. 38 (2): 321, 1921.

** Lophocolea dentiflora Steph., Bull. Herb. Boissier (sér. 2) 6 (7): 550 (64), 1906.

** Lophocolea difformis Nees, Syn. Hepat. 2: 166, 1845.

** Lophocolea discedens (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 2: 167, 1845. Bas.: Jungermannia discedens Lehm. et Lindenb., Nov. Stirp. Pug. 5: 3, 1833.

* Lophocolea dusenii Steph., Cat. Afr. Pl. (Hiern) 2 (2): 314, 1901.

*** Lophocolea erosa (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013. Bas.: Chiloscyphus erosus J.J.Engel, Phytologia 83 (1): 43, 1997 [1998].

*** Lophocolea excisifolia Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 118, 1914.

*** Lophocolea fertilis (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013. Bas.: Chiloscyphus fertilis J.J.Engel, Phytologia 83 (1): 43, 1997 [1998].

** Lophocolea flavicans Steph., Sp. Hepat. (Stephani) 6: 300, 1922.

*** Lophocolea floribunda Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 886 (111), 1906.

** Lophocolea foliicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 428, 1885.

** Lophocolea fragillima Steph., Sp. Hepat. (Stephani) 6: 273, 1922.

*** Lophocolea fragmentissima R.M.Schust., Phytologia 39 (4): 245, 1978.

*** Lophocolea fragrans (Moris et De Not.) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 166, 1845. Bas.: Jungermannia fragrans Moris et De Not., Fl. Caprariae: 177, 1839.

** Lophocolea glaziovii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 961 (141), 1906.

** Lophocolea gollanii (Steph.) Váňa, Phytotaxa 112 (1): 26, 2013. Bas.: Chiloscyphus gollanii Steph., Bull. Herb. Boissier (sér. 2) 7 (10): 837 (209), 1907.

** Lophocolea granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 126, 1864.

** Lophocolea griffithiana Steph., Sp. Hepat. (Stephani) 6: 274, 1922.

** Lophocolea hahnii Steph., Bull. Herb. Boissier (sér. 2) 6 (8): 660 (76), 1906.

** Lophocolea haskarliana Gottsche, Syn. Hepat. 2: 153, 1845.

*** Lophocolea hattorii (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013. Bas.: Chiloscyphus hattorii J.J.Engel, J. Hattori Bot. Lab. 74: 29, 1993.

** Lophocolea hawaica Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 945 (125), 1906.

** Lophocolea heterodonta Steph., Sp. Hepat. (Stephani) 6: 275, 1922.

** Lophocolea heteromorpha Steph., Sp. Hepat. (Stephani) 6: 275, 1922.

*** Lophocolea heterophylla (Schrad.) Dumort., Recueil Observ. Jungerm.: 17, 1835. Bas.: Jungermannia heterophylla Schrad., J. Bot. (Schrader) 5: 66, 1802 [1803].

** Lophocolea heterophylla subsp. cladogyna R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 223, 1980.

** Lophocolea horikawana S.Hatt., Bull. Tokyo Sci. Mus. 11: 50, 1944.

** Lophocolea howeana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 118, 1914.

* Lophocolea humifusa (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 695, 1847. Bas.: Jungermannia humifusa Hook.f. et Taylor, London J. Bot. 3: 472, 1844.

** Lophocolea humistrata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 701, 1847. Bas.: Jungermannia humistrata Hook.f. et Taylor, London J. Bot. 4: 82, 1845.

** Lophocolea itoana Inoue, J. Jap. Bot. 31 (11): 340, 1956.

** Lophocolea javanica Schiffn., Hep. Fl. Buitenzorg: 178, 1900.

** Lophocolea koponenii (Piippo) Váňa, Phytotaxa 112 (1): 26, 2013. Bas.: Chiloscyphus koponenii Piippo, Ann. Bot. Fenn. 35 (1): 55, 1998.

** Lophocolea kurzii Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 296, 1864.

* Lophocolea kurzii var. siamensis N.Kitag., Acta Phytotax. Geobot. 30 (1/3): 33, 1979.

** Lophocolea laceristipula Steph., Sp. Hepat. (Stephani) 6: 281, 1922.

** Lophocolea latistipula Steph., Sp. Hepat. (Stephani) 6: 281, 1922.

*** Lophocolea lauterbachii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 938 (118), 1906.

* Lophocolea laxissima Herzog, Ann. Bryol. 5: 77, 1932.

** Lophocolea ledermannii Steph., Sp. Hepat. (Stephani) 6: 300, 1922.

*** Lophocolea lenta (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 162, 1845. Bas.: Jungermannia lenta Hook.f. et Taylor, London J. Bot. 3: 379, 1844.

*** Lophocolea leptantha (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 694, 1847. Bas.: Jungermannia leptantha Hook.f. et Taylor, London J. Bot. 3: 471, 1844.

*** Lophocolea liebmanniana Gottsche, Mexik. Leverm.: 113, 1863.

** Lophocolea lindmannii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 960 (140), 1906.

*** Lophocolea longiciliata Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 96, 1952.

** Lophocolea lucida (Spreng.) Mont., Voy. Amér. Mérid., Bot. 7 (2): 78, 1839. Bas.: Jungermannia lucida Spreng. Nov. Stirp. Pug. 5: 2, 1833.

** Lophocolea madagascariensis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 344, 1882.

** Lophocolea magna (Udar et V.Nath) Váňa, Phytotaxa 183 (4): 291, 2014. Bas.: Cephaloziella magna Udar et V.Nath, Geophytology 6 (1): 105, 1976.

*** Lophocolea mediinfrons (J.J.Engel et Braggins) L.Söderstr., Phytotaxa 112 (1): 26, 2013. Bas.: Chiloscyphus mediinfrons J.J.Engel et Braggins, Fieldiana, Bot. (n.ser.) 48: 119, 2010.

** Lophocolea micronesica Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 4, 1968.

* Lophocolea microstipula Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 43, 1900. 150

*** Lophocolea minor Nees, Naturgesch. Eur. Leberm. 2: 330, 1836.

** Lophocolea minutistipula Steph., Sp. Hepat. (Stephani) 6: 283, 1922.

* Lophocolea mollis (Nees) Nees, Syn. Hepat. 2: 158, 1845. Bas.: Jungermannia mollis Nees, Enum. Pl. Crypt. Javae: 24, 1830. 151

* Lophocolea morobeana Piippo, Acta Bot. Fenn. 131: 160, 1985. 152

** Lophocolea muhavurensis (S.W.Arnell) S.W.Arnell ex Pócs, Acta Bot. Acad. Sci. Hung. 25 (3/4): 227, 1979 [1980]. Bas.: Chiloscyphus muhavurensis S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 526, 1956.

*** Lophocolea muricata (Lehm.) Nees, Syn. Hepat. 2: 169, 1845. Bas.: Jungermannia muricata Lehm., Linnaea 4: 363, 1829.

* Lophocolea muricata var. major Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 10, 1893.

** Lophocolea nakajimae S.Hatt. et Inoue, J. Hattori Bot. Lab. 21: 221, 1959.

*** Lophocolea novae-zeelandiae (Lehm. et Lindenb.) Nees, Syn. Hepat. 2: 168, 1845. Bas.: Jungermannia novae-zeelandiae Lehm. et Lindenb., Nov. Stirp. Pug. 6: 33, 1834.

*** Lophocolea novae-zeelandiae var. meridionalis (Steph.) L.Söderstr., Phytotaxa 112 (1): 26, 2013. Bas.: Lophocolea meridionalis Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 888 (113), 1906.

** Lophocolea orbigniana Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 55, 1836.

** Lophocolea papulimarginata H.A.Mill., Phytologia 47 (4): 323, 1981. Nom. nov. pro Lophocolea papulosa Steph., Sp. Hepat. (Stephani) 6: 286, 1922, nom. illeg.

** Lophocolea parca (Gottsche) Fulford et Sharp, Mem. New York Bot. Gard. 63: 19, 1990. Bas.: Jungermannia parca Gottsche, Mexik. Leverm.: 94, 1863.

** Lophocolea parva Steph., Sp. Hepat. (Stephani) 6: 287, 1922.

*** Lophocolea parvispinea (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013. Bas.: Chiloscyphus parvispineus J.J.Engel, Phytologia 83 (1): 44, 1997 [1998].

** Lophocolea parvistipula Steph., Sp. Hepat. (Stephani) 6: 287, 1922.

*** Lophocolea patulistipa Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 50, 1911.

*** Lophocolea perpusilla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 163, 1845. Bas.: Jungermannia perpusilla Hook.f. et Taylor, London J. Bot. 3: 380, 1844.

** Lophocolea piacenzai (Gola) Váňa, Phytotaxa 112 (1): 26, 2013. Bas.: Lophozia piacenzai Gola, Atti Reale Accad. Sci. Torino, Cl. Sci. Fis. Mat. Nat. 49: 759, 1914.

** Lophocolea pilistipula Steph., Sp. Hepat. (Stephani) 6: 288, 1922.

* Lophocolea pinnatistipula Steph., Biblioth. Bot. 87 (2): 220, 1916.

** Lophocolea platensis C.Massal., Atti Accad. Sci. Med. Nat. Ferrara 80 (3/4): 12, 1906.

** Lophocolea purpurea Steph., Sp. Hepat. (Stephani) 6: 289, 1922.

** Lophocolea pusilla Steph., Sp. Hepat. (Stephani) 6: 290, 1922.

** Lophocolea randii S.W.Arnell, Svensk Bot. Tidskr. 47 (3): 420, 1953.

** Lophocolea rara Steph., Sp. Hepat. (Stephani) 6: 290, 1922.

** Lophocolea rectangularis Herzog, Rev. Bryol. Lichénol. 23 (1/2): 43, 1954.

** Lophocolea rectangulata Mitt., Fl. vit.: 404, 1871 [1873]. 153

*** Lophocolea rupicola Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 874 (99), 1906.

*** Lophocolea sabuletorum (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 697, 1847. Bas.: Jungermannia sabuletorum Hook.f. et Taylor, London J. Bot. 3: 469, 1844.

** Lophocolea salacensis Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 943 (123), 1906.

** Lophocolea savesiana Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 942 (122), 1906.

*** Lophocolea semiteres (Lehm.) Mitt., J. Linn. Soc., Bot. 16 (91): 188, 1877. Bas.: Jungermannia semiteres Lehm., Linnaea 4: 363, 1829. 154

*** Lophocolea semiteres var. retusa (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 27, 2013. Bas.: Chiloscyphus semiteres var. retusus J.J.Engel, Phytologia 83 (1): 44, 1997 [1998].

** Lophocolea serrata Mitt., St. Helena: 368, 1875.

** Lophocolea siamensis Steph., Sp. Hepat. (Stephani) 6: 293, 1922.

** Lophocolea sikkimensis (Steph.) Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 164, 1958 [1959]. Bas.: Herpocladium sikkimense Steph., Sp. Hepat. (Stephani) 6: 349, 1922.

* Lophocolea silvestris Gottsche, Abh. Naturwiss. Vereins Bremen 7: 345, 1882.

** Lophocolea steetziae De Not., Epat. Borneo: 20, 1874.

*** Lophocolea striatella (C.Massal.) Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 13, 1890. Bas.: Chiloscyphus striatellus C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 232, 1885.

** Lophocolea subbidentata Herzog, Rev. Bryol. Lichénol. 23 (1/2): 43, 1954.

** Lophocolea subcostata Steph., Sp. Hepat. (Stephani) 6: 295, 1922.

*** Lophocolea subporosa Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 137, 1854.

** Lophocolea subporosa var. inflexifolia (Steph.) L.Söderstr., Phytotaxa 112 (1): 27, 2013. Bas.: Lophocolea inflexifolia Steph., Sp. Hepat. (Stephani) 6: 278, 1922.

** Lophocolea subviridis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 699, 1847. Bas.: Jungermannia subviridis Hook.f. et Taylor, London J. Bot. 3: 473, 1844.

** Lophocolea sumatrana Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 195, 1900 [1901].

*** Lophocolea sylvatica Mitt., Rep. Challenger, Bot. 1 (3, 1): 84, 1884.

** Lophocolea tenera Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 79, 1876 [1877].

** Lophocolea tenerrima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 439, 1885.

** Lophocolea teptepensis Piippo, Acta Bot. Fenn. 131: 163, 1985.

*** Lophocolea textilis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 696, 1847. Bas.: Jungermannia textilis Hook.f. et Taylor, London J. Bot. 3: 468, 1844.

** Lophocolea textiloidea J.J.Engel, Phytologia 41 (5): 311, 1979. Nom. nov. pro Chiloscyphus lucidus Mitt., J. Linn. Soc., Bot. 15 (82): 64, 1876, nom. illeg.

*** Lophocolea trichocoleoides (Glenny, J.J.Engel et He-Nygrén) L.Söderstr., Phytotaxa 112 (1): 27, 2013. Bas.: Chiloscyphus trichocoleoides Glenny, J.J.Engel et He-Nygrén, J. Bryol. 31 (2): 100, 2009.

** Lophocolea tricuspidata Herzog, Rev. Bryol. Lichénol. 11 (1): 17, 1938 [1939].

*** Lophocolea tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 19, 1958.

* Lophocolea undulata Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 351, 1845.

*** Lophocolea villosa Mitt., Sp. Hepat. (Stephani) 6: 299, 1922.

** Lophocolea wacei (S.W.Arnell ex J.J.Engel et Váňa) Váňa et L.Söderstr., Phytotaxa 112 (1): 27, 2013. Bas.: Chiloscyphus wacei S.W.Arnell ex J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 48, 2013.

*** Lophocolea wambana Piippo, Acta Bot. Fenn. 131: 163, 1985.

** Lophocolea werthii (J.J.Engel et R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 27, 2013. Bas.: Chiloscyphus werthii J.J.Engel et R.M.Schust., Nova Hedwigia 39: 425, 1984 [1985].

** Lophocolea widgrenii Steph., Bull. Herb. Boissier (sér. 2) 7 (1): 66 (154), 1907. Nom. nov. pro Lophocolea pallida Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 80, 1876 [1877], nom. illeg.

** Otoscyphus J.J.Engel, Bardat et Thouvenot, Cryptog. Bryol. 33 (3): 280, 2012.

*** Otoscyphus crassicaulis (Steph.) J.J.Engel, Bardat et Thouvenot, Cryptog. Bryol. 33 (3): 280, 2012. Bas.: Lophocolea crassicaulis Steph., Sp. Hepat. (Stephani) 6: 268, 1922.

*** Pachyglossa Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 150, 1958 [1959].

*** Pachyglossa austrigena (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 24, 2013. Bas.: Jungermannia austrigena Hook.f. et Taylor, London J. Bot. 3: 466, 1844.

*** Pachyglossa austrigena subsp. okaritana (Steph.) L.Söderstr., Phytotaxa 112 (1): 24, 2013. Bas.: Lophocolea okaritana Steph., Bull. Herb. Boissier (sér. 2) 6 (9): 785 (85), 1906.

*** Pachyglossa boveana (C.Massal.) L.Söderstr., Phytotaxa 112 (1): 24, 2013. Bas.: Lophocolea boveana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 225, 1885.

** Pachyglossa dissitifolia Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 155, 1958 [1959].

* Pachyglossa exilis (Herzog et Grolle) Hässel et Solari, Transecta botánica de la Patagonia austral: 324, 1985. Bas.: Pachyglossa spegazziniana var. exilis Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 159, 1958 [1959]. 155

** Pachyglossa fissa (Mitt.) Herzog et Grolle, Rev. Bryol. Lichénol. 28 (3/4): 346, 1959 [1960]. Bas.: Herpocladium fissum Mitt., J. Linn. Soc., Bot. 15 (82): 69, 1876.

*** Pachyglossa gottscheoides (Besch. et C.Massal.) L.Söderstr., Phytotaxa 112 (1): 25, 2013. Bas.: Lophocolea gottscheoides Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 631, 1886.

** Pachyglossa grolleana Váňa, Cryptog. Bryol. 26 (1): 86, 2005.

** Pachyglossa otiphylla (Hook.f. et Taylor) Váňa, Phytotaxa 112 (1): 25, 2013. Bas.: Jungermannia otiphylla Hook.f. et Taylor, London J. Bot. 3: 466, 1844.

** Pachyglossa spegazziniana (C.Massal.) Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 159, 1958 [1959]. Bas.: Lophocolea spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 225, 1885.

** Pachyglossa tenacifolia (Hook.f. et Taylor) Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 153, 1958 [1959]. Bas.: Jungermannia tenacifolia Hook.f. et Taylor, Bot. Antarct. Voy. I (Fl. Antarct. 1): 152, 1845.

** Perdusenia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 7 (2): 11, 1989.

** Perdusenia rheophila Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 7 (2): 11, 1989.

** Pigafettoa C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 237, 1885.

** Pigafettoa crenulata C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 237, 1885.

** Platycaulis R.M.Schust., Phytologia 39 (4): 245, 1978.

*** Platycaulis renifolius R.M.Schust., Phytologia 39 (4): 245, 1978.

*** Stolonivector J.J.Engel, J. Hattori Bot. Lab. 69: 80, 1991.

** Stolonivector clasmatocoleoides J.J.Engel, Nova Hedwigia 88 (3/4): 339, 2009.

*** Stolonivector fiordlandiae (E.A.Hodgs.) J.J.Engel, J. Hattori Bot. Lab. 69: 82, 1991. Bas.: Lophocolea fiordlandiae E.A.Hodgs., Trans. Roy. Soc. New Zealand 80 (3/4): 340, 1952 [1953].

** Stolonivector fiordlandiae var. nodulosus J.J.Engel, Nova Hedwigia 93 (3/4): 403, 2011.

*** Stolonivector gremmenii (Váňa) Váňa, Phytotaxa 112 (1): 28, 2013. Bas.: Chiloscyphus gremmenii Váňa, Cryptog. Bryol. 26 (1): 81, 2005.

** Stolonivector obtusilobus J.J.Engel, Nova Hedwigia 88 (3/4): 337, 2009.

** Stolonivector waipouensis J.J.Engel, J. Hattori Bot. Lab. 93: 70, 2003.

** Xenocephalozia R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965.

** Xenocephalozia navicularis (Steph.) R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965. Bas.: Lophocolea navicularis Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 43, 1900.

*** Mastigophoraceae R.M.Schust.

** Dendromastigophora R.M.Schust., Mem. New York Bot. Gard. 45: 738, 1987.

*** Dendromastigophora flagellifera (Hook.) R.M.Schust., Mem. New York Bot. Gard. 45: 738, 1987. Bas.: Jungermannia flagellifera Hook., Musci Exot. 1: tab. 59, 1818.

*** Mastigophora Nees, Naturgesch. Eur. Leberm. 3: 89, 1838 nom. conserv. 156

* Mastigophora appendiculata Steph., Sp. Hepat. (Stephani) 6: 368, 1922.

* Mastigophora attenuata (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877. Bas.: Lepidozia attenuata Taylor, London J. Bot. 5: 369, 1846.

** Mastigophora caledonica Steph., Rev. Bryol. 35 (2): 31, 1908.

*** Mastigophora diclados (Brid. ex F.Weber) Nees, Naturgesch. Eur. Leberm. 3: 18, 1838. Bas.: Jungermannia diclados Brid. ex F.Weber, Hist. Musc. Hepat. Prodr.: 56, 1815.

** Mastigophora diclados var. borneensis (De Not.) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 251, 1893. Bas.: Sendtnera diclados var. borneensis De Not., Epat. Borneo: 42, 1874.

** Mastigophora diclados var. ramentifissa Herzog, Trans. Brit. Bryol. Soc. 1 (4): 315, 1950.

* Mastigophora diclados var. villosa Herzog, Ann. Bryol. 5: 82, 1932.

* Mastigophora guineensis Steph., Sp. Hepat. (Stephani) 6: 369, 1923.

* Mastigophora humillima (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877. Bas.: Lepidozia humillima Taylor, London J. Bot. 5: 369, 1846.

* Mastigophora pyramidana Steph., Sp. Hepat. (Stephani) 6: 369, 1923.

** Mastigophora sepikiana Piippo, Ann. Bot. Fenn. 23 (1): 2, 1986.

** Mastigophora tuberculata D.H.Mill. et H.A.Mill., J. Hattori Bot. Lab. 75: 181, 1994.

* Mastigophora valida Steph., Sp. Hepat. (Stephani) 6: 369, 1923.

** Mastigophora viridula (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877. Bas.: Jungermannia viridula Nees, Flora 6 (2): 30, 1823.

*** Mastigophora woodsii (Hook.) Nees, Naturgesch. Eur. Leberm. 3: 95, 1838. Bas.: Jungermannia woodsii Hook., Brit. Jungermann.: tab. 66, 1814.

*** Plagiochilaceae Müll.Frib.

by L. Söderström

The placement of Pedinophyllopsis in Plagiochilaceae follows. The inclusion of Pseudolophocolea in the family follows. The subgeneric division of Plagiochila follows the review by.

** Acrochila R.M.Schust., J. Hattori Bot. Lab. 26: 285, 1963.

*** Acrochila biserialis (Lehm. et Lindenb.) Grolle, J. Jap. Bot. 39 (8): 236, 1964. Bas.: Plagiochila biserialis Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 5: 126, 1843.

** Acrochila caledonica (Steph.) Inoue, J. Jap. Bot. 42 (6): 182, 1967. Bas.: Plagiochila caledonica Steph., Rev. Bryol. 35 (2): 32, 1908.

** Chiastocaulon Carl, Flora 126: 58, 1931.

*** Chiastocaulon dendroides (Nees) Carl, Flora 126: 59, 1931. Bas.: Jungermannia dendroides Nees, Enum. Pl. Crypt. Javae: 77, 1830.

** Dinckleria Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 421, 1877.

*** Dinckleria fruticella (Hook.f. et Taylor) J.J.Engel et Heinrichs, Cryptog. Bryol. 29 (2): 194, 2008. Bas.: Jungermannia fruticella Hook.f. et Taylor, London J. Bot. 3: 565, 1844.

*** Dinckleria pleurata (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 421, 1877. Bas.: Jungermannia pleurata Hook.f. et Taylor, London J. Bot. 3: 372, 1844.

** Pedinophyllopsis R.M.Schust. et Inoue, Phytologia 47 (4): 311, 1981.

** Pedinophyllopsis abdita (Sull.) R.M.Schust. et Inoue, Phytologia 47 (4): 311, 1981. Bas.: Plagiochila abdita Sull., Hooker’s J. Bot. Kew Gard. Misc. 2: 317, 1850.

** Pedinophyllum Lindb. ex Nordst., Bot. Not. 1874: 156, 1874.

*** Pedinophyllum autoicum (Steph.) Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 9 (4): 575, 1966. Bas.: Plagiochila autoica Steph., Sp. Hepat. (Stephani) 6: 126, 1917.

*** Pedinophyllum interruptum (Nees) Kaal., Nyt Mag. Naturvidensk. 33 (1): 190, 1893. Bas.: Jungermannia interrupta Nees, Naturgesch. Eur. Leberm. 1: 165, 1833.

*** Pedinophyllum monoicum (Steph.) Grolle, Nova Hedwigia 2: 287, 1960. Bas.: Plagiochila monoica Steph., Bull. Herb. Boissier (sér. 2) 3 (4): 331 (315), 1903.

** Pedinophyllum truncatum (Steph.) Inoue, J. Hattori Bot. Lab. 23: 35, 1960 [1961]. Bas.: Clasmatocolea truncata Steph., Bull. Herb. Boissier 5 (2): 87, 1897.

*** Plagiochila (Dumort.) Dumort., Recueil Observ. Jungerm.: 14, 1835 nom. conserv. Bas.: Radula sect. Plagiochila Dumort., Syll. Jungerm. Europ.: 42, 1831.

*** Plagiochila heteromalla (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 83, 1840. Bas.: Jungermannia heteromalla Lehm. et Lindenb., Nov. Stirp. Pug. 6: 62, 1834.

*** sect. Adianthoideae Lindenb., Monogr. hep. gen. Plagiochilae: xx, 1844.

*** Plagiochila adianthoides (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 77, 1840. Bas.: Jungermannia adianthoides Sw., Prodr. (Swartz): 142, 1788.

* Plagiochila adianthoides var. aspergillifera Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 474, 1885.

*** Plagiochila cristata (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 1: 33, 1839. Bas.: Jungermannia cristata Sw., Prodr. (Swartz): 143, 1788.

*** Plagiochila grandicrista Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 931 (581), 1905.

*** Plagiochila herminieri Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 748 (547), 1905.

*** sect. Africanae Heinrichs, Taxon 54 (2): 319, 2005.

*** Plagiochila barteri Mitt., J. Linn. Soc., Bot. 22 (146): 320, 1886.

** Plagiochila barteri var. valida (Steph.) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 73, 1981. Bas.: Plagiochila valida Steph., Bull. Herb. Boissier (sér. 2) 4 (6): 587 (438), 1904.

*** Plagiochila colorans Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 116, 1911.

*** sect. Arrectae Carl, Ann. Bryol., Suppl. 2: 52, 1931.

* Plagiochila arnelliana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 861 (230), 1902. 157

*** Plagiochila badia Mitt., Rep. Challenger, Bot. 1 (3, 1): 84, 1884.

*** Plagiochila bidens Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 322, 1857.

*** Plagiochila bifaria (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 5: 127, 1843. Bas.: Jungermannia bifaria Sw., Prodr. (Swartz): 145, 1788.

** Plagiochila bifaria var. rosea (R.M.Schust.) Heinrichs, Org. Divers. Evol. 4 (1/2): 112, 2004. Bas.: Rhodoplagiochila rosea R.M.Schust., Phytologia 39 (4): 247, 1978.

*** Plagiochila chacabucensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 27, 1911.

*** Plagiochila emeiensis Grolle et M.L.So, Bryologist 101 (2): 282, 1998.

* Plagiochila fragilis Taylor, London J. Bot. 7: 198, 1848. 158

*** Plagiochila lunata S.W.Arnell, Bot. Not. 115: 204, 1962.

*** Plagiochila pachyloma Taylor, London J. Bot. 5: 267, 1846.

* Plagiochila pachyloma var. elatior Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 480, 1885.

** Plagiochila papillifolia Steph., Biblioth. Bot. 87 (2): 207, 1916.

*** Plagiochila parviramifera Inoue, J. Hattori Bot. Lab. 46: 317, 1979.

*** Plagiochila pseudoattenuata S.W.Arnell, Bot. Not. 115: 206, 1962.

*** Plagiochila punctata (Taylor) Taylor, London J. Bot. 5: 261, 1846. Bas.: Jungermannia punctata Taylor, Trans. Bot. Soc. Edinburgh 1: 179, 1844.

*** Plagiochila renauldii Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 156 (408), 1904.

*** Plagiochila retrorsa Gottsche, Mexik. Leverm.: 67, 1863.

*** Plagiochila rubescens (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 46, 1840. Bas.: Jungermannia rubescens Lehm. et Lindenb., Nov. Stirp. Pug. 6: 63, 1834.

*** Plagiochila sichuanensis Grolle et M.L.So, Bryologist 101 (2): 284, 1998.

*** Plagiochila spinulosa (Dicks.) Dumort., Recueil Observ. Jungerm.: 15, 1835. Bas.: Jungermannia spinulosa Dicks., Fasc. Pl. Crypt. Brit. 2: 14, 1790.

** Plagiochila sticticola Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 198, 1856.

*** Plagiochila stricta Lindenb., Sp. Hepat. (Lindenberg) 1: 20, 1839.

*** Plagiochila tronadoris Herzog, Darwiniana 11 (2): 214, 1957.

*** Plagiochila uncialis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 628, 1847. Bas.: Jungermannia uncialis Hook.f. et Taylor, London J. Bot. 3: 459, 1844.

*** Plagiochila wilmsiana Steph., Sp. Hepat. (Stephani) 6: 240, 1921.

* sect. Caducifoliae J.J.Engel et G.L.Merr., Nova Hedwigia 96 (3/4): 407, 2013.

** Plagiochila caducifolia Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 71, 1971.

* sect. Cardotiae Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 386, 1965.

*** Plagiochila cumingiana Steph., Bull. Herb. Boissier (sér. 2) 4 (1): 32 (404), 1904.

*** Plagiochila denticulata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 95, 1860 [1861].

*** Plagiochila fragillima Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 522 (326), 1903.

*** Plagiochila pseudorenitens Schiffn., Österr. Bot. Z. 49 (4): 132, 1899.

*** Plagiochila stevensiana Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 110 (290), 1903.

* sect. Cobanae Carl, Ann. Bryol., Suppl. 2: 79, 1931.

** Plagiochila cobana Steph., Sp. Hepat. (Stephani) 6: 138, 1918.

** Plagiochila detecta M.L.So et Grolle, Nova Hedwigia 71 (3/4): 391, 2000.

*** Plagiochila singularis Schiffn., Hep. Fl. Buitenzorg: 158, 1900.

*** Plagiochila tagawae Inoue, J. Hattori Bot. Lab. 38: 561, 1974.

** Plagiochila tixieri Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 87, 1975.

*** Plagiochila zhuensis Grolle et M.L.So, Bryologist 102 (2): 200, 1999.

*** sect. Cucullatae Schiffn., Hep. Fl. Buitenzorg: 107, 1900.

*** Plagiochila bantamensis (Reinw., Blume et Nees) Mont., Voy. Amér. Mérid., Bot. 7 (2): 82, 1839. Bas.: Jungermannia bantamensis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 235, 1824 [1825].

*** Plagiochila blepharophora (Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 102, 1840. Bas.: Jungermannia blepharophora Nees, Enum. Pl. Crypt. Javae: 71, 1830.

** Plagiochila chauviniana Mont., Ann. Sci. Nat. Bot. (sér. 3) 11: 34, 1849.

** Plagiochila clavatosaccata Steph., Bull. Herb. Boissier (sér. 2) 4 (1): 25 (397), 1904.

** Plagiochila grossispina Steph., Sp. Hepat. (Stephani) 6: 162, 1918.

*** Plagiochila integerrima Steph., Bot. Jahrb. Syst. 8 (2): 83, 1886.

* Plagiochila integrilobula Schiffn., Hep. Fl. Buitenzorg: 170, 1900. 159

** Plagiochila johannis-winkleri Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 187, 1931.

* Plagiochila kuhliana Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 292, 1864. 160

** Plagiochila kurzii Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 112 (292), 1903.

** Plagiochila reischeckiana Steph., Bull. Herb. Boissier (sér. 2) 3 (4): 331 (315), 1903.

*** Plagiochila sandei Dozy ex Sande Lac., Plagiochila Sandei: 5, 1856.

*** Plagiochila sciophila Nees, Sp. Hepat. (Lindenberg) 2-4: 100, 1840.

** Plagiochila sciophila subsp. ciliigera (R.M.Schust.) L.Söderstr., Phytotaxa 208 (1): 84, 2015. Bas.: Plagiochila japonica subsp. ciliigera R.M.Schust., Amer. Midl. Naturalist 62 (2): 354, 1959.

* Plagiochila stephanii Schiffn., Hep. Fl. Buitenzorg: 166, 1900. 161

** Plagiochila subplana Lindenb., Sp. Hepat. (Lindenberg) 2-4: 73, 1840.

** Plagiochila vitiensis Mitt., Bonplandia 9 (24): 367, 1861.

*** sect. Denticulatae Schiffn., Hep. Fl. Buitenzorg: 106, 1900.

*** Plagiochila alternans Lindenb. et Gottsche, Syn. Hepat. 5: 648, 1847.

** Plagiochila ansata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 649, 1847. Bas.: Jungermannia ansata Hook.f. et Taylor, London J. Bot. 3: 457, 1844.

*** Plagiochila banksiana Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 329, 1857.

** Plagiochila banksiana var. echinophora Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 62, 1971.

** Plagiochila chonotica Taylor, London J. Bot. 5: 260, 1846.

** Plagiochila equitans Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 331, 1857.

* Plagiochila fragmentissima Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 155, 1971. 162

*** Plagiochila gigantea Lindenb., Sp. Hepat. (Lindenberg) 2-4: 115, 1840. Nom. nov. pro Jungermannia gigantea Hook., Musci Exot. 1: tab. 93, 1818, nom. illeg.

** Plagiochila gigantea var. inermis J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 513, 2010.

*** Plagiochila gregaria (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 654, 1847. Bas.: Jungermannia gregaria Hook.f. et Taylor, London J. Bot. 3: 564, 1844.

** Plagiochila hookeriana Lindenb., Sp. Hepat. (Lindenberg) 2-4: 81, 1840.

** Plagiochila latifrons Gottsche et Hampe, Linnaea 27 (5): 553, 1854.

** Plagiochila minutula (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 652, 1847. Bas.: Jungermannia minutula Hook.f. et Taylor, London J. Bot. 3: 459, 1844.

*** Plagiochila nobilis Gottsche, Bot. Zeitung (Berlin) Beil. 16: 37, 1858.

** Plagiochila obovata Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 33, 1911.

*** Plagiochila ovata Lindenb. et Gottsche, Syn. Hepat. 5: 656, 1847.

*** Plagiochila retrospectans Lindenb., Sp. Hepat. (Lindenberg) 5: 123, 1843. Nom. nov. pro Jungermannia retrospectans Nees, Linnaea 6 (4): 619, 1831, nom. illeg.

** Plagiochila riparia Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 34, 1911.

** Plagiochila rutlandii Steph., Bull. Herb. Boissier (sér. 2) 4 (8): 776 (454), 1904.

** Plagiochila subpectinata Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 628, 1886.

*** sect. Durae Carl, Ann. Bryol., Suppl. 2: 123, 1931.

** Plagiochila acanthocaulis Sull., Hooker’s J. Bot. Kew Gard. Misc. 2: 317, 1850.

** Plagiochila angulata Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 26, 1900.

** Plagiochila bicornuta Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 305, 1896.

** Plagiochila crozetensis Kaal., Nyt Mag. Naturvidensk. 49 (2/3): 92, 1911.

*** Plagiochila deltoidea Lindenb., Sp. Hepat. (Lindenberg) 5: 132, 1843.

** Plagiochila deltoidea var. densa J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 506, 2010.

** Plagiochila dura De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 214, 1857.

** Plagiochila heterodonta (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 638, 1847. Bas.: Jungermannia heterodonta Hook.f. et Taylor, London J. Bot. 3: 460, 1844.

*** Plagiochila ramosissima (Hook.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 87, 1840. Bas.: Jungermannia ramosissima Hook., Musci Exot. 1: tab. 92, 1818.

*** sect. Duseniae Carl, Ann. Bryol., Suppl. 2: 126, 1931.

** Plagiochila dusenii Steph., Bull. Herb. Boissier (sér. 2) 4 (10): 979 (475), 1904.

** Plagiochila elata Taylor, London J. Bot. 5: 259, 1846.

** Plagiochila lechleri Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 325, 1857.

** Plagiochila validissima Steph., Biblioth. Bot. 87 (2): 214, 1916.

* sect. Flexicaules Carl, Ann. Bryol., Suppl. 2: 127, 1931.

** Plagiochila flexicaulis Mont., Syn. Hepat. 5: 629, 1847.

*** sect. Fruticosae Inoue, Gen. Plagiochila SE Asia: 50, 1984.

*** Plagiochila assamica Steph., Sp. Hepat. (Stephani) 6: 125, 1917.

** Plagiochila benitoi Inoue ex Piippo, J. Hattori Bot. Lab. 72: 122, 1992.

*** Plagiochila frondescens (Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 52, 1840. Bas.: Jungermannia frondescens Nees, Linnaea 6 (4): 610, 1831.

*** Plagiochila fruticosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 94, 1860 [1861].

** Plagiochila mauiensis Steph., Sp. Hepat. (Stephani) 6: 184, 1921.

*** Plagiochila pulcherrima Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 63, 1931.

** Plagiochila tamiensis Steph., Sp. Hepat. (Stephani) 6: 235, 1921.

*** sect. Fuscoluteae Carl, Ann. Bryol., Suppl. 2: 46, 1931.

*** Plagiochila aerea Taylor, London J. Bot. 5: 263, 1846.

*** Plagiochila dependula Taylor, London J. Bot. 5: 265, 1846.

*** Plagiochila fuscolutea Taylor, London J. Bot. 5: 263, 1846.

** Plagiochila heterophylla Lindenb., Nov. Stirp. Pug. 10: 2, 1857.

** Plagiochila heterophylla var. beauverdii (Steph.) Heinrichs, Bryophyt. Biblioth. 58: 148, 2002. Bas.: Plagiochila beauverdii Steph., Biblioth. Bot. 87 (2): 191, 1916.

*** Plagiochila longiramea Steph., Biblioth. Bot. 87 (2): 204, 1916.

*** Plagiochila paraphyllina Herzog, Hedwigia 74 (2): 89, 1934.

*** Plagiochila rudischusteri H.Rob., Beih. Nova Hedwigia 90: 199, 1988.

*** Plagiochila subbidentata Taylor, Ann. Mag. Nat. Hist. 20 (135): 381, 1847.

*** Plagiochila tabinensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 862 (231), 1902.

*** sect. Glaucescentes Carl, Ann. Bryol., Suppl. 2: 70, 1931.

*** Plagiochila buchtiniana Steph., Sp. Hepat. (Stephani) 6: 135, 1918.

*** Plagiochila diversifolia Lindenb. et Gottsche, Syn. Hepat. 5: 640, 1847.

*** Plagiochila longispina Lindenb. et Gottsche, Syn. Hepat. 5: 642, 1847.

*** sect. Hylacoetes Carl, Ann. Bryol., Suppl. 2: 50, 1931.

*** Plagiochila amicta Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 895 (561), 1905.

*** Plagiochila boryana Gottsche, Bull. Soc. Roy. Bot. Belgique 31: 118, 1892.

*** Plagiochila breuteliana Lindenb., Sp. Hepat. (Lindenberg) 5: 150, 1843.

*** Plagiochila canelensis Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 926 (576), 1905.

*** Plagiochila cucullifolia J.B.Jack et Steph., Hedwigia 31 (1): 24, 1892.

*** Plagiochila cucullifolia var. anomala Heinrichs et Gradst., Pl. Syst. Evol. 242 (1/4): 208, 2003.

*** Plagiochila dimorpha Lindenb. et Gottsche, Syn. Hepat. 5: 627, 1847.

*** Plagiochila dominicensis Taylor, London J. Bot. 5: 270, 1846.

*** Plagiochila ecuadorica (Inoue) L.Söderstr., Phytotaxa 208 (1): 84, 2015. Bas.: Steereochila ecuadorica Inoue, Mem. New York Bot. Gard. 45: 279, 1987.

*** Plagiochila ensiformis Taylor, London J. Bot. 5: 265, 1846.

*** Plagiochila flabelliflora Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 880 (249), 1902.

** Plagiochila guevarii H.Rob., Bryologist 70 (1): 48, 1967.

** Plagiochila husnotii Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 178 (506), 1905.

*** Plagiochila macrostachya Lindenb., Sp. Hepat. (Lindenberg) 2-4: 75, 1840.

*** Plagiochila patriciae Heinrichs et H.Anton, Bryophyt. Biblioth. 58: 107, 2002.

*** Plagiochila superba (Nees ex Spreng.) Mont. et Nees, Voy. Amér. Mérid., Bot. 7 (2): 81, 1839. Bas.: Jungermannia superba Nees ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 326, 1827.

*** Plagiochila superba var. macrotricha (Spruce) Heinrichs, Bryophyt. Biblioth. 58: 114, 2002. Bas.: Plagiochila macrotricha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 476, 1885.

*** Plagiochila turgida Herzog, Hedwigia 72 (6): 196, 1932.

*** Plagiochila vincentina Lindenb., Sp. Hepat. (Lindenberg) 2-4: 39, 1840.

* sect. Jacquinotiae Hässel, Nova Hedwigia 89 (1/2): 85, 2009.

** Plagiochila jacquinotii Mont., Voy. Pole Sud, Bot. 1: 273, 1845.

* sect. Kaalaasiae Carl, Ann. Bryol., Suppl. 2: 103, 1931.

** Plagiochila amboynensis Taylor, London J. Bot. 5: 260, 1846.

* sect. Longiflorae Carl, Ann. Bryol., Suppl. 2: 130, 1931.

** Plagiochila longiflora Mont., Syn. Hepat. 5: 651, 1847.

* sect. Oligodontes Carl, Ann. Bryol., Suppl. 2: 130, 1931.

** Plagiochila lophocoleoides Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 348, 1845.

*** sect. Peculiares Schiffn., Hep. Fl. Buitenzorg: 107, 1900.

** Plagiochila aspericaulis Grolle et M.L.So, Syst. Bot. 24 (3): 307, 1999.

** Plagiochila caulimammillosa Grolle et M.L.So, J. Bryol. 20 (1): 42, 1998.

*** Plagiochila devexa Steph., Bull. Herb. Boissier (sér. 2) 3 (4): 340 (324), 1903. Nom. nov. pro Plagiochila deflexa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861], nom. illeg.

*** Plagiochila durelii Schiffn., Österr. Bot. Z. 49 (4): 131, 1899.

** Plagiochila durelii subsp. guizhouensis Grolle et M.L.So, Syst. Bot. 24 (3): 304, 1999.

** Plagiochila grollei Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 384, 1965.

** Plagiochila hoei Inoue, J. Hattori Bot. Lab. 40: 421, 1976.

** Plagiochila hyalodermica Grolle et M.L.So, Bryologist 100 (4): 470, 1997.

** Plagiochila magna Inoue, J. Hattori Bot. Lab. 28: 216, 1965.

** Plagiochila paraphyllosa Grolle et M.L.So, Syst. Bot. 24 (3): 298, 1999.

** Plagiochila peculiaris Schiffn., Hep. Fl. Buitenzorg: 157, 1900.

*** Plagiochila perserrata Herzog, Symb. Sin. 5: 19, 1930.

** Plagiochila philippinensis Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 526 (330), 1903.

*** Plagiochila pseudopoeltii Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 382, 1965.

*** Plagiochila renitens (Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 90, 1840. Bas.: Jungermannia renitens Nees, Enum. Pl. Crypt. Javae: 76, 1830.

** Plagiochila semidecurrens (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 5: 142, 1843. Bas.: Jungermannia semidecurrens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 21, 1832.

** Plagiochila semidecurrens var. alaskana (A.Evans) Inoue, J. Hattori Bot. Lab. 28: 216, 1965. Bas.: Plagiochila alaskana A.Evans, Bull. Torrey Bot. Club 41 (12): 590, 1914 [1915].

*** Plagiochila vexans Schiffn. ex Steph., Sp. Hepat. (Stephani) 6: 237, 1921.

*** Plagiochila zangii Grolle et M.L.So, Bryologist 100 (4): 467, 1997.

*** Plagiochila zonata Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 225, 1894.

*** sect. Plagiochila

** Plagiochila arctica Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 41, 1906.

* Plagiochila arctica var. intermedia R.M.Schust., Amer. Midl. Naturalist 62 (1): 152, 1959.

*** Plagiochila asplenioides (L.) Dumort., Recueil Observ. Jungerm.: 14, 1835. Bas.: Jungermannia asplenioides L., Sp. Pl. 1: 1131, 1753.

*** Plagiochila bischleriana Grolle et M.L.So, Cryptog. Bryol. Lichénol. 18 (3): 191, 1997.

** Plagiochila britannica Paton, J. Bryol. 10 (3): 245, 1979.

*** Plagiochila chinensis Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 223, 1894.

*** Plagiochila circumdentata Steph., Bull. Herb. Boissier (sér. 2) 4 (8): 778 (456), 1904.

** Plagiochila circumdentata var. carinata J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 511, 2010.

** Plagiochila circumserrata Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 5 (1): 27, 1979.

** Plagiochila columbiana A.Evans, Bot. Gaz. 21 (4): 189, 1896.

*** Plagiochila delavayi Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 224, 1894.

*** Plagiochila elegans Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861].

*** Plagiochila hakkodensis Steph., Bull. Herb. Boissier 5 (2): 103, 1897.

** Plagiochila korthalsiana Molk., Ned. Kruidk. Arch. 3: 416, 1854 [1855].

** Plagiochila loriana Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 608 (354), 1903.

** Plagiochila microdentata M.L.So, New Zealand J. Bot. 38 (3): 426, 2000.

** Plagiochila mundaliensis Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 536 (340), 1903.

** Plagiochila orbicularis (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 3: 26, 1948 [1950]. Bas.: Plagiochila ovalifolia var. orbicularis S.Hatt., Bull. Tokyo Sci. Mus. 11: 61, 1944.

*** Plagiochila ovalifolia Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 193, 1891.

*** Plagiochila porelloides (Torr. ex Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 61, 1840. Bas.: Jungermannia porelloides Torr. ex Nees, Naturgesch. Eur. Leberm. 1: 170, 1833.

** Plagiochila porelloides var. norvegica (H.H.Blom et Holten) Schumacker et Váňa, Identif. keys liverw. hornw. Europe: 131, 2005. Bas.: Plagiochila norvegica H.H.Blom et Holten, Lindbergia 14 (1): 8, 1988.

** Plagiochila porelloides var. subarctica (Jørg.) Lammes, Fl. Fenn. 6: 54, 1977. Bas.: Plagiochila asplenioides var. subarctica Jørg., Bergens Mus. Skr. (n.ser.) 16: 173, 1934.

** Plagiochila schofieldiana Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 15 (1): 183, 1972.

*** Plagiochila secretifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 98, 1860 [1861].

** Plagiochila spinosa M.L.So, New Zealand J. Bot. 38 (3): 428, 2000.

** Plagiochila sumatrana Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 183, 1900 [1901].

** Plagiochila taiwanensis Inoue, Bull. Natl. Sci. Mus. Tokyo, B 8 (4): 136, 1982.

*** Plagiochila trapezoidea Lindenb., Sp. Hepat. (Lindenberg) 2-4: 112, 1840. Nom. nov. pro Jungermannia asplenioides β australis Nees, Enum. Pl. Crypt. Javae: 73, 1830.

** Plagiochila uniformis Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 98, 1860 [1861].

** sect. Poeltiae Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 394, 1965.

*** Plagiochila biondiana C.Massal., Hepat. Shen-si: 15, 1897.

*** Plagiochila carringtonii (Balf. ex Carrington) Grolle, Trans. Brit. Bryol. Soc. 4 (4): 656, 1964. Bas.: Adelanthus carringtonii Balf. ex Carrington, Trans. Bot. Soc. Edinburgh 10: 380, 1870.

** Plagiochila carringtonii subsp. lobuchensis Grolle, Trans. Brit. Bryol. Soc. 4 (4): 660, 1964.

*** Plagiochila duthiana Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 527 (331), 1903.

** Plagiochila erlangensis M.L.So, Haussknechtia, Beih. 9: 350, 1999.

*** Plagiochila poeltii Inoue et Grolle, Trans. Brit. Bryol. Soc. 4 (4): 656, 1964.

*** Plagiochila recurvata (W.E.Nicholson) Grolle, Trans. Brit. Bryol. Soc. 4 (4): 654, 1964. Bas.: Jamesoniella carringtonii var. recurvata W.E.Nicholson, Symb. Sin. 5: 13, 1930.

*** Plagiochila retusa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 96, 1860 [1861].

*** Plagiochila wallichiana Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 523 (327), 1903.

** Plagiochila wangii Inoue, J. Jap. Bot. 37 (6): 187, 1962.

** Plagiochila yulungensis Piippo, Ann. Bot. Fenn. 34 (4): 283, 1997.

*** sect. Rutilantes Carl, Ann. Bryol., Suppl. 2: 83, 1931.

*** Plagiochila bicuspidata Gottsche, Mexik. Leverm.: 43, 1863.

** Plagiochila caduciloba H.L.Blomq., Bryologist 42 (5): 114, 1939.

** Plagiochila chimborazensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 469, 1885.

* Plagiochila cuneata Lindenb. et Gottsche, Syn. Hepat. 5: 632, 1847. 163

*** Plagiochila debilis Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861].

*** Plagiochila deflexa Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 192, 1856.

*** Plagiochila defolians Grolle et M.L.So, Syst. Bot. 23 (4): 459, 1998 [1999].

*** Plagiochila exigua (Taylor) Taylor, London J. Bot. 5: 264, 1846. Bas.: Jungermannia exigua Taylor, Trans. Bot. Soc. Edinburgh 1: 179, 1844.

** Plagiochila fracta Pócs, Phytotaxa 195 (2): 183, 2015.

*** Plagiochila ghatiensis Steph., Sp. Hepat. (Stephani) 6: 159, 1918.

*** Plagiochila grossa Grolle et M.L.So, Syst. Bot. 23 (4): 461, 1998 [1999].

*** Plagiochila gymnocalycina (Lehm. et Lindenb.) Mont. et Nees, Voy. Amér. Mérid., Bot. 7 (2): 81, 1839. Bas.: Jungermannia gymnocalycina Lehm. et Lindenb., Nov. Stirp. Pug. 5: 28, 1833.

** Plagiochila gymnocalycina var. surinamensis (Molk.) Heinrichs et D.S.Rycroft, J. Hattori Bot. Lab. 100: 139, 2006. Bas.: Plagiochila surinamensis Molk., Syn. hepat. jav.: 103, 1856 [1857].

** Plagiochila loriloba Herzog ex Carl, Ann. Bryol., Suppl. 2: 47, 1931.

** Plagiochila maderensis Gottsche ex Steph., Bull. Herb. Boissier (sér. 2) 4 (4): 350 (426), 1904.

** Plagiochila oresitropha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 467, 1885.

*** Plagiochila pectinata Lindenb., Sp. Hepat. (Lindenberg) 1: 14, 1839.

*** Plagiochila rutilans Lindenb., Sp. Hepat. (Lindenberg) 2-4: 47, 1840.

*** Plagiochila rutilans var. moritziana (Lindenb. et Gottsche) Heinrichs, Bryologist 105 (2): 197, 2002. Bas.: Plagiochila moritziana Lindenb. et Gottsche, Linnaea 20 (3): 323, 1847.

*** Plagiochila rutilans var. standleyi (Herzog ex Carl) Heinrichs et D.S.Rycroft, Bryologist 104 (3): 357, 2001. Bas.: Plagiochila standleyi Herzog ex Carl, Ann. Bryol., Suppl. 2: 80, 1931.

* Plagiochila steyermarkii H.Rob., Bryologist 68 (1): 93, 1965. 164

*** Plagiochila trichostoma Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 113, 1864.

* sect. Strombifoliae Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 130, 1971.

** Plagiochila strombifolia Taylor, Nov. Stirp. Pug. 8: 5, 1844.

*** sect. Tayloriae Carl, Ann. Bryol., Suppl. 2: 140, 1931.

*** Plagiochila annotina Lindenb., Sp. Hepat. (Lindenberg) 1: 34, 1839.

*** Plagiochila baylisii Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 150, 1971.

*** Plagiochila bazzanioides J.J.Engel et G.L.Merr., Novon 9 (1): 29, 1999 (Engel and Smith Merrill 1999b).

*** Plagiochila chenii Grolle et M.L.So, Syst. Bot. 25 (1): 6, 2000.

*** Plagiochila circinalis (Lehm. et Lindenb.) Lehm., Sp. Hepat. (Lindenberg) 5: 124, 1843. Bas.: Jungermannia circinalis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 64, 1832.

** Plagiochila circinalis var. hemicardia (Hook.f. et Taylor) J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 512, 2010. Bas.: Jungermannia hemicardia Hook.f. et Taylor, London J. Bot. 3: 371, 1844.

*** Plagiochila colensoi Hook.f. et Taylor, London J. Bot. 5: 269, 1846.

** Plagiochila colensoi var. quinquespina (Steph.) J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 504, 2010. Bas.: Plagiochila quinquespina Steph., Bull. Herb. Boissier (sér. 2) 3 (4): 328 (312), 1903.

** Plagiochila corticola Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 224, 1894.

** Plagiochila fasciculata Lindenb., Sp. Hepat. (Lindenberg) 1: 7, 1839.

*** Plagiochila fusca Sande Lac., Ned. Kruidk. Arch. 3: 417, 1854 [1855].

** Plagiochila fuscella (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 648, 1847. Bas.: Jungermannia fuscella Hook.f. et Taylor, London J. Bot. 3: 373, 1844.

** Plagiochila fuscella var. novae-zelandiae (E.A.Hodgs.) J.J.Engel et G.L.Merr., Nova Hedwigia 89 (3/4): 294, 2009. Bas.: Plagiochila retrospectans var. novae-zelandiae E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 73 (4): 293, 1944.

*** Plagiochila gracilis Lindenb. et Gottsche, Syn. Hepat. 5: 632, 1847.

*** Plagiochila gymnoclada Sande Lac., Plagiochila Sandei: 6, 1856.

*** Plagiochila himalayana Schiffn., Österr. Bot. Z. 49 (4): 131, 1899.

** Plagiochila incurvicolla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 651, 1847. Bas.: Jungermannia incurvicolla Hook.f. et Taylor, London J. Bot. 3: 564, 1844.

** Plagiochila incurvicolla var. lonchoscypha (Herzog) J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 505, 2010. Bas.: Plagiochila lonchoscypha Herzog, Trans. & Proc. Roy. Soc. New Zealand 68 (1): 42, 1938.

** Plagiochila microdictyon Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 131, 1854.

** Plagiochila monospiris Inoue et Grolle, J. Hattori Bot. Lab. 36: 489, 1972 [1973].

*** Plagiochila nitens Inoue, Willdenowia 18 (2): 561, 1989.

** Plagiochila pseudocapillaris Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (3): 302, 1968.

*** Plagiochila pseudofirma Herzog, Symb. Sin. 5: 17, 1930.

** Plagiochila radiculosa Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 132, 1854.

** Plagiochila spathulifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 96, 1860 [1861].

*** Plagiochila stephensoniana Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 133, 1854.

*** sect. Trabeculatae S.Hatt. ex Inoue, J. Hattori Bot. Lab. 20: 75, 1958.

** Plagiochila austinii A.Evans, Rhodora 16 (184): 68, 1914.

*** Plagiochila flexuosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 94, 1860 [1861].

** Plagiochila sullivantii Gottsche, Bot. Gaz. 21 (4): 191, 1896.

** Plagiochila sullivantii var. spinigera R.M.Schust., Amer. Midl. Naturalist 62 (2): 323, 1959.

*** Plagiochila trabeculata Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 103 (283), 1903.

*** sect. Vagae Lindenb., Monogr. hep. gen. Plagiochilae: xv, 1844.

*** Plagiochila abietina (Nees) Mont. et Nees, Voy. Amér. Mérid., Bot. 7 (2): 81, 1839. Bas.: Jungermannia abietina Nees, Enum. Pl. Crypt. Javae: 76, 1830.

** Plagiochila abrupta Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 106, 1840.

** Plagiochila aequatorialis Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 334, 1857.

*** Plagiochila africana Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 973 (263), 1902.

*** Plagiochila akiyamae Inoue, Bull. Natl. Sci. Mus. Tokyo, B 12 (3): 73, 1986.

*** Plagiochila angusta Lindenb., Sp. Hepat. (Lindenberg) 5: 148, 1843.

*** Plagiochila angustitexta Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 977 (267), 1902.

*** Plagiochila arbuscula (Brid. ex Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 1: 23, 1839. Bas.: Jungermannia arbuscula Brid. ex Lehm. et Lindenb., Nov. Stirp. Pug. 4: 63, 1832.

** Plagiochila arbuscula var. rekohuensis J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 509, 2010.

* Plagiochila aspera Steph., Sp. Hepat. (Stephani) 6: 125, 1917. 165

** Plagiochila aspleniformis R.M.Schust., Amer. Midl. Naturalist 63 (1): 51, 1960.

** Plagiochila beddomei Steph., Bull. Herb. Boissier (sér. 2) 3 (10): 876 (361), 1903.

*** Plagiochila boivinii Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 987 (277), 1902.

*** Plagiochila bryopteroides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 499, 1885.

** Plagiochila contigua Gottsche, Mexik. Leverm.: 30, 1863.

*** Plagiochila corrugata (Nees) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 52, 1836. Bas.: Jungermannia corrugata Nees, Fl. Bras. (Martius) 1 (1): 378, 1833.

*** Plagiochila cuspidata Steph., Sp. Hepat. (Stephani) 6: 144, 1918.

** Plagiochila cymata Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 95, 1975.

** Plagiochila deflexirama Taylor, London J. Bot. 5: 262, 1846.

*** Plagiochila dichotoma (P.Beauv.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 53, 1836. Bas.: Carpolepidum dichotomum P.Beauv., Fl. Oware 1 (3): 23, 1805.

* Plagiochila dichotoma var. fluitans Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 490, 1885.

*** Plagiochila dissecta Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 600 (346), 1903.

*** Plagiochila disticha (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 107, 1840. Bas.: Jungermannia disticha Lehm. et Lindenb., Nov. Stirp. Pug. 6: 64, 1834.

** Plagiochila distinctifolia Lindenb., Sp. Hepat. (Lindenberg) 1: 17, 1839.

*** Plagiochila divergens Steph., Hedwigia 30 (6): 268, 1891.

*** Plagiochila drepanophylla Sande Lac., Syn. hepat. jav.: 103, 1856 [1857].

* Plagiochila drepanophylla var. minor Gottsche, Abh. Naturwiss. Vereins Bremen 7: 341, 1882.

*** Plagiochila effusa Steph., Bot. Jahrb. Syst. 20 (3): 310, 1895.

* Plagiochila effusa var. decurrens Steph., Bot. Jahrb. Syst. 20 (3): 310, 1895.

*** Plagiochila ericicola Steph., Bull. Herb. Boissier (sér. 2) 4 (6): 590 (441), 1904.

** Plagiochila exinnovata Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 600 (346), 1903.

** Plagiochila fastigiata Lindenb. et Gottsche, Syn. Hepat. 5: 657, 1847.

*** Plagiochila flabellata Steph., Bot. Jahrb. Syst. 8 (2): 82, 1886.

** Plagiochila floridana A.Evans, Bot. Gaz. 21 (4): 190, 1896.

*** Plagiochila fordiana Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 104 (284), 1903.

** Plagiochila francana Steph., Sp. Hepat. (Stephani) 6: 157, 1918.

*** Plagiochila furcifolia Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 194, 1891.

*** Plagiochila fusifera Taylor, London J. Bot. 5: 268, 1846.

*** Plagiochila hampeana Gottsche, Bot. Zeitung (Berlin) Beil. 16: 38, 1858.

* Plagiochila heterospina Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 128, 1914.

* Plagiochila heterostipa Steph., Hedwigia 31 (3): 129, 1892. 166

** Plagiochila incerta Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 324, 1857.

*** Plagiochila indica Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 532 (336), 1903.

** Plagiochila invisa (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 513, 1980. Bas.: Plagiochila ludoviciana var. invisa R.M.Schust., Amer. Midl. Naturalist 63 (1): 101, 1960.

*** Plagiochila javanica (Sw.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 52, 1836. Bas.: Jungermannia javanica Sw., Meth. Musc.: 35, 1781.

*** Plagiochila junghuhniana Sande Lac., Ned. Kruidk. Arch. 3: 416, 1854 [1855].

*** Plagiochila khasiana Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 95, 1860 [1861].

** Plagiochila kiaeri Gottsche, Abh. Naturwiss. Vereins Bremen 7: 341, 1882.

** Plagiochila kiaeri var. capensis (Steph.) M.Wigginton et Grolle, Bryophyt. Biblioth. 50: 182, 1996. Bas.: Plagiochila capensis Steph., Bull. Herb. Boissier (sér. 2) 4 (4): 350 (426), 1904.

** Plagiochila kiaeri var. myriocarpa (Pearson) Pócs, J. E. Afr. Nat. Hist. 96 (1): 38, 2007. Bas.: Plagiochila myriocarpa Pearson, Ark. Bot. 19 (5): 5, 1924.

*** Plagiochila kunmingensis Piippo, Ann. Bot. Fenn. 34 (4): 281, 1997.

** Plagiochila kurokawae Inoue, J. Hattori Bot. Lab. 32: 104, 1969.

*** Plagiochila laetevirens Lindenb., Sp. Hepat. (Lindenberg) 2-4: 101, 1840.

** Plagiochila lamellistipula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 491, 1885.

*** Plagiochila lastii Mitt., J. Linn. Soc., Bot. 22 (146): 320, 1886.

** Plagiochila latifolia Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 742 (541), 1905.

* Plagiochila loloënsis Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 166 (418), 1904. 167

** Plagiochila manillana Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 189, 1856.

** Plagiochila massalongoana Schiffn., Hep. Fl. Buitenzorg: 136, 1900.

** Plagiochila mastigophoroides Inoue, J. Hattori Bot. Lab. 32: 99, 1969.

** Plagiochila metcalfii Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 533 (337), 1903.

** Plagiochila micropteryx Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 107, 1864.

** Plagiochila miradorensis Gottsche, Mexik. Leverm.: 31, 1863.

** Plagiochila miradorensis var. convoluta R.M.Schust., Amer. Midl. Naturalist 63 (1): 113, 1960.

* Plagiochila moenkemeyeri Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 160 (412), 1904. 168

*** Plagiochila montagnei Nees, Ann. Sci. Nat. Bot. (sér. 2) 5: 53, 1836.

** Plagiochila morobensis Inoue et Piippo, Ann. Bot. Fenn. 26 (2): 203, 1989.

** Plagiochila multipinnula Herzog et S.Hatt., J. Hattori Bot. Lab. 14: 36, 1955.

*** Plagiochila neckeroidea Mitt., Trans. Linn. Soc. London 23 (1): 57, 1860.

*** Plagiochila nepalensis Lindenb., Sp. Hepat. (Lindenberg) 2-4: 93, 1840.

** Plagiochila norfolkiensis Steph., Bull. Herb. Boissier (sér. 2) 3 (10): 877 (362), 1903.

*** Plagiochila obtusa Lindenb., Sp. Hepat. (Lindenberg) 2-4: 42, 1840.

* Plagiochila owaihiensis Nees et Lindenb., Sp. Hepat. (Lindenberg) 1: 30, 1839. 169

** Plagiochila pacifica Mitt., Fl. vit.: 407, 1871 [1873].

** Plagiochila parallela Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 686 (223), 1902.

*** Plagiochila parvifolia Lindenb., Sp. Hepat. (Lindenberg) 1: 28, 1839.

*** Plagiochila patentissima Lindenb., Sp. Hepat. (Lindenberg) 2-4: 64, 1840.

*** Plagiochila patula (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 1: 21, 1839. Bas.: Jungermannia patula Sw., Fl. Ind. Occid. 3: 1844, 1806.

** Plagiochila patula var. brevifolia Gottsche, Mexik. Leverm.: 10, 1863.

** Plagiochila paucidens Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 117 (297), 1903.

** Plagiochila pensilis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 497, 1885.

** Plagiochila peradenyensis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 172, 1900 [1901].

** Plagiochila perdentata M.L.So et Grolle, Syst. Bot. 26 (3): 460, 2001.

*** Plagiochila pinniflora Steph., Hedwigia 30 (5): 212, 1891.

*** Plagiochila praemorsa Steph., Bot. Jahrb. Syst. 8 (2): 92, 1886.

** Plagiochila propinqua Sande Lac., Plagiochila Sandei: 5, 1856.

** Plagiochila purpurascens Steph., Sp. Hepat. (Stephani) 6: 197, 1921.

*** Plagiochila raddiana Lindenb., Sp. Hepat. (Lindenberg) 1: 9, 1839.

* Plagiochila ragazzii Gola, Ann. Bot. (Rome) 13 (1): 67, 1914.

*** Plagiochila repanda (Schwägr.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 62, 1840. Bas.: Jungermannia repanda Schwägr., Hist. Musc. Hepat. Prodr.: 26, 1814.

** Plagiochila repanda var. perrotana (Steph.) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 65, 1981. Bas.: Plagiochila perrotana Steph., Bull. Herb. Boissier (sér. 2) 4 (6): 586 (437), 1904.

*** Plagiochila rodriguezii Steph., Bot. Gaz. 15 (11): 290, 1890.

*** Plagiochila rudolfii Pócs, Beih. Nova Hedwigia 90: 223, 1988.

*** Plagiochila salacensis Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 576, 1853.

*** Plagiochila salvadorica Steph., Hedwigia 30 (6): 272, 1891.

** Plagiochila serrialata Herzog, Hedwigia 72 (6): 212, 1932.

*** Plagiochila shangaica Steph., Sp. Hepat. (Stephani) 6: 216, 1921.

*** Plagiochila simplex (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 54, 1840. Bas.: Jungermannia simplex Sw., Prodr. (Swartz): 143, 1788.

*** Plagiochila squamulosa Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 165, 1863.

** Plagiochila squamulosa var. crispulocaudata (Gottsche) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 74, 1981. Bas.: Plagiochila crispulocaudata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 340, 1882.

** Plagiochila squamulosa var. sinuosa (Mitt.) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 75, 1981. Bas.: Plagiochila sinuosa Mitt., J. Linn. Soc., Bot. 22 (146): 319, 1886.

** Plagiochila streimannii Inoue, J. Jap. Bot. 63 (11): 365, 1988.

*** Plagiochila strictifolia Steph., Hedwigia 30 (5): 210, 1891.

** Plagiochila subflabellata Colenso, Trans. & Proc. New Zealand Inst. 21: 51, 1889.

** Plagiochila subjavanica M.L.So, Austral. Syst. Bot. 13 (5): 804, 2000.

*** Plagiochila subtropica Steph., Bull. Soc. Roy. Bot. Belgique 38 (1): 46, 1899.

** Plagiochila tamariscina Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 685 (222), 1902.

** Plagiochila tecta Inoue et Grolle, J. Hattori Bot. Lab. 33: 319, 1970.

*** Plagiochila terebrans Nees et Mont., Sp. Hepat. (Lindenberg) 2-4: 98, 1840.

*** Plagiochila teysmannii Sande Lac., Plagiochila Sandei: 6, 1856.

** Plagiochila thyoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 498, 1885.

** Plagiochila tocarema Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 106, 1864.

** Plagiochila ulata Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 91, 1975.

** Plagiochila undata Sull., Amer. J. Sci. Arts (ser. 2) 1 (1): 73, 1846.

** Plagiochila undata subsp. crispata (Gottsche) R.M.Schust., Amer. Midl. Naturalist 63 (1): 122, 1960. Bas.: Plagiochila crispata Gottsche, Mexik. Leverm.: 71, 1863.

** Plagiochila ungarangana Sande Lac., Syn. hepat. jav.: 10, 1856 [1857].

*** Plagiochila virginica A.Evans, Prelim. cat. fl. W. Virginia: 497, 1892.

** Plagiochila virginica var. caroliniana R.M.Schust., Amer. Midl. Naturalist 63 (1): 15, 1960.

** Plagiochila virginica var. euryphylla R.M.Schust., Amer. Midl. Naturalist 63 (1): 21, 1960.

*** Plagiochila wightii Nees, Sp. Hepat. (Lindenberg) 2-4: 43, 1840.

** Plagiochila wilhelmina Inoue, J. Hattori Bot. Lab. 33: 317, 1970.

* sect. Zanteniae (Inoue) Inoue, Gen. Plagiochila SE Asia: 45, 1984. Bas.: Plagiochila subsect. Zanteniae Inoue, J. Hattori Bot. Lab. 32: 109, 1969.

** Plagiochila zantenii Inoue, J. Hattori Bot. Lab. 32: 107, 1969.

Incertae sedis

* Plagiochila abscendens Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 104, 1864.

** Plagiochila aculeata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 627, 1847. Bas.: Jungermannia aculeata Hook.f. et Taylor, London J. Bot. 3: 578, 1844.

** Plagiochila acuta Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 607 (353), 1903.

* Plagiochila albertii Steph., Biblioth. Bot. 87 (2): 188, 1916.

* Plagiochila aliena Gottsche, Mexik. Leverm.: 22, 1863. 170

** Plagiochila allionii Steph., Sp. Hepat. (Stephani) 6: 120, 1917.

* Plagiochila ambigua Lindenb. et Hampe, Linnaea 24 (6): 640, 1851 [1852].

* Plagiochila ampliata Steph., Biblioth. Bot. 87 (2): 189, 1916.

** Plagiochila andicola Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 187, 1856.

** Plagiochila andina Steph., Sp. Hepat. (Stephani) 6: 121, 1917.

* Plagiochila angolensis Steph., Sp. Hepat. (Stephani) 6: 122, 1917.

* Plagiochila angusteoblonga Steph., Biblioth. Bot. 87 (2): 189, 1916.

** Plagiochila angustisedens Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 743 (542), 1905.

** Plagiochila angustispina Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 671 (208), 1902.

** Plagiochila apicalis Gottsche, Mexik. Leverm.: 29, 1863.

** Plagiochila appalachiana Inoue, J. Hattori Bot. Lab. 40: 415, 1976. Nom. nov. pro Plagiochila yokogurensis subsp. fragilifolia R.M.Schust., J. Hattori Bot. Lab. 18: 18, 1957.

* Plagiochila arcta S.Winkl., Rev. Bryol. Lichénol. 42 (3): 818, 1976.

** Plagiochila arcuata Lindenb., Sp. Hepat. (Lindenberg) 2-4: 91, 1840.

** Plagiochila arenacea Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 53, 1964.

* Plagiochila artsii Pócs, J. Hattori Bot. Lab. 100: 334, 2006.

** Plagiochila atrovirens Taylor, London J. Bot. 5: 266, 1846.

** Plagiochila balansae Gottsche, Contr. Étude Plagiochila: 149, 1928.

** Plagiochila baldwinii Austin, Trans. Connecticut Acad. Arts 8 (15): 257, 1891.

* Plagiochila bamingensis Steph., Bull. Mus. Natl. Hist. Nat. 18 (2): 117, 1912.

** Plagiochila bancroftii Steph., Sp. Hepat. (Stephani) 6: 127, 1917.

*** Plagiochila barbadensis Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 897 (563), 1905.

* Plagiochila barbata Steph., Biblioth. Bot. 87 (2): 190, 1916.

* Plagiochila barbeyi Steph., Biblioth. Bot. 87 (2): 190, 1916.

* Plagiochila batava Steph., Sp. Hepat. (Stephani) 6: 128, 1917.

* Plagiochila berggrenii Steph., Biblioth. Bot. 87 (2): 191, 1916.

** Plagiochila beskeana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 863 (232), 1902.

** Plagiochila bialata Mitt., Fl. vit.: 407, 1871 [1873].

* Plagiochila biapiculata Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 891 (557), 1905.

* Plagiochila bicaudata Steph., Sp. Hepat. (Stephani) 6: 130, 1918.

** Plagiochila biciliata Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 529 (333), 1903.

** Plagiochila bicornis Hampe et Gottsche, Linnaea 25 (3): 338, 1852 [1853].

* Plagiochila bidentula Steph., Sp. Hepat. (Stephani) 6: 130, 1918.

** Plagiochila binghamiae A.Evans, Trans. Connecticut Acad. Arts 18 (5): 304, 1914.

** Plagiochila binominata Steph., Sp. Hepat. (Stephani) 6: 131, 1918.

** Plagiochila bitexta Dugas, Contr. Étude Plagiochila: 58, 1928.

** Plagiochila blepharobasis Herzog, Hedwigia 72 (6): 216, 1932.

** Plagiochila bogotensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 98, 1864.

* Plagiochila boliviana Spruce, Mem. Torrey Bot. Club 1 (3): 137, 1890.

* Plagiochila borneensis Steph., Sp. Hepat. (Stephani) 6: 132, 1918. 171

** Plagiochila brassii Inoue et Grolle, J. Hattori Bot. Lab. 36: 492, 1972 [1973].

* Plagiochila brevicalycina Lindenb. et Gottsche, Linnaea 20 (3): 322, 1847.

** Plagiochila brunneola Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 164 (416), 1904.

*** Plagiochila bryhnii Steph., Biblioth. Bot. 87 (2): 192, 1916.

** Plagiochila bunburii Taylor, London J. Bot. 5: 269, 1846.

* Plagiochila byssacea Hampe, Linnaea 20 (3): 326, 1847.

** Plagiochila caducidentata R.M.Schust., Phytologia 39 (4): 247, 1978.

** Plagiochila caldana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 879 (248), 1902.

** Plagiochila callaensis Steph., Sp. Hepat. (Stephani) 6: 136, 1918.

* Plagiochila camusii Steph., Biblioth. Bot. 87 (2): 193, 1916.

* Plagiochila capillicaulis Steph., Biblioth. Bot. 87 (2): 193, 1916.

** Plagiochila caribbeania R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 412, 1980.

* Plagiochila cava Steph., Bull. Herb. Boissier (sér. 2) 4 (12): 1213 (501), 1904.

** Plagiochila ceylanica Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 98, 1860 [1861].

** Plagiochila chacoënsis Herzog, Feddes Repert. Spec. Nov. Regni Veg. 55 (1): 7, 1952.

** Plagiochila chiloënsis Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 27, 1900.

** Plagiochila chinantlana Gottsche, Mexik. Leverm.: 12, 1863.

* Plagiochila chiovendae Gola, Ann. Bot. (Rome) 13 (1): 67, 1914.

* Plagiochila cinchonae Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 920 (570), 1905.

* Plagiochila circumvoluta Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 475, 1947.

*** Plagiochila cleefii Inoue, Stud. Cryptog. S. Peru: 95, 1987.

* Plagiochila concava Nees, Sp. Hepat. (Lindenberg) 2-4: 70, 1840.

** Plagiochila conduplicata Steph., Sp. Hepat. (Stephani) 6: 139, 1918.

* Plagiochila conferta Steph., Sp. Hepat. (Stephani) 6: 139, 1918.

** Plagiochila confertissima Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 182 (510), 1905.

** Plagiochila connata Lindenb. et Gottsche, Syn. Hepat. 5: 645, 1847.

** Plagiochila contorta Lindenb. et Hampe, Linnaea 24 (3): 301, 1851 [1852].

** Plagiochila convoluta Steph., Sp. Hepat. (Stephani) 6: 141, 1918.

** Plagiochila convolutifolia Steph., Sp. Hepat. (Stephani) 6: 142, 1918.

*** Plagiochila corymbulosa Pearson, Forh. Vidensk.-Selsk. Kristiania 1887 (9): 14, 1887.

** Plagiochila costariensis Horik., Acta Phytotax. Geobot. 13: 213, 1943. Nom. nov. pro Plagiochila pinnata Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 749 (548), 1905, nom. illeg.

** Plagiochila crispabilis Lindenb., Sp. Hepat. (Lindenberg) 1: 15, 1839.

* Plagiochila crispabilis var. minima Lindenb., Sp. Hepat. (Lindenberg) 5: 155, 1843.

** Plagiochila cristophylla Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 117 (297), 1903.

*** Plagiochila cuatrecasii H.Rob., Bryologist 70 (1): 47, 1967.

** Plagiochila cubensis Steph., Sp. Hepat. (Stephani) 6: 143, 1918.

** Plagiochila cucullata Lindenb. et Gottsche, Syn. Hepat. 5: 642, 1847.

* Plagiochila cuervina Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 96, 1864.

** Plagiochila delapsa Inoue, Beih. Nova Hedwigia 90: 171, 1988.

* Plagiochila delognei Steph., Biblioth. Bot. 87 (2): 196, 1916.

** Plagiochila denigrata Inoue, Willdenowia 18 (2): 557, 1989.

** Plagiochila densa Herzog, Hedwigia 72 (6): 222, 1932.

** Plagiochila densiflora Herzog, Hedwigia 72 (6): 226, 1932.

** Plagiochila densiramosa Steph., Biblioth. Bot. 87 (2): 196, 1916.

** Plagiochila deppeana Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 886 (552), 1905.

** Plagiochila desciscens Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 867 (236), 1902.

* Plagiochila dilatata Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 887 (553), 1905.

* Plagiochila distans Colenso, Trans. & Proc. New Zealand Inst. 19: 283, 1887. 172

** Plagiochila divaricata Lindenb., Sp. Hepat. (Lindenberg) 5: 147, 1843.

* Plagiochila doerfleri Steph., Biblioth. Bot. 87 (2): 196, 1916.

** Plagiochila dolichoblasta Herzog, Trans. Brit. Bryol. Soc. 1 (4): 286, 1950.

** Plagiochila dussiana Steph., Symb. Antill. (Urban) 3 (2): 277, 1902.

** Plagiochila echinata R.M.Schust., Amer. Midl. Naturalist 62 (2): 341, 1959.

** Plagiochila echinella Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 332, 1857.

* Plagiochila ecuadorensis Steph., Sp. Hepat. (Stephani) 6: 149, 1918.

* Plagiochila effuseintricata Steph., Biblioth. Bot. 87 (2): 197, 1916.

** Plagiochila eggersii Inoue, Bull. Natl. Sci. Mus. Tokyo, B 14 (4): 138, 1988.

** Plagiochila ekmanii S.W.Arnell, Bryologist 59 (4): 274, 1956.

* Plagiochila electa Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 739 (538), 1905.

* Plagiochila elegantula Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 161, 1955.

* Plagiochila emarginatobidentula Steph., Biblioth. Bot. 87 (2): 197, 1916.

* Plagiochila erythraeae Herzog, Hedwigia 78 (3/4): 224, 1938.

** Plagiochila estrellensis Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 23, 1925.

** Plagiochila eurydictyon Herzog, Hedwigia 72 (6): 207, 1932.

* Plagiochila excisa Steph., Sp. Hepat. (Stephani) 6: 153, 1918.

** Plagiochila exesa Lindenb. et Gottsche, Syn. Hepat. 5: 629, 1847.

* Plagiochila exilis Colenso, Trans. & Proc. New Zealand Inst. 19: 282, 1887. 173

** Plagiochila expansa Gottsche, Mexik. Leverm.: 18, 1863.

** Plagiochila facallonia Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 871 (240), 1902.

* Plagiochila falcato-oblonga Steph., Biblioth. Bot. 87 (2): 198, 1916.

** Plagiochila fallax Lindenb. et Hampe, Linnaea 24 (3): 300, 1851 [1852].

** Plagiochila faxinensis Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 60, 1964.

** Plagiochila fendleri Mont., Ann. Sci. Nat. Bot. (sér. 4) 6: 198, 1856.

* Plagiochila filicicola Steph., Bull. Herb. Boissier (sér. 2) 4 (4): 351 (427), 1904.

** Plagiochila fissidentoides Taylor, London J. Bot. 5: 264, 1846.

* Plagiochila flabellifrons Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 488, 1885.

** Plagiochila flava Steph., Sp. Hepat. (Stephani) 6: 156, 1918.

** Plagiochila flavescens (Gottsche, Lindenb. et Nees) Gottsche, Mexik. Leverm.: 52, 1863. Bas.: Plagiochila guilleminiana β flavescens Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 644, 1847.

* Plagiochila flavorufescens Steph., Biblioth. Bot. 87 (2): 199, 1916.

** Plagiochila footei A.Evans, Trans. Connecticut Acad. Arts 18 (5): 306, 1914.

* Plagiochila formosa Nees, Contr. Étude Plagiochila: 147, 1928.

** Plagiochila fragmentata R.M.Schust., Phytologia 45 (5): 421, 1980.

* Plagiochila frausa Gottsche, Mexik. Leverm.: 66, 1863.

* Plagiochila frausa var. boliviana Spruce, Mem. Torrey Bot. Club 1 (3): 134, 1890.

** Plagiochila frayjorgensis Hässel, Bol. Soc. Argent. Bot. 22 (1/4): 103, 1983. Nom. nov. pro Plagiochila modesta Herzog, Rev. Bryol. Lichénol. 23 (1/2): 34, 1954, nom. illeg.

* Plagiochila gaffatensis Gottsche ex Schweinf., Beitr. Fl. Aethiop.: 227, 1866 [1867].

** Plagiochila gaudichaudii Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 193, 1856.

** Plagiochila geniculata Lindenb., Sp. Hepat. (Lindenberg) 5: 131, 1843.

* Plagiochila geppii Steph., Biblioth. Bot. 87 (2): 199, 1916.

** Plagiochila germana Gottsche, Mexik. Leverm.: 34, 1863.

** Plagiochila germanii Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 938 (588), 1905.

* Plagiochila gibbiflora Steph., Bull. Herb. Boissier (sér. 2) 4 (6): 590 (441), 1904.

** Plagiochila gittinsii Inoue, Bull. Natl. Sci. Mus. Tokyo, B 3 (4): 139, 1977.

* Plagiochila glauca Carl, Ann. Bryol., Suppl. 2: 129, 1931. 174

* Plagiochila gracilicaulis Spruce, Mem. Torrey Bot. Club 1 (3): 132, 1890.

** Plagiochila gracillima Austin, Trans. Connecticut Acad. Arts 8 (15): 256, 1891.

* Plagiochila granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 111, 1864.

** Plagiochila granditexta Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 165 (417), 1904.

** Plagiochila grateloupii Mont., Ann. Sci. Nat. Bot. (sér. 4) 6: 188, 1856.

** Plagiochila guatemalensis Steph., Sp. Hepat. (Stephani) 6: 163, 1918.

** Plagiochila guttisquama Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 5 (1): 29, 1979.

** Plagiochila gymnocalyx Inoue, Trop. Bryol. 1: 34, 1989.

** Plagiochila haeseliae Inoue, Stud. Cryptog. S. Chile: 97, 1984.

* Plagiochila hans-meyeri Steph., Sp. Hepat. (Stephani) 6: 164, 1918.

** Plagiochila harlingii S.W.Arnell, Svensk Bot. Tidskr. 56 (2): 346, 1962.

** Plagiochila haumanii Herzog, Repert. Spec. Nov. Regni Veg. 41 (14/25): 285, 1937.

** Plagiochila hawaica Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 598 (344), 1903.

** Plagiochila heteracantha Steph., Sp. Hepat. (Stephani) 6: 166, 1918.

* Plagiochila heterofolia Steph., Biblioth. Bot. 87 (2): 200, 1916.

* Plagiochila hieronymii Steph., Biblioth. Bot. 87 (2): 201, 1916.

** Plagiochila hiroshiana Pócs, J. Hattori Bot. Lab. 100: 335, 2006.

** Plagiochila hoehnei Herzog, Hedwigia 72 (6): 220, 1932.

* Plagiochila holstii Steph., Hedwigia 45 (4): 214, 1906. Nom. nov. pro Plagiochila prostrata Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 168 (420), 1904, nom. illeg.

** Plagiochila horrida Gottsche, Mexik. Leverm.: 74, 1863.

* Plagiochila huatuscana Gottsche, Mexik. Leverm.: 24, 1863.

** Plagiochila huerlimannii Inoue, J. Hattori Bot. Lab. 33: 307, 1970.

* Plagiochila humboldtiana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 112, 1864.

** Plagiochila hyalina Lindenb., Syn. Hepat. 5: 640, 1847.

* Plagiochila incisa Dugas, Contr. Étude Plagiochila: 112, 1928.

** Plagiochila inflata Steph., Bull. Herb. Boissier (sér. 2) 3 (11): 961 (376), 1903.

* Plagiochila informifolia Steph., Biblioth. Bot. 87 (2): 202, 1916.

** Plagiochila infuscata Mitt., J. Linn. Soc., Bot. 15 (82): 63, 1876.

* Plagiochila injasutiensis S.W.Arnell, Bot. Not. 110: 404, 1957.

** Plagiochila inouei Grolle, J. Bryol. 10 (3): 269, 1979. Nom. nov. pro Plagiochila nudiuscula Inoue, Bull. Natl. Sci. Mus. Tokyo, B 3 (2): 45, 1977, nom. illeg.

** Plagiochila insularia Mitt., St. Helena: 366, 1875.

** Plagiochila intertexta Hook.f. et Taylor, London J. Bot. 5: 267, 1846.

* Plagiochila inuensis Steph., Biblioth. Bot. 87 (2): 202, 1916.

** Plagiochila irregularis Gottsche, Mexik. Leverm.: 14, 1863.

** Plagiochila itatiajensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 874 (243), 1902.

** Plagiochila jamaicensis Lindenb. et Hampe, Linnaea 24 (3): 302, 1851 [1852].

** Plagiochila jaramillii H.Rob., Bryologist 70 (1): 48, 1967.

* Plagiochila karstenii Steph., Hedwigia 45 (4): 214, 1906. Nom. nov. pro Plagiochila patentispina Steph., Bull. Herb. Boissier (sér. 2) 4 (1): 28 (400), 1904, nom. illeg. 175

* Plagiochila kaulfussiana Steph., Sp. Hepat. (Stephani) 6: 172, 1918.

** Plagiochila keckiana Steph., Bull. Herb. Boissier (sér. 2) 5 (4): 358 (526), 1905.

** Plagiochila kerneriana S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 60, 1964.

** Plagiochila koponenii Inoue et Piippo, Ann. Bot. Fenn. 26 (2): 216, 1989.

* Plagiochila lacerifolia Steph., Biblioth. Bot. 87 (2): 202, 1916.

* Plagiochila lachenaudii Steph., Biblioth. Bot. 87 (2): 202, 1916.

* Plagiochila laciniosa Dugas, Contr. Étude Plagiochila: 68, 1928.

* Plagiochila laevifolia Gola, Ann. Bot. (Rome) 6 (2): 273, 1907.

** Plagiochila lansbergii Gottsche, Mexik. Leverm.: 13, 1863.

** Plagiochila latitrigona Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 48, 1964.

* Plagiochila laxiramea Steph., Biblioth. Bot. 87 (2): 203, 1916.

* Plagiochila lecontei Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 165 (417), 1904. 176

* Plagiochila ledermanniana Beauverd, Sp. Hepat. (Stephani) 6: 572, 1924. Nom. nov. pro Plagiochila cucullifolia Steph., Sp. Hepat. (Stephani) 6: 243, 1922, nom. illeg. 177

* Plagiochila ledieui Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 161 (413), 1904. 178

** Plagiochila lehmanniana S.W.Arnell, Svensk Bot. Tidskr. 55 (1): 205, 1961.

** Plagiochila liebmanniana Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 97, 1840.

* Plagiochila lignicola Spruce, Mem. Torrey Bot. Club 1 (3): 135, 1890.

* Plagiochila lindaui Steph., Biblioth. Bot. 87 (2): 204, 1916.

* Plagiochila lindigiana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 105, 1864.

** Plagiochila lingua Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 677 (214), 1902.

* Plagiochila longifissa Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 891 (557), 1905.

* Plagiochila lotsyana Steph., Biblioth. Bot. 87 (2): 205, 1916.

* Plagiochila luteola Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 175 (503), 1905.

** Plagiochila lutescens Steph., Sp. Hepat. (Stephani) 6: 179, 1921.

** Plagiochila luzonensis Grolle et M.L.So, Bryologist 102 (1): 69, 1999.

* Plagiochila macra Taylor, London J. Bot. 7: 198, 1848.

** Plagiochila macrifolia Taylor, London J. Bot. 5: 270, 1846.

* Plagiochila macrifolia var. angustifolia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 470, 1885.

*** Plagiochila martiana (Nees) Lindenb., Sp. Hepat. (Lindenberg) 1: 12, 1839. Bas.: Jungermannia martiana Nees, Linnaea 6 (4): 617, 1831.

* Plagiochila matanga Steph., Sp. Hepat. (Stephani) 6: 183, 1921.

** Plagiochila maunakeana Steph., Sp. Hepat. (Stephani) 6: 186, 1921.

** Plagiochila maximiliana Gottsche, Mexik. Leverm.: 18, 1863.

** Plagiochila meghalayensis K.K.Rawat et S.C.Srivast., Geophytology 35 (1/2): 49, 2005.

** Plagiochila meridana Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 328, 1857.

** Plagiochila microdonta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861].

* Plagiochila mildbreadiana Beauverd, Sp. Hepat. (Stephani) 6: 571, 1924. Nom. nov. pro Plagiochila andongensis Steph., Sp. Hepat. (Stephani) 6: 122, 1917, nom. illeg.

** Plagiochila minarum Herzog, Hedwigia 72 (6): 207, 1932.

** Plagiochila minutiretis Reimers, Repert. Spec. Nov. Regni Veg. 21 (8/20): 264, 1925.

** Plagiochila miqueliana Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 95, 1840.

** Plagiochila molliuscula Inoue, Ann. Bot. Fenn. 13 (3): 134, 1976.

** Plagiochila mollusca Lehm., Nov. Stirp. Pug. 10: 4, 1857.

** Plagiochila moniliformis R.M.Schust., Phytologia 39 (4): 247, 1978.

** Plagiochila muelleriana Gottsche, Mexik. Leverm.: 38, 1863.

* Plagiochila multiflora Steph., Pflanzenw. Ost-Afrikas C: 64, 1895. 179

** Plagiochila mutila Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 327, 1857.

* Plagiochila naranjoënsis Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 687 (224), 1902.

** Plagiochila neesiana Lindenb., Sp. Hepat. (Lindenberg) 2-4: 71, 1840.

* Plagiochila neglecta Steph., Bull. Herb. Boissier (sér. 2) 5 (4): 351 (519), 1905.

* Plagiochila negrensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 466, 1885.

* Plagiochila negrii Gola, Ann. Bot. (Rome) 13 (1): 66, 1914.

* Plagiochila nigricaulis Steph., Biblioth. Bot. 87 (2): 206, 1916.

* Plagiochila notha Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 737 (536), 1905.

* Plagiochila nova Steph., Sp. Hepat. (Stephani) 6: 189, 1921.

** Plagiochila nudifolia (Steph.) Grolle, Feddes Repert. 82 (1): 89, 1971. Bas.: Jamesoniella nudifolia Steph., Biblioth. Bot. 87 (2): 184, 1916.

** Plagiochila nutans Steph., Bull. Herb. Boissier (sér. 2) 3 (11): 960 (375), 1903.

* Plagiochila obliquetruncata Steph., Biblioth. Bot. 87 (2): 207, 1916.

** Plagiochila oblonga Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 398, 1965.

** Plagiochila oerstediana Lindenb. et Hampe, Linnaea 24 (3): 301, 1851 [1852].

** Plagiochila olivacea Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 190 (518), 1905.

* Plagiochila ornata Wilson ex Lindenb., Sp. Hepat. (Lindenberg) 5: 164, 1843.

* Plagiochila ovato-obconica Steph., Sp. Hepat. (Stephani) 6: 191, 1921.

* Plagiochila ovifolia Steph., Sp. Hepat. (Stephani) 6: 191, 1921.

** Plagiochila palangiensis S.C.Srivast., K.K.Rawat et P.K.Verma, Natl. Acad. Sci. Lett. 29 (7/8): 267, 2006.

** Plagiochila panamensis Inoue, Bull. Natl. Sci. Mus. Tokyo, B 15 (3): 94, 1989.

** Plagiochila paramicola Herzog, Beih. Bot. Centralbl. 61B (3): 563, 1942.

* Plagiochila parvitexta Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 674 (211), 1902.

** Plagiochila parvivittata Inoue, Bull. Natl. Sci. Mus. Tokyo, B 13 (2): 50, 1987.

* Plagiochila parvula Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 726, 1913.

* Plagiochila pastasensis Steph., Sp. Hepat. (Stephani) 6: 197, 1921.

** Plagiochila patentispina Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 877 (246), 1902.

** Plagiochila patuloides Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 58, 1964.

** Plagiochila paucidentata Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 197, 1856.

** Plagiochila paupercula Gottsche, Mexik. Leverm.: 22, 1863.

* Plagiochila pearceana Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 742 (541), 1905.

** Plagiochila pellucida Lindenb. et Gottsche, Linnaea 20 (3): 321, 1847.

** Plagiochila perrottetiana Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 195, 1856.

** Plagiochila perrottetiana var. minor Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 101, 1864.

** Plagiochila pilifera Steph., Sp. Hepat. (Stephani) 6: 198, 1921.

** Plagiochila pinnatidens Steph., Sp. Hepat. (Stephani) 6: 199, 1921.

** Plagiochila pittieri Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 673 (210), 1902.

** Plagiochila platyphylla Herzog, Hedwigia 72 (6): 223, 1932.

** Plagiochila polopolensis Herzog, Hedwigia 67 (6): 261, 1927.

* Plagiochila prostrata Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 668 (205), 1902.

* Plagiochila pseudopatula Herzog, Hedwigia 72 (6): 227, 1932.

** Plagiochila pseudoradicans Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 24, 1925.

** Plagiochila ptychanthoidea Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 121 (301), 1903.

** Plagiochila pulchella Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 871 (240), 1902.

** Plagiochila pulvinata Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 526 (330), 1903.

** Plagiochila ratkowskiana Inoue, Brunonia 3 (1): 141, 1980.

** Plagiochila regeliana Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 675 (212), 1902.

** Plagiochila remyana Steph., Bull. Herb. Boissier (sér. 2) 3 (11): 963 (378), 1903.

*** Plagiochila revolvens Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851.

** Plagiochila rigidula Lindenb. et Gottsche, Linnaea 20 (3): 323, 1847.

** Plagiochila rosana Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 892 (558), 1905.

* Plagiochila rubricaulis Steph., Bot. Jahrb. Syst. 20 (3): 311, 1895.

* Plagiochila rufifolia Steph., Biblioth. Bot. 87 (2): 209, 1916.

** Plagiochila rufoviridis Spruce, Mem. Torrey Bot. Club 1 (3): 136, 1890.

* Plagiochila rusbyi Spruce, Mem. Torrey Bot. Club 1 (3): 135, 1890.

* Plagiochila sabensis Steph., Sp. Hepat. (Stephani) 6: 214, 1921.

** Plagiochila sachapatensis Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 679 (216), 1902.

** Plagiochila salazariae Inoue, Bull. Natl. Sci. Mus. Tokyo, B 15 (3): 91, 1989.

** Plagiochila saltuensis Spruce ex Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 927 (577), 1905.

** Plagiochila saltuensis var. spinosissima Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 163, 1955.

* Plagiochila samoana Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 85: 197, 1910.

** Plagiochila sarmentosa (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 86, 1840. Bas.: Jungermannia sarmentosa Lehm. et Lindenb., Linnaea 9 (4): 427, 1835.

* Plagiochila schiffneri Steph., Biblioth. Bot. 87 (2): 209, 1916.

* Plagiochila schinzei Steph., Biblioth. Bot. 87 (2): 209, 1916.

* Plagiochila schmidtii Steph., Biblioth. Bot. 87 (2): 209, 1916.

* Plagiochila schraderbergii Steph., Sp. Hepat. (Stephani) 6: 244, 1922. 180

** Plagiochila schubertiana Steph., Sp. Hepat. (Stephani) 6: 208, 1921.

** Plagiochila schusteri Inoue et Grolle, J. Hattori Bot. Lab. 33: 326, 1970.

** Plagiochila scissifolia Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 865 (234), 1902.

* Plagiochila scotica Macvicar ex Steph., Sp. Hepat. (Stephani) 6: 209, 1921. 181

** Plagiochila semiamplexicaulis Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 936 (586), 1905.

** Plagiochila semiermis Dugas, Contr. Étude Plagiochila: 66, 1928.

** Plagiochila silvatica Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 108, 1864.

* Plagiochila similis Steph., Biblioth. Bot. 87 (2): 210, 1916.

** Plagiochila sisparensis Steph., Sp. Hepat. (Stephani) 6: 207, 1921.

* Plagiochila slateri Steph., Biblioth. Bot. 87 (2): 211, 1916.

** Plagiochila solitaria Gottsche, Sp. Hepat. (Stephani) 6: 225, 1921.

*** Plagiochila spinifera Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 113, 1873.

* Plagiochila sprucei Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 860 (229), 1902.

* Plagiochila staudtiana Steph., Bull. Herb. Boissier (sér. 2) 4 (4): 352 (428), 1904.

** Plagiochila stipata Steph., Sp. Hepat. (Stephani) 6: 209, 1921.

* Plagiochila stolzii Steph., Sp. Hepat. (Stephani) 6: 244, 1922.

** Plagiochila subcontigua Herzog, Hedwigia 72 (6): 230, 1932.

* Plagiochila subconvoluta Gottsche, Mexik. Leverm.: 24, 1863.

* Plagiochila subdenudata Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 673 (210), 1902.

** Plagiochila subedentata Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 679 (216), 1902.

** Plagiochila subfragilis Inoue, Stud. Cryptog. S. Peru: 98, 1987.

* Plagiochila subhyalina Steph., Biblioth. Bot. 87 (2): 212, 1916.

** Plagiochila subligulata Steph., Sp. Hepat. (Stephani) 6: 224, 1921.

** Plagiochila sublyallii M.L.So, New Zealand J. Bot. 39 (1): 109, 2001.

* Plagiochila subrara Herzog, Hedwigia 74 (2): 87, 1934.

* Plagiochila subrotundifolia Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 935 (585), 1905.

** Plagiochila subundulata Lindenb., Sp. Hepat. (Lindenberg) 5: 137, 1843.

* Plagiochila suringarii Steph., Sp. Hepat. (Stephani) 6: 208, 1921.

** Plagiochila sylvicultrix Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 486, 1885.

* Plagiochila symmetrica Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 876 (245), 1902.

** Plagiochila tabinana Gottsche, Mexik. Leverm.: 46, 1863.

** Plagiochila talinayi S.W.Arnell, Svensk Bot. Tidskr. 50 (2): 312, 1956.

** Plagiochila tambillensis Loitl., Diagn. pl. nov.: 22, 1894.

** Plagiochila tarapotensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 863 (232), 1902.

* Plagiochila tenera Steph., Sp. Hepat. (Stephani) 6: 231, 1921.

** Plagiochila tenuis Lindenb., Sp. Hepat. (Lindenberg) 2-4: 50, 1840.

* Plagiochila tenuispica Steph., Sp. Hepat. (Stephani) 6: 234, 1921.

** Plagiochila thamniopsis Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cc, 1889 [1890].

* Plagiochila thollonii Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 980 (270), 1902.

** Plagiochila thrausta Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 5 (1): 34, 1979.

* Plagiochila tonduzana Steph., Sp. Hepat. (Stephani) 6: 227, 1921.

** Plagiochila tortuosa Lindenb. et Gottsche, Mexik. Leverm.: 70, 1863.

* Plagiochila tovarina Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 102, 1864.

* Plagiochila trabeculatospinosa Herzog, Hedwigia 72 (6): 213, 1932.

* Plagiochila trabutii Steph., Biblioth. Bot. 87 (2): 213, 1916.

** Plagiochila trianae Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 114, 1864.

** Plagiochila trichomanes Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cc, 1889 [1890].

* Plagiochila tricuspis Steph., Sp. Hepat. (Stephani) 6: 229, 1921.

** Plagiochila tristaniana Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 89, 2013. Nom. nov. pro Plagiochila fragilifolia S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 25, 1958, nom. illeg.

** Plagiochila tristis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 880 (249), 1902.

** Plagiochila trollii Herzog, Hedwigia 74 (2): 90, 1934.

** Plagiochila truncata Gottsche, Mexik. Leverm.: 25, 1863.

** Plagiochila uleana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 868 (237), 1902.

** Plagiochila umbrosioides L.Söderstr., Phytotaxa 208 (1): 85, 2015. Nom. nov. pro Plagiochila umbrosa Steph., Sp. Hepat. (Stephani) 6: 235, 1921, nom. illeg.

* Plagiochila unduavensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 882 (251), 1902.

* Plagiochila usambarana Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 980 (270), 1902.

** Plagiochila variedentata Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 887 (256), 1902.

** Plagiochila vastifolia Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 680 (217), 1902.

** Plagiochila velata Inoue et Piippo, Ann. Bot. Fenn. 26 (2): 207, 1989.

* Plagiochila venezuelana Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 749 (548), 1905.

* Plagiochila ventricosotrigona Steph., Biblioth. Bot. 87 (2): 214, 1916.

* Plagiochila verrucosa Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 885 (551), 1905.

* Plagiochila vetustisilva Steph., Sp. Hepat. (Stephani) 6: 245, 1922. 182

* Plagiochila viminea Spruce, Mem. Torrey Bot. Club 1 (3): 134, 1890.

* Plagiochila viridis Steph., Sp. Hepat. (Stephani) 6: 239, 1921.

** Plagiochila viridonigra (E.A.Hodgs.) Inoue, Bryologist 68 (2): 218, 1965. Bas.: Syzygiella viridonigra E.A.Hodgs., Rec. Domin. Mus. 4 (11): 120, 1962.

** Plagiochila vittiana Inoue, Beih. Nova Hedwigia 90: 171, 1988.

** Plagiochila vittifolia Steph., Sp. Hepat. (Stephani) 6: 239, 1921.

** Plagiochila vulcanica Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 671 (208), 1902.

** Plagiochila wacei S.W.Arnell ex Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 89, 2013.

* Plagiochila wallisiana Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 936 (586), 1905.

** Plagiochila wattsiana J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 511, 2010. Nom. nov. pro Plagiochila wattsii Steph., Sp. Hepat. (Stephani) 6: 240, 1921, nom. illeg.

** Plagiochila wettsteiniana S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 53, 1964.

* Plagiochila weymouthiana Steph., Biblioth. Bot. 87 (2): 215, 1916.

** Plagiochila wiemanniana S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 54, 1964.

*** Plagiochila winteri Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 981 (271), 1902.

** Plagiochila wrightii Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 681 (218), 1902.

** Plagiochila xalapensis Gottsche, Mexik. Leverm.: 21, 1863.

* Plagiochila xanthochroma Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 489, 1885.

* Plagiochila yoshinagana Steph., Sp. Hepat. (Stephani) 6: 242, 1922.

** Plagiochila zacuapana Gottsche, Mexik. Leverm.: 20, 1863.

** Plagiochilidium Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 186, 1931.

** Plagiochilidium bidentulum (Steph.) Grolle, J. Hattori Bot. Lab. 65: 408, 1988. Bas.: Tylimanthus bidentulus Steph., Bull. Herb. Boissier (sér. 2) 5 (12): 1134 (6), 1905.

** Plagiochilion S.Hatt., Biosphaera 1 (1): 7, 1947.

*** Plagiochilion braunianum (Nees) S.Hatt., Biosphaera 1 (1): 7, 1947. Bas.: Jungermannia brauniana Nees, Enum. Pl. Crypt. Javae: 80, 1830.

** Plagiochilion combinatum (Mitt.) Inoue, J. Hattori Bot. Lab. 27: 55, 1964. Bas.: Plagiochila combinata Mitt., Fl. vit.: 408, 1871 [1873].

*** Plagiochilion conjugatum (Hook.) R.M.Schust., J. Hattori Bot. Lab. 26: 285, 1963. Bas.: Jungermannia conjugata Hook., Musci Exot. 1: tab. 91, 1818.

** Plagiochilion fimbriatum (Mitt.) Inoue, J. Hattori Bot. Lab. 27: 57, 1964. Bas.: Plagiochila fimbriata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861].

** Plagiochilion giulianettii (Steph.) Inoue, J. Hattori Bot. Lab. 27: 57, 1964. Bas.: Plagiochila giulianettii Steph., Bull. Herb. Boissier (sér. 2) 4 (1): 30 (402), 1904.

** Plagiochilion herzogii Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (2): 270, 1971.

** Plagiochilion intermedium R.M.Schust., Phytologia 45 (5): 421, 1980.

** Plagiochilion mayebarae S.Hatt., J. Hattori Bot. Lab. 3: 39, 1948 [1950].

*** Plagiochilion oppositum (Reinw., Blume et Nees) S.Hatt., Biosphaera 1 (1): 7, 1947. Bas.: Jungermannia opposita Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 236, 1824 [1825].

** Plagiochilion pachycephalum (De Not.) Inoue, J. Hattori Bot. Lab. 27: 55, 1964. Bas.: Plagiochila pachycephala De Not., Epat. Borneo: 14, 1874.

*** Plagiochilion proliferum (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 285, 1963. Bas.: Plagiochila prolifera Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 130, 1854.

** Plagiochilion theriotanum (Steph.) Inoue, J. Hattori Bot. Lab. 27: 59, 1964. Bas.: Plagiochila theriotana Steph., Sp. Hepat. (Stephani) 6: 228, 1921.

** Pseudolophocolea R.M.Schust. et J.J.Engel, Lindbergia 8 (2): 71, 1982. Based on: Pseudolophocolea R.M.Schust. et J.J.Engel, Phytologia 47 (4): 310, 1981.

** Pseudolophocolea denticulata R.M.Schust. et J.J.Engel, Lindbergia 8 (2): 73, 1982. Based on: Pseudolophocolea denticulata R.M.Schust. et J.J.Engel, Phytologia 47 (4): 311, 1981, nom. inval.

** Xenochila R.M.Schust., Amer. Midl. Naturalist 62 (1): 15, 1959.

** Xenochila integrifolia (Mitt.) Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 6 (4): 373, 1963. Bas.: Plagiochila integrifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 96, 1860 [1861].

*** Pseudolepicoleaceae Fulford et J.Taylor

by M. von Konrat

Blepharostoma was recognized as an element within Pseudolepicoleaceae by, but we follow the concept of with Blepharostoma as the single genus in Blepharostomataceae.

*** Archeophylla R.M.Schust., J. Hattori Bot. Lab. 26: 263, 1963.

** Archeophylla paradoxa R.M.Schust., Trans. Brit. Bryol. Soc. 4 (5): 810, 1965.

** Archeophylla pungens (Herzog) R.M.Schust., Candollea 21 (1): 86, 1966. Bas.: Blepharostoma pungens Herzog, Rev. Bryol. Lichénol. 29 (3/4): 189, 1960 [1961].

*** Archeophylla schusteri (E.A.Hodgs. et Allison) R.M.Schust., J. Hattori Bot. Lab. 26: 263, 1963. Bas.: Temnoma schusteri E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand, Bot. 1 (12): 147, 1962.

*** Castanoclobos J.J.Engel et Glenny, Novon 17 (4): 424, 2007.

*** Castanoclobos julaceus (Hatcher ex J.J.Engel) J.J.Engel et Glenny, Novon 17 (4): 425, 2007. Bas.: Leiomitra julacea Hatcher ex J.J.Engel, Novon 9 (1): 26, 1999.

** Chaetocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 346, 1885.

*** Chaetocolea palmata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 346, 1885.

*** Herzogiaria Fulford ex Hässel, Lindbergia 7 (1): 23, 1981.

*** Herzogiaria teres (Steph.) Fulford ex Hässel, Lindbergia 7 (1): 24, 1981. Bas.: Lepicolea teres Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 26, 1901.

** Isophyllaria E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand, Bot. 3 (4): 68, 1965.

*** Isophyllaria attenuata (Rodway) E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 111, 1972. Bas.: Isotachis attenuata Rodway, Pap. & Proc. Roy. Soc. Tasmania 1916: 47, 1917.

** Isophyllaria fuegiana (Hässel) R.M.Schust., Beih. Nova Hedwigia 118: 141, 2000. Bas.: Fulfordiella fuegiana Hässel, Comun. Mus. Argent. Ci. Nat. “Bernardino Rivadavia,” Ci. Bot. 2 (9): 48, 1974.

*** Pseudolepicolea Fulford et J.Taylor, Nova Hedwigia 1 (3/4): 412, 1959 [1960].

** Pseudolepicolea andoi (R.M.Schust.) Inoue, Bull. Natl. Sci. Mus. Tokyo, B 4 (3): 94, 1978. Bas.: Lophochaete andoi R.M.Schust., J. Hattori Bot. Lab. 26: 261, 1963.

** Pseudolepicolea fryei (Perss.) Grolle et Ando, Hikobia 3 (3): 180, 1963. Bas.: Lepicolea fryei Perss., Bryologist 49 (2): 47, 1946.

*** Pseudolepicolea grolleana (R.M.Schust.) Grolle, Ann. Bot. Fenn. 21 (1): 30, 1984. Bas.: Archeochaete grolleana R.M.Schust., Nova Hedwigia 15: 441, 1968.

*** Pseudolepicolea kuehnemannii (R.M.Schust.) Hässel, Fl. Criptog. Tierra del Fuego 15: 125, 1975. Bas.: Archeochaete kuehnemannii R.M.Schust., J. Hattori Bot. Lab. 26: 262, 1963.

*** Pseudolepicolea quadrilaciniata (Sull.) Fulford et J.Taylor, Nova Hedwigia 1 (3/4): 413, 1959 [1960]. Bas.: Sendtnera quadrilaciniata Sull., Hooker’s J. Bot. Kew Gard. Misc. 2: 317, 1850.

*** Pseudolepicolea temnomoides (R.M.Schust.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 92, 2013. Bas.: Archeochaete temnomoides R.M.Schust., Candollea 21 (1): 129, 1966.

** Pseudolepicolea trollii (Herzog) Grolle et Ando, Hikobia 3 (3): 177, 1963. Bas.: Blepharostoma trollii Herzog, Ann. Bryol. 12: 80, 1939.

* Pseudolepicolea trollii var. darjeelingensis S.Hatt. et Mizut., J. Hattori Bot. Lab. 31: 252, 1968.

*** Temnoma Mitt., Handb. N. Zeal. fl. 2: 750, 1867.

*** Temnoma angustifolium R.M.Schust., Candollea 21 (2): 279, 1966 [1967].

*** Temnoma chaetophyllum R.M.Schust., Phytologia 39 (4): 239, 1978.

** Temnoma palmatum (Lindb. ex Pearson) R.M.Schust., Bryologist 62 (4): 240, 1959 [1960]. Bas.: Blepharostoma palmatum Lindb. ex Pearson, J. Bot. 25: 193, 1887.

** Temnoma palmatum var. cuneatum R.M.Schust., Candollea 21 (2): 347, 1966 [1967].

** Temnoma palmatum var. laxifolium R.M.Schust., Candollea 21 (2): 345, 1966 [1967].

** Temnoma palmatum var. pseudospiniferum R.M.Schust., Candollea 21 (2): 348, 1966 [1967].

** Temnoma patagonicum R.M.Schust., Candollea 21 (2): 313, 1966 [1967].

*** Temnoma paucisetigerum R.M.Schust., Candollea 21 (2): 266, 1966 [1967].

*** Temnoma pilosum (A.Evans) R.M.Schust., Bryologist 62 (4): 240, 1959 [1960]. Bas.: Blepharostoma pilosum A.Evans, Bull. Torrey Bot. Club 25 (8): 413, 1898.

*** Temnoma pulchellum (Hook.) Mitt., Handb. N. Zeal. fl. 2: 753, 1867. Bas.: Jungermannia pulchella Hook., Musci Exot. 1: tab. 94, 1818.

*** Temnoma quadrifidum (Mitt.) Mitt. ex E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand, Bot. 1 (12): 142, 1962. Bas.: Jungermannia quadrifida Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 128, 1854.

*** Temnoma quadripartitum (Hook.) Mitt., J. Linn. Soc., Bot. 15 (82): 68, 1876. Bas.: Jungermannia quadripartita Hook., Musci Exot. 2: tab. 117, 1820.

** Temnoma quadripartitum var. pseudopungens R.M.Schust., Candollea 21 (2): 312, 1966 [1967].

** Temnoma quadripartitum var. randii (S.W.Arnell) R.M.Schust., Candollea 21 (2): 307, 1966 [1967]. Bas.: Lepidozia randii S.W.Arnell, Svensk Bot. Tidskr. 47 (3): 417, 1953.

*** Temnoma setigerum (Lindenb.) R.M.Schust., Nova Hedwigia 5: 35, 1963. Bas.: Jungermannia setigera Lindenb., Syn. Hepat. 1: 131, 1844.

** Temnoma setigerum var. hawaiicum Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 17 (3): 228, 1974.

** Temnoma townrowii R.M.Schust., Candollea 21 (2): 351, 1966 [1967].

*** Trichocoleaceae Nakai

by T. Katagiri

The treatment of Trichocoleaceae follows what was outlined in and, ). Recent nomenclatural and taxonomic notes can also be found in.

*** Eotrichocolea R.M.Schust., J. Hattori Bot. Lab. 26: 252, 1963.

*** Eotrichocolea furukii T.Katag., Bryologist 115 (4): 519, 2012.

*** Eotrichocolea polyacantha (Hook.f. et Taylor) R.M.Schust., J. Hattori Bot. Lab. 26: 264, 1963. Bas.: Jungermannia polyacantha Hook.f. et Taylor, London J. Bot. 3: 290 [390], 1844.

*** Leiomitra Lindb., Acta Soc. Sci. Fenn. 10: 515, 1875.

** subg. Brachygyna R.M.Schust., Nova Hedwigia 73 (3/4): 480, 2001.

** Leiomitra mastigophoroides R.M.Schust., Phytologia 45 (5): 416, 1980.

** subg. Leiomitra, Nova Hedwigia 73 (3/4): 480, 2001.

*** Leiomitra breviseta (Steph.) R.M.Schust., Beih. Nova Hedwigia 118: 152, 2000. Bas.: Trichocolea breviseta Steph., Sp. Hepat. (Stephani) 4: 60, 1909.

*** Leiomitra capillata Lindb., Acta Soc. Sci. Fenn. 10: 515, 1875.

*** Leiomitra elegans (Lehm.) Hässel, Novon 12 (4): 465, 2002. Bas.: Trichocolea elegans Lehm., Nov. Stirp. Pug. 10: 8, 1857.

** Leiomitra elliottii (Steph.) R.M.Schust., Nova Hedwigia 73 (3/4): 469, 2001. Bas.: Trichocolea elliottii Steph., Sp. Hepat. (Stephani) 4: 55, 1909.

*** Leiomitra flaccida Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 349, 1885.

** Leiomitra hirticaulis R.M.Schust., Nova Hedwigia 73 (3/4): 472, 2001.

*** Leiomitra lanata (Hook.) R.M.Schust., Phytologia 45 (5): 417, 1980. Bas.: Jungermannia lanata Hook., Musci Exot. 2: tab. 116, 1820.

*** Leiomitra merrillana (Steph.) T.Katag., Bryologist 115 (4): 488, 2012. Bas.: Trichocolea merrillana Steph., Sp. Hepat. (Stephani) 6: 374, 1923.

*** Leiomitra paraphyllina Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 350, 1885.

** Leiomitra robusta (Steph.) R.M.Schust., Nova Hedwigia 73 (3/4): 469, 2001. Bas.: Trichocolea robusta Steph., Sp. Hepat. (Stephani) 4: 58, 1909.

** Leiomitra smaragdina Hässel, Novon 12 (4): 467, 2002.

*** Leiomitra tomentosa (Sw.) Lindb., Acta Soc. Sci. Fenn. 10: 515, 1875. Bas.: Jungermannia tomentosa Sw., Prodr. (Swartz): 145, 1788.

*** Trichocolea Dumort., Commentat. Bot. (Dumortier): 113, 1822 nom. conserv.

** Trichocolea argentea Herzog, Arch. Bot. São Paulo 1 (2): 40, 1925.

*** Trichocolea brevifissa Steph., Sp. Hepat. (Stephani) 4: 54, 1909.

** Trichocolea comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 27, 1922.

*** Trichocolea filicaulis Steph., Sp. Hepat. (Stephani) 4: 59, 1909.

* Trichocolea floccosa Herzog et Hatcher, Lloydia 20 (3): 148, 1957 [1958]. 183

** Trichocolea geniculata Pearson, J. Linn. Soc., Bot. 46 (305): 28, 1922.

** Trichocolea gracillima Austin, Bot. Gaz. 3 (1): 6, 1878.

*** Trichocolea hatcheri E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 69, 1965.

*** Trichocolea iriomotensis T.Katag., Hattoria 4: 6, 2013.

*** Trichocolea japonica T.Katag., Bryologist 114 (4): 744, 2011.

*** Trichocolea magna T.Katag., Hattoria 4: 7, 2013.

** Trichocolea minutifolia Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 134, 1914.

*** Trichocolea mollissima (Hook.f. et Taylor) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 132, 1864. Bas.: Jungermannia mollissima Hook.f. et Taylor, London J. Bot. 3: 290 [390], 1844.

*** Trichocolea pluma (Reinw., Blume et Nees) Mont., Voy. Bonite, Bot. 1: 238, 1846. Bas.: Jungermannia pluma Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 209, 1824 [1825].

*** Trichocolea rigida R.M.Schust., Nova Hedwigia 15: 447, 1968.

*** Trichocolea rudimentaris Steph., Sp. Hepat. (Stephani) 6: 376, 1923.

* Trichocolea sprucei Steph., Sp. Hepat. (Stephani) 4: 59, 1909. Nom. nov. pro Trichocolea gracillima Spruce, J. Linn. Soc., Bot. 30 (210): 353, 1895, nom. illeg. 184

*** Trichocolea tomentella (Ehrh.) Dumort., Syll. Jungerm. Europ.: 67, 1831. Bas.: Jungermannia tomentella Ehrh., Hannover. Mag. 21 (18): 277, 1783.

** Trichocolea udarii D.K.Singh, Bull. Bot. Surv. India 25: 177, 1983 [1985].

** Trichocolea wattsiana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 135, 1914.

*** Myliaceae Schljakov

by L. Söderström

outlines the controversy as to the placement of Mylia anomala where they advocate recognition at the subgeneric level. It has previously been recognized as a segregate genus, Leiomylia J.J.Engel and Braggins (cf. also ).

*** Mylia Gray, Nat. Arr. Brit. Pl. 1: 693, 1821 nom. conserv.

** subg. Anomalae (R.M.Schust. ex Potemkin) L.Söderstr., Phytotaxa 202 (1): 70, 2015. Bas.: Mylia sect. Anomalae R.M.Schust. ex Potemkin, Arctoa 2: 1, 1993.

*** Mylia anomala (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 693, 1821. Bas.: Jungermannia anomala Hook., Brit. Jungermann.: tab. 34, 1813.

** subg. Mylia

*** Mylia taylorii (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 693, 1821. Bas.: Jungermannia taylorii Hook., Brit. Jungermann.: tab. 57, 1813.

*** Mylia verrucosa Lindb., Acta Soc. Sci. Fenn. 10: 236, 1872 [1873].

*** Mylia verrucosa subsp. nuda (Inoue et B.Y.Yang) Potemkin et Kazan., Arctoa 2: 5, 1993. Bas.: Mylia nuda Inoue et B.Y.Yang, Taiwania 12 (1): 35, 1966.

*** Schistochilaceae H.Buch

by X. He and D. Glenny

provided an historical account of Schistochilaceae summarizing studies that have showed that the phylogenetic structure of the family does not match units that have resulted from morphologically-based investigations. Further studies are needed until a natural division of the family can be proposed. Thus, we follow the broad concept of and here, treating Schistochilaceae as comprising a single genus as discussed by.

*** Schistochila Dumort., Recueil Observ. Jungerm.: 15, 1835.

** Schistochila acuminata Steph., Sp. Hepat. (Stephani) 4: 81, 1909.

*** Schistochila aequiloba Steph., Sp. Hepat. (Stephani) 4: 80, 1909.

*** Schistochila alata (Lehm.) Schiffn., Hepat. (Engl.-Prantl): 111, 1893. Bas.: Jungermannia alata Lehm., Linnaea 4: 359, 1829.

*** Schistochila aligera (Nees et Blume) J.B.Jack et Steph., Hedwigia 31 (1): 12, 1892. Bas.: Jungermannia aligera Nees et Blume, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 11: 135, 1823.

* Schistochila aligera var. laxa (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 212, 1898. Bas.: Jungermannia aligera γ laxa Nees, Enum. Pl. Crypt. Javae: 68, 1830.

*** Schistochila altissima E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 85, 1965.

** Schistochila altissima subsp. polystratosa R.M.Schust. et J.J.Engel, Phytologia 30 (4): 241, 1975.

*** Schistochila antara Grolle, J. Hattori Bot. Lab. 29: 249, 1966.

*** Schistochila appendiculata (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877. Bas.: Jungermannia appendiculata Hook., Musci Exot. 1: tab. 15, 1818.

* Schistochila baileyana Steph., Sp. Hepat. (Stephani) 4: 85, 1909. 185

*** Schistochila balfouriana (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 4: 91, 1909. Bas.: Jungermannia balfouriana Hook.f. et Taylor, London J. Bot. 3: 556, 1844.

*** Schistochila beccariana (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877. Bas.: Gottschea beccariana De Not., Epat. Borneo: 9, 1874.

*** Schistochila berggrenii (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010. Bas.: Pachyschistochila berggrenii J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 58: 476, 1985.

*** Schistochila berteroana (Hook.) Steph., Sp. Hepat. (Stephani) 4: 96, 1909. Bas.: Jungermannia berteroana Hook., Bot. Misc. 2: 148, 1831.

*** Schistochila blumei (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877. Bas.: Jungermannia blumei Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 11: 136, 1823.

** Schistochila brassii Grolle, J. Hattori Bot. Lab. 31: 5, 1968.

** Schistochila caledonica Steph., Sp. Hepat. (Stephani) 4: 77, 1909.

*** Schistochila carnosa (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 93, 1909. Bas.: Gottschea carnosa Mitt., J. Linn. Soc., Bot. 15 (82): 72, 1876.

*** Schistochila caudata R.M.Schust. et J.J.Engel, Phytologia 30 (4): 242, 1975.

*** Schistochila childii (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010. Bas.: Pachyschistochila childii R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 471, 1985.

*** Schistochila chlorophylla (Colenso) E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 71 (3): 192, 1941. Bas.: Gottschea chlorophylla Colenso, Trans. & Proc. New Zealand Inst. 18: 240, 1886.

*** Schistochila ciliata (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 87, 1909. Bas.: Gottschea ciliata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 151, 1854.

*** Schistochila compacta (Colenso) E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 71 (3): 192, 1941. Bas.: Gottschea compacta Colenso, Trans. & Proc. New Zealand Inst. 16: 349, 1884.

*** Schistochila conchophylla Herzog ex E.A.Hodgs. et Allison, Trans. & Proc. Roy. Soc. New Zealand 71 (3): 191, 1941.

** Schistochila conchophylla var. multidentata (J.J.Engel) Xiao L.He et Glenny, Phytotaxa 173 (1): 92, 2014. Bas.: Gottschea conchophylla var. multidentata J.J.Engel, Nova Hedwigia 93 (3/4): 407, 2011.

*** Schistochila congoana Steph., Sp. Hepat. (Stephani) 4: 71, 1909.

** Schistochila cookei (H.A.Mill.) R.M.Schust., J. Hattori Bot. Lab. 42: 274, 1977. Bas.: Fulfordistria cookei H.A.Mill., Phytologia 20 (5): 320, 1970.

** Schistochila crinita Grolle, J. Hattori Bot. Lab. 31: 7, 1968.

*** Schistochila cristata Steph., Hedwigia 28 (4): 274, 1889.

*** Schistochila cunninghamii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 27, 1901.

** Schistochila doriae (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877. Bas.: Gottschea doriae De Not., Epat. Borneo: 10, 1874.

** Schistochila engleriana Steph., Sp. Hepat. (Stephani) 4: 69, 1909.

*** Schistochila exalata Herzog, Rev. Bryol. Lichénol. 29 (3/4): 191, 1960 [1961].

** Schistochila fijiensis H.Buch et Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 94, 1952.

*** Schistochila glaucescens (Hook.) A.Evans, Rev. Bryol. 32 (4): 60, 1905. Bas.: Jungermannia glaucescens Hook., Musci Exot. 1: tab. 39, 1818.

** Schistochila hattorii Grolle, J. Hattori Bot. Lab. 29: 247, 1966.

** Schistochila integerrima Steph., Sp. Hepat. (Stephani) 6: 492, 1924.

** Schistochila isotachyphylla (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010. Bas.: Paraschistochila isotachyphylla J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 58: 429, 1985.

*** Schistochila kirkiana Steph., Sp. Hepat. (Stephani) 4: 86, 1909.

*** Schistochila kunkelii S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 12, 1957.

** Schistochila lacerata Steph., Sp. Hepat. (Stephani) 6: 492, 1924.

*** Schistochila lamellata (Hook.) Dumort. ex A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892. Bas.: Jungermannia lamellata Hook., Musci Exot. 1: tab. 49, 1818.

*** Schistochila laminigera (Hook.f. et Taylor) A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892. Bas.: Jungermannia laminigera Hook.f. et Taylor, London J. Bot. 3: 456, 1844.

*** Schistochila latiloba (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010. Bas.: Pachyschistochila latiloba R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 493, 1985.

*** Schistochila lehmanniana (Lindenb.) Steph., Sp. Hepat. (Stephani) 4: 86, 1909. Bas.: Jungermannia lehmanniana Lindenb., Nov. Stirp. Pug. 4: 60, 1832.

*** Schistochila leucophylla (Lehm. ex Gottsche, Lindenb. et Nees) Steph., Sp. Hepat. (Stephani) 4: 98, 1910. Bas.: Gottschea leucophylla Lehm. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 1: 17, 1844.

** Schistochila macrodonta W.E.Nicholson, Symb. Sin. 5: 29, 1930.

** Schistochila minor C.Gao et Y.H.Wu, J. Hattori Bot. Lab. 95: 267, 2004.

*** Schistochila monticola R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (4): 623, 1971.

*** Schistochila muricata E.A.Hodgs. et Allison, Trans. & Proc. Roy. Soc. New Zealand 71 (3): 186, 1941.

*** Schistochila nadeaudiana Steph., Sp. Hepat. (Stephani) 4: 76, 1909.

** Schistochila neesii (Mont.) Lindb., J. Linn. Soc., Bot. 13 (67): 194, 1872 [1873]. Bas.: Gottschea neesii Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 244, 1843.

*** Schistochila nitidissima R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 27, 1968.

*** Schistochila nivicola (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010. Bas.: Pachyschistochila nivicola R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 512, 1985.

*** Schistochila nobilis (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877. Bas.: Jungermannia nobilis Hook., Musci Exot. 1: tab. 11, 1818.

*** Schistochila nuda Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 215, 1934.

*** Schistochila pachyphylla (Lehm.) Steph., Sp. Hepat. (Stephani) 4: 99, 1910. Bas.: Jungermannia pachyphylla Lehm., Nov. Stirp. Pug. 6: 61, 1834.

*** Schistochila papillifera R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 27, 1968.

*** Schistochila parvistipula Rodway, Pap. & Proc. Roy. Soc. Tasmania 1916: 47, 1917.

*** Schistochila pellucida R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 286, 1985.

*** Schistochila piligera Steph., Bot. Gaz. 15 (11): 291, 1890.

*** Schistochila pinnatifolia (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877. Bas.: Jungermannia pinnatifolia Hook., Musci Exot. 2: tab. 114, 1820.

*** Schistochila pluriciliata R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 395, 1985.

*** Schistochila pseudociliata R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (4): 623, 1971.

*** Schistochila quadrifida A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892.

** Schistochila ramentacea Steph., Sp. Hepat. (Stephani) 6: 494, 1924.

*** Schistochila reflexa (Mont.) Steph., Sp. Hepat. (Stephani) 4: 97, 1910. Bas.: Gottschea reflexa Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 347, 1845.

*** Schistochila reflexistipula J.J.Engel et R.M.Schust., Phytologia 30 (4): 245, 1975.

*** Schistochila reinwardtii (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 218, 1898. Bas.: Jungermannia reinwardtii Nees, Enum. Pl. Crypt. Javae: 66, 1830.

*** Schistochila repleta (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 4: 90, 1909. Bas.: Jungermannia repleta Hook.f. et Taylor, London J. Bot. 3: 557, 1844.

** Schistochila rubriseta Steph., Sp. Hepat. (Stephani) 4: 78, 1909.

** Schistochila schultzei Steph., Sp. Hepat. (Stephani) 6: 494, 1924.

*** Schistochila sciophila R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (4): 632, 1971.

*** Schistochila sciurea (Nees) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 251, 1893. Bas.: Jungermannia sciurea Nees, Enum. Pl. Crypt. Javae: 34, 1830.

*** Schistochila simulans (C.Massal.) Xiao L.He et Yu Sun, Polish Bot. J. 58 (2): 473, 2013. Bas.: Cephalozia simulans C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 236, 1885.

*** Schistochila spegazziniana (C.Massal.) Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 60, 1900. Bas.: Gottschea spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 206, 1885.

*** Schistochila sphagnoides (Schwägr.) Lindb. ex Steph., Sp. Hepat. (Stephani) 4: 70, 1909. Bas.: Jungermannia sphagnoides Schwägr., Hist. Musc. Hepat. Prodr.: 23, 1814.

*** Schistochila splachnophylla (Hook.f. et Taylor) Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 28, 1901. Bas.: Jungermannia splachnophylla Hook.f. et Taylor, London J. Bot. 3: 455, 1844.

** Schistochila stratosa (Mont.) A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892. Bas.: Gottschea stratosa Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 346, 1845.

*** Schistochila subhyalina R.M.Schust., Phytologia 30 (4): 246, 1975.

** Schistochila subhyalina var. grandidentata (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010. Bas.: Pachyschistochila subhyalina var. grandidentata J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 58: 507, 1985.

*** Schistochila subimmersa J.J.Engel et R.M.Schust., Phytologia 30 (4): 247, 1975.

*** Schistochila succulenta (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010. Bas.: Pachyschistochila succulenta J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 58: 517, 1985.

*** Schistochila tasmanica Steph., Sp. Hepat. (Stephani) 4: 86, 1909.

*** Schistochila trispiralis R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 28, 1968.

*** Schistochila tuloides (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 4: 89, 1909. Bas.: Jungermannia tuloides Hook.f. et Taylor, London J. Bot. 3: 558, 1844.

*** Schistochila undulatifolia Piippo, Ann. Bot. Fenn. 23 (1): 8, 1986.

*** Schistochila virescens R.M.Schust., Phytologia 30 (4): 248, 1975.

*** Schistochila vitreocincta (Herzog) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010. Bas.: Perssoniella vitreocincta Herzog, Ark. Bot. (n.ser.) 2 (4): 265, 1952.

** Schistochila volans Grolle, J. Hattori Bot. Lab. 29: 244, 1966.

*** Schistochila yakushimensis N.Ohnishi et Deguchi, Bryologist 106 (3): 451, 2003.

** Schistochila zantenii Grolle, J. Hattori Bot. Lab. 29: 243, 1966.

*** Frullaniaceae Lorch

by M. von Konrat, J. Hentschel, J. Uribe, P. Sukkharak, J. Heinrichs, J. Larraín, R. Stotler and L. Zhang

The subgeneric treatment of Frullaniaceae follows, which includes a slight departure from formal infrageneric ranks where they apply clades referred to as Diastaloba I, II, III, and IV that appear to represent distinct subgenera.

*** Frullania Raddi, Jungermanniogr. Etrusca: 9, 1818.

*** subg. Chonanthelia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 8, 1884.

* Frullania flammea Taylor, Trans. & Proc. Bot. Soc. Edinburgh 15: 29, 1884.

*** Frullania lindmanii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 19, 1897.

** Frullania paranensis Steph., Sp. Hepat. (Stephani) 4: 607, 1911.

** Frullania spegazzinii M.E.Reiner, Bol. Soc. Argent. Bot. 25 (3/4): 310, 1988.

*** sect. Chonanthelia (Spruce) Yuzawa ex Hentschel et von Konrat, Phytotaxa 220 (2): 129, 2015. Bas.: Frullania subg. Chonanthelia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 8, 1884.

*** Frullania gibbosa Nees, Syn. Hepat. 3: 411, 1845.

*** sect. Cladocarpicae Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 43, 1884.

*** Frullania beauverdii Steph., Biblioth. Bot. 87 (2): 241, 1916.

*** Frullania blepharozia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 18, 1884.

*** Frullania bogotensis Steph., Sp. Hepat. (Stephani) 4: 327, 1910.

* Frullania brachycarpa Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cciv, 1889 [1890].

*** Frullania confertiloba Steph., Sp. Hepat. (Stephani) 4: 326, 1910.

*** Frullania decidua Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 30, 1884.

*** Frullania ecklonii (Spreng.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 413, 1845. Bas.: Jungermannia ecklonii Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 324, 1827.

* Frullania ecklonii var. robustior (Gottsche, Lindenb. et Nees) Sim, Trans. Roy. Soc. South Africa 15 (1): 39, 1926. Bas.: Frullania ecklonii a robustior Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 771, 1847.

* Frullania ecklonii var. rufescens Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 771, 1847.

* Frullania ecklonii var. tenerior (Gottsche, Lindenb. et Nees) Sim, Trans. Roy. Soc. South Africa 15 (1): 39, 1926. Bas.: Frullania ecklonii b tenerior Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 771, 1847.

*** Frullania holostipula S.Hatt. et D.G.Griffin, Misc. Bryol. Lichenol. 8 (3): 47, 1978 (Hattori and Griffin 1978).

*** Frullania megalostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 15, 1884.

*** Frullania obscura (Sw.) Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 333, 1840. Bas.: Jungermannia obscura Sw., Fl. Ind. Occid. 3: 1869, 1806.

*** Frullania obscura var. spiniloba (Steph.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015. Bas.: Frullania spiniloba Steph., Sp. Hepat. (Stephani) 4: 336, 1910.

*** Frullania ringens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 17, 1884.

*** Frullania rio-janeirensis (Raddi) Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 88, 1876 [1877]. Bas.: Frullanoides rio-janeirensis Raddi, Critt. Brasil.: 13, 1822.

*** Frullania sphaerocephala Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 17, 1884.

** Frullania tunguraguana L.Clark et Frye, Bryologist 55 (2): 133, 1952. Nom. nov. pro Frullania brachyclada Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 15, 1884, nom. illeg.

*** sect. Pluricarinatae (Yuzawa, Mues et S.Hatt.) Hentschel et von Konrat, Phytotaxa 220 (2): 129, 2015. Bas.: Frullania ser. Pluricarinatae Yuzawa, Mues et S.Hatt., J. Hattori Bot. Lab. 63: 428, 1987.

*** Frullania albertii Steph., Biblioth. Bot. 87 (2): 237, 1916.

*** Frullania arsenii Steph., Sp. Hepat. (Stephani) 6: 530, 1924.

** Frullania bonariensis M.E.Reiner, Bol. Soc. Argent. Bot. 25 (3/4): 313, 1988.

*** Frullania cuencensis Taylor, London J. Bot. 5: 406, 1846.

*** Frullania depressa Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 168, 1863.

*** Frullania dusenii Steph., Arch. Mus. Nac. Rio de Janeiro 13: 115 (9), 1905.

*** Frullania gradsteinii Yuzawa, Mues et S.Hatt., J. Hattori Bot. Lab. 63: 429, 1987.

*** Frullania haematocysta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 54, 1884.

*** Frullania jelskii Loitl., Diagn. pl. nov.: 18, 1894.

*** Frullania laxiflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 26, 1884.

* Frullania laxiflora var. crossii Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 27, 1884.

*** Frullania neurota Taylor, London J. Bot. 5: 400, 1846.

*** Frullania planifolia Steph., Sp. Hepat. (Stephani) 4: 337, 1910.

*** Frullania pluricarinata Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 168, 1864.

** Frullania sandvicensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 28, 1872.

*** Frullania standaertii Steph., Sp. Hepat. (Stephani) 4: 342, 1910.

*** Frullania stenostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 29, 1884.

*** Frullania tetraptera Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 47, 1838.

** Frullania trinervis (Lehm.) Drège, Flora, Beig. 26: 186, 1843. Bas.: Jungermannia trinervis Lehm., Linnaea 9 (4): 426, 1835. 186

*** Frullania winteri Steph., Sp. Hepat. (Stephani) 4: 338, 1910.

*** Frullania winteri var. vanderhammenii (Haarbrink) Yuzawa, J. Hattori Bot. Lab. 70: 233, 1991. Bas.: Frullania vanderhammenii Haarbrink, Lindbergia 7 (1): 56, 1981.

*** subg. Diastaloba Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 55, 1884.

** Frullania antaresensis S.Hatt., J. Hattori Bot. Lab. 47: 92, 1980.

* Frullania armatifolia Verd., Bull. Jard. Bot. Buitenzorg (sér. 3) 12 (1): 61, 1932.

*** Frullania curvilobula Schäf.-Verw., D.F.Peralta et S.M.Siqueira, Phytotaxa 57 (4): 27, 2012.

** Frullania gracilicaulis S.Hatt., J. Hattori Bot. Lab. 43: 421, 1977 [1978].

** Frullania humbertii Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 24, 1976.

** Frullania hypoleucula S.Hatt., J. Hattori Bot. Lab. 57: 413, 1984.

** Frullania incurva S.Hatt., J. Hattori Bot. Lab. 65: 431, 1988.

** Frullania klotzschii Nees, Sp. Hepat. (Stephani) 4: 558, 1911.

** Frullania letestui Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 38, 1976.

* Frullania miradorensis Lindenb. et Gottsche, Syn. Hepat. 5: 781, 1847.

* Frullania odontostipa Spruce, Mem. Torrey Bot. Club 1 (3): 120, 1890.

*** Frullania pilibracteola S.Hatt., J. Hattori Bot. Lab. 43: 428, 1977 [1978].

*** Frullania pilistipula Steph., Sp. Hepat. (Stephani) 4: 648, 1911.

*** Frullania ramuligera (Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 14, 1842. Bas.: Jungermannia ramuligera Nees, Enum. Pl. Crypt. Javae: 52, 1830.

** Frullania subpilibracteola S.Hatt., J. Hattori Bot. Lab. 43: 434, 1977 [1978].

** Frullania subtilissima (Nees ex Mont.) Lindenb., Syn. Hepat. 3: 443, 1845. Bas.: Frullania atrata β subtilissima Nees ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 333, 1840.

** Frullania vandenberghenii Pócs, Acta Bot. Acad. Sci. Hung. 25 (3/4): 229, 1979 [1980]. Nom. nov. pro Frullania epiphylla Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 30, 1976, nom. illeg.

grp. Diastaloba I

** Frullania akiyamae S.Hatt., J. Hattori Bot. Lab. 60: 240, 1986.

** Frullania apiculata (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 452, 1845. Bas.: Jungermannia apiculata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 222, 1824 [1825].

** Frullania apiculata var. goebelii Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 222, 1893.

*** Frullania armitiana Steph., Sp. Hepat. (Stephani) 4: 538, 1911.

** Frullania armitiana var. inflexula S.Hatt., J. Hattori Bot. Lab. 65: 415, 1988.

** Frullania aterrima (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 450, 1845. Bas.: Jungermannia aterrima Hook.f. et Taylor, London J. Bot. 3: 395, 1844.

*** Frullania attenuata Steph., Sp. Hepat. (Stephani) 4: 538, 1911.

*** Frullania bella Steph., Sp. Hepat. (Stephani) 4: 643, 1911.

** Frullania changii S.Hatt. et C.Gao, J. Jap. Bot. 60 (1): 1, 1985.

* Frullania claviloba Steph., Sp. Hepat. (Stephani) 4: 651, 1911. 187

*** Frullania colliculosa von Konrat, Braggins, Hentschel et Heinrichs, Nova Hedwigia 91 (3/4): 494, 2010.

*** Frullania cordistipula (Reinw., Blume et Nees) Nees, Voy. Amér. Mérid. 7 (2): 68, 1839. Bas.: Jungermannia cordistipula Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 220, 1824 [1825]. 188

** Frullania cordistipula var. dentistipula S.Hatt., J. Hattori Bot. Lab. 60: 242, 1986.

* Frullania cordistipula var. mutica (Gottsche, Lindenb. et Nees) Schiffn., Consp. Hepat. Arch. Ind.: 323, 1898. Bas.: Frullania cordistipula β mutica Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 454, 1845.

* Frullania crenatiloba Steph., Sp. Hepat. (Stephani) 6: 551, 1924. 189

*** Frullania curvistipula Steph., Sp. Hepat. (Stephani) 4: 548, 1911.

** Frullania curvistipula var. falcatidentata S.Hatt., Misc. Bryol. Lichenol. 9 (6): 124, 1982.

** Frullania curvistipula var. lamii Verd., Ann. Bryol., Suppl. 1: 91, 1930.

** Frullania cuspidifolia Steph., Sp. Hepat. (Stephani) 4: 543, 1911.

* Frullania degelii S.W.Arnell, Svensk Bot. Tidskr. 53 (4): 503, 1959.

** Frullania dentifera S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 102, 1985.

*** Frullania dentiloba S.Hatt., J. Jap. Bot. 50 (6): 161, 1975.

** Frullania durifolia Steph., Hedwigia 33 (3): 162, 1894.

** Frullania exilis Taylor, London J. Bot. 5: 405, 1846.

** Frullania gabonensis Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 81, 1976.

*** Frullania gracilis (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 452, 1845. Bas.: Jungermannia gracilis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 221, 1824 [1825].

* Frullania gracilis var. brevior (Gottsche, Lindenb. et Nees) Schiffn., Consp. Hepat. Arch. Ind.: 327, 1898. Bas.: Frullania gracilis β brevior Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 453, 1845.

** Frullania gracilis var. vittata S.Hatt., J. Hattori Bot. Lab. 60: 243, 1986.

** Frullania gracilis subsp. zennoskei S.Hatt. et Thaithong, J. Hattori Bot. Lab. 44: 183, 1978.

*** Frullania hasskarliana Lindenb., Syn. Hepat. 3: 453, 1845.

** Frullania hasskarliana var. gracilis S.Hatt., J. Hattori Bot. Lab. 60: 243, 1986.

** Frullania hasskarliana var. parvidentata S.Hatt., J. Hattori Bot. Lab. 60: 245, 1986.

*** Frullania hattorii von Konrat et Braggins, New Zealand J. Bot. 41 (1): 56, 2003.

*** Frullania hodgsoniae von Konrat, Braggins, Hentschel et Heinrichs, Nova Hedwigia 91 (3/4): 492, 2010. Nom. nov. pro Frullania aterrima var. lepida E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 386, 1949.

** Frullania hottana S.Hatt., J. Hattori Bot. Lab. 40: 479, 1976.

*** Frullania inconstans Verd., Ann. Bryol., Suppl. 1: 83, 1930.

** Frullania inconstans var. grossedentata Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 530, 1973.

*** Frullania johnsonii Steph., Hedwigia 33 (3): 163, 1894.

** Frullania macgregorii Steph., Hedwigia 33 (3): 154, 1894.

** Frullania macgregorii var. rostellula (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 220, 1982. Bas.: Frullania reimersii var. rostellula S.Hatt., J. Hattori Bot. Lab. 38: 258, 1974.

** Frullania madens Steph., Sp. Hepat. (Stephani) 6: 553, 1924.

** Frullania mehrana S.Hatt., Recent Adv. Bot.: 66, 1976.

** Frullania motoyana Steph., Sp. Hepat. (Stephani) 4: 646, 1911.

* Frullania multilacera Steph., Sp. Hepat. (Stephani) 4: 650, 1911. 190

** Frullania multilacera subsp. gracilior S.Hatt., Mem. New York Bot. Gard. 45: 547, 1987.

** Frullania multilacera var. lacerissima S.Hatt., J. Hattori Bot. Lab. 39: 294, 1975.

** Frullania multilaceroides S.Hatt., Mem. New York Bot. Gard. 45: 549, 1987.

** Frullania neosheana S.Hatt., J. Hattori Bot. Lab. 45: 350, 1979.

** Frullania papillata Steph., Sp. Hepat. (Stephani) 4: 615, 1911.

** Frullania pulogensis Steph., Sp. Hepat. (Stephani) 4: 545, 1911.

*** Frullania purpurea Steph., Sp. Hepat. (Stephani) 4: 626, 1911.

** Frullania reimersii Verd., Ann. Bryol., Suppl. 1: 84, 1930.

** Frullania saepisdentata S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 116, 1985.

** Frullania schiffneri Verd., Ann. Bryol. 2: 150, 1929.

** Frullania schusterana S.Hatt., J. Hattori Bot. Lab. 36: 411, 1972 [1973].

** Frullania seriatifolia Steph., Hedwigia 33 (3): 167, 1894.

** Frullania serrata Gottsche, Syn. Hepat. 3: 453, 1845.

** Frullania serrata var. ceramensis S.Hatt., J. Hattori Bot. Lab. 60: 247, 1986.

** Frullania serrata subsp. grolleana (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 225, 1982. Bas.: Frullania grolleana S.Hatt., J. Hattori Bot. Lab. 36: 416, 1972 [1973].

** Frullania serrata var. hamatispina (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 225, 1982. Bas.: Frullania serrata subsp. hamatispina S.Hatt., J. Hattori Bot. Lab. 38: 262, 1974.

** Frullania serrata var. pertenuis (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 342, 1898. Bas.: Jungermannia cordistipula γ pertenuis Nees, Enum. Pl. Crypt. Javae: 49, 1830.

** Frullania serrata subsp. spinistipula S.Hatt., J. Hattori Bot. Lab. 51: 225, 1982. Nom. nov. pro Frullania spinistipula S.Hatt., J. Hattori Bot. Lab. 36: 413, 1972 [1973], nom. illeg.

** Frullania setacea S.Hatt., J. Hattori Bot. Lab. 65: 447, 1988.

** Frullania sheana S.Hatt., J. Hattori Bot. Lab. 45: 356, 1979.

** Frullania simmondsii Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 109, 1914.

*** Frullania sinuata Sande Lac., Ned. Kruidk. Arch. 3: 424, 1854 [1855].

** Frullania steereana S.Hatt., Mem. New York Bot. Gard. 45: 553, 1987.

** Frullania stipatiloba Steph., Hedwigia 33 (3): 168, 1894.

** Frullania subdentata Steph., Sp. Hepat. (Stephani) 4: 545, 1911.

** Frullania subdentata var. latistipula (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 228, 1982. Bas.: Frullania curvistipula var. latistipula S.Hatt., J. Hattori Bot. Lab. 44: 530, 1978.

** Frullania submultilacera S.Hatt. et Koike, J. Hattori Bot. Lab. 75: 190, 1994.

** Frullania subocellata S.Hatt., J. Hattori Bot. Lab. 60: 248, 1986.

*** Frullania taxodiocola R.M.Schust., Phytologia 53 (5): 364, 1983.

*** Frullania ternatensis Gottsche, Syn. Hepat. 4: 465, 1846.

** Frullania ternatensis var. non-appendiculata S.Hatt., J. Hattori Bot. Lab. 38: 174, 1974.

*** Frullania trichodes Mitt., Bonplandia 10 (2): 19, 1862.

** Frullania vaga Mitt., Fl. vit.: 418, 1871 [1873].

*** Frullania vaginata (Sw.) Nees, Syn. Hepat. 4: 465, 1846. Bas.: Jungermannia vaginata Sw., Meth. Musc.: 35, 1781.

* Frullania vaginata var. nigricans (Gottsche, Lindenb. et Nees) Schiffn., Consp. Hepat. Arch. Ind.: 348, 1898. Bas.: Frullania vaginata β nigricans Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 465, 1846.

* Frullania van-zantenii Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 528, 1973. 191

** Frullania venusta S.Hatt., J. Hattori Bot. Lab. 38: 217, 1974.

** Frullania verdoorniana S.Hatt., J. Hattori Bot. Lab. 37: 122, 1973.

*** Frullania vitalii Yuzawa et S.Hatt., J. Jap. Bot. 63 (1): 30, 1988.

** Frullania vittata S.Hatt., J. Hattori Bot. Lab. 38: 270, 1974.

** Frullania vivipara Pócs, Fieldiana, Bot. (n.ser.) 47: 151, 2008.

** Frullania wairua von Konrat et Braggins, New Zealand J. Bot. 43 (4): 886, 2005.

** Frullania warnckeana S.Hatt., J. Hattori Bot. Lab. 38: 213, 1974.

** Frullania warnckeana var. dentosa S.Hatt., Misc. Bryol. Lichenol. 7 (8): 162, 1977.

grp. Diastaloba II

*** Frullania baumannii S.Hatt., J. Hattori Bot. Lab. 43: 410, 1977 [1978].

*** Frullania congesta Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 451, 1845. Nom. nov. pro Jungermannia congesta Hook.f. et Taylor, London J. Bot. 3: 396, 1844, nom. illeg.

** Frullania ocellata S.Hatt. et Kamim., J. Hattori Bot. Lab. 37: 531, 1973.

*** Frullania ptychantha Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 257, 1843.

*** Frullania repandistipula Sande Lac., Ned. Kruidk. Arch. 3: 422, 1854 [1855].

** Frullania repandistipula subsp. queenslandica S.Hatt., Mem. New York Bot. Gard. 45: 550, 1987.

** Frullania repandistipula subsp. spinibractea S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 150, 1975.

grp. Diastaloba III

** Frullania grossiclava Steph., Sp. Hepat. (Stephani) 4: 384, 1910.

** Frullania loricata Pearson, Forh. Vidensk.-Selsk. Kristiania 1890 (2): 6, 1891.

* Frullania loricata var. laxa Pearson, Forh. Vidensk.-Selsk. Kristiania 1890 (2): 8, 1891.

*** Frullania usambarana Schiffn., Hedwigia 33 (3): 160, 1894.

** Frullania usambarana var. reducta Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 91, 1976.

grp. Diastaloba IV

* Frullania brunea (Spreng.) Drège, Flora, Beig. 26: 186, 1843. Bas.: Jungermannia brunea Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827. 192

*** Frullania caulisequa (Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 12: 51, 1839. Bas.: Jungermannia caulisequa Nees, Fl. Bras. (Martius) 1 (1): 373, 1833.

*** Frullania grossifolia Steph., Sp. Hepat. (Stephani) 4: 633, 1911.

*** Frullania hypoleuca Nees, Observ. bot.: 470, 1843.

*** Frullania lindenbergii Lehm., Nov. Stirp. Pug. 8: 17, 1844.

* Frullania lindenbergii var. fusca Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 780, 1847.

** Frullania ponapensis S.Hatt. et Koike, J. Hattori Bot. Lab. 75: 186, 1994.

* Frullania tricarinata Sande Lac., Plagiochila Sandei: 10, 1856. 193

* subg. Diversitextae (Kamim.) S.Hatt., J. Hattori Bot. Lab. 59: 154, 1985. Bas.: Frullania subsect. Diversitextae Kamim., J. Hattori Bot. Lab. 24: 80, 1961.

*** Frullania diversitexta Steph., Bull. Herb. Boissier 5 (2): 89, 1897.

*** subg. Frullania

* Frullania amamiensis Kamim., Bull. Kochi Gakuen Jun. Coll. 1: 51, 1970.

** Frullania amplicrania Steph., Sp. Hepat. (Stephani) 4: 404, 1910.

*** Frullania ampullifera J.B.Jack et Steph., Hedwigia 33 (3): 139, 1894.

*** Frullania anderssonii Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 144, 1873.

** Frullania angustistipa Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 297, 1907.

** Frullania aposinensis S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 131, 1985. Nom. nov. pro Frullania chinensis Steph., Sp. Hepat. (Stephani) 4: 469, 1911, nom. illeg.

** Frullania appendistipula S.Hatt., J. Hattori Bot. Lab. 36: 424, 1972 [1973].

** Frullania appendistipula var. spinifera S.Hatt., J. Hattori Bot. Lab. 38: 226, 1974.

** Frullania auriculata S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 11 (1): 11, 1985.

** Frullania benjaminiana Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 109, 1975.

** Frullania bergmanii S.Hatt., J. Hattori Bot. Lab. 38: 192, 1974.

*** Frullania berthoumieui Steph., Hedwigia 33 (3): 140, 1894.

** Frullania bhutanensis S.Hatt., Fl. E. Himalaya 2: 232, 1971.

** Frullania blastopetala S.Hatt., J. Hattori Bot. Lab. 57: 407, 1984.

*** Frullania bonincola S.Hatt., J. Hattori Bot. Lab. 44: 551, 1978. Nom. nov. pro Frullania viridis Horik., Sci. Rep. Tôhoku Imp. Univ., Ser. 4, Biol. 5 (4): 646, 1929 [1930], nom. illeg.

* Frullania brevicalycina Steph., Hedwigia 33 (3): 141, 1894.

** Frullania brittoniae A.Evans, Trans. Connecticut Acad. Arts 10 (1): 15, 1899.

*** Frullania bullata Steph., Sp. Hepat. (Stephani) 4: 371, 1910.

*** Frullania caffraria Steph., Hedwigia 33 (3): 141, 1894.

*** Frullania calcarata Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 137, 1873.

** Frullania carrii Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 520, 1973.

** Frullania chenii S.Hatt. et P.J.Lin, J. Jap. Bot. 60 (4): 106, 1985.

** Frullania chilensis Steph., Hedwigia 33 (3): 142, 1894.

*** Frullania chodatii Beauverd, Sp. Hepat. (Stephani) 6: 533, 1924. Based on: Frullania grandistipula Steph., Sp. Hepat. (Stephani) 6: 533, 1924, nom. inval.

*** Frullania cobrensis Gottsche, Hedwigia 33 (3): 142, 1894.

** Frullania consociata Steph., Sp. Hepat. (Stephani) 4: 461, 1911.

* Frullania contracta Steph., Sp. Hepat. (Stephani) 4: 469, 1911. 194

* Frullania cornuta Steph., Sp. Hepat. (Stephani) 4: 467, 1911. 195

*** Frullania crassitexta Steph., Sp. Hepat. (Stephani) 4: 423, 1910.

*** Frullania crispiplicata Yuzawa et S.Hatt., J. Jap. Bot. 58 (2): 43, 1983.

** Frullania cristata S.Hatt., J. Hattori Bot. Lab. 49: 165, 1981.

* Frullania cuneiloba Nees, Syn. Hepat. 3: 427, 1845.

** Frullania cyparioides (Schwägr.) Nees, Naturgesch. Eur. Leberm. 3: 210, 1838. Bas.: Jungermannia cyparioides Schwägr., Hist. Musc. Hepat. Prodr.: 14, 1814.

*** Frullania davurica Hampe ex Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 422, 1845.

** Frullania debilis Steph. ex S.Hatt., J. Hattori Bot. Lab. 38: 190, 1974.

** Frullania deppii Lehm., Nov. Stirp. Pug. 8: 15, 1844.

** Frullania diptera (Lehm.) Drège, Flora, Beig. 26: 186, 1843. Bas.: Jungermannia diptera Lehm., Linnaea 9 (4): 425, 1835.

*** Frullania duthiana Steph., Sp. Hepat. (Stephani) 4: 351, 1910.

** Frullania duthiana var. appendiculata S.Hatt., Fl. E. Himalaya 2: 234, 1971.

** Frullania duthiana var. laevis S.Hatt., Bull. Univ. Mus. Univ. Tokyo 8: 233, 1975.

** Frullania duthiana var. szechuanensis S.Hatt. et C.Gao, J. Jap. Bot. 60 (1): 2, 1985.

** Frullania echinantha S.Hatt., J. Hattori Bot. Lab. 38: 233, 1974.

** Frullania echinatella S.Hatt., J. Hattori Bot. Lab. 65: 423, 1988.

* Frullania elegans Lehm., Nov. Stirp. Pug. 10: 16, 1857.

*** Frullania elephantum S.Hatt., J. Hattori Bot. Lab. 43: 416, 1977 [1978].

** Frullania epiphylla S.Hatt., J. Hattori Bot. Lab. 38: 235, 1974.

* Frullania epiphylla subsp. fijiensis S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 11 (1): 14, 1985.

** Frullania erostrata S.Hatt., J. Hattori Bot. Lab. 38: 237, 1974.

* Frullania esenbeckiana Beauverd, Sp. Hepat. (Stephani) 6: 570, 1924. Nom. nov. pro Frullania grossiloba Steph., Sp. Hepat. (Stephani) 4: 531, 1911, nom. illeg.

** Frullania evelynae S.Hatt. et Thaithong, J. Hattori Bot. Lab. 44: 179, 1978.

* Frullania evelynae var. devendrae Sushil K.Singh et Barbhuiya, Taiwania 57 (2): 109, 2012.

* Frullania evelynae var. srivastavae Sushil K.Singh et Barbhuiya, Taiwania 57 (2): 112, 2012.

*** Frullania eymae S.Hatt., J. Hattori Bot. Lab. 39: 284, 1975.

** Frullania eymae var. crispidentata S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 105, 1985.

*** Frullania falciloba Lehm., Nov. Stirp. Pug. 8: 20, 1844.

** Frullania falsicornuta S.Hatt., J. Hattori Bot. Lab. 60: 212, 1986.

** Frullania fauriana Steph., Hedwigia 33 (3): 144, 1894.

** Frullania fengyangshanensis R.L.Zhu et M.L.So, Bryologist 100 (3): 356, 1997.

** Frullania ferdinandi-muelleri Steph., Sp. Hepat. (Stephani) 4: 417, 1910.

* Frullania flexuosa S.Hatt., J. Hattori Bot. Lab. 54: 146, 1983. 196

** Frullania fuscovirens Steph., Sp. Hepat. (Stephani) 4: 401, 1910.

** Frullania fuscovirens var. gemmipara (R.M.Schust. et S.Hatt.) S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 135, 1985. Bas.: Frullania gemmipara R.M.Schust. et S.Hatt., J. Hattori Bot. Lab. 44: 547, 1978.

** Frullania gaoligongensis X.L.Bai et C.Gao, Hikobia 13 (1): 87, 1999.

** Frullania gemmulosa S.Hatt. et Thaithong, J. Hattori Bot. Lab. 43: 449, 1977 [1978].

*** Frullania gigantea Steph., Sp. Hepat. (Stephani) 4: 467, 1911.

** Frullania giraldiana C.Massal., Hepat. Shen-si: 41, 1897. 197

** Frullania giraldiana var. handelii (Verd.) S.Hatt., J. Hattori Bot. Lab. 36: 123, 1972 [1973]. Bas.: Frullania nepalensis var. handelii Verd., Symb. Sin. 5: 41, 1930.

** Frullania globosa S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 107, 1985.

** Frullania grandilobula S.Hatt. et Piippo, Acta Bot. Fenn. 133: 45, 1986.

*** Frullania grandistipula Lindenb., Syn. Hepat. 3: 430, 1845.

** Frullania hainanensis S.Hatt. et P.J.Lin, J. Jap. Bot. 61 (10): 307, 1986.

** Frullania hamatiloba Steph., Sp. Hepat. (Stephani) 4: 400, 1910.

** Frullania handelii Verd., Symb. Sin. 5: 36, 1930.

** Frullania handel-mazzettii S.Hatt., J. Hattori Bot. Lab. 49: 150, 1981.

** Frullania hattoriantha Udar et V.Nath, Misc. Bryol. Lichenol. 9 (2): 44, 1981.

* Frullania hebridensis Steph., Sp. Hepat. (Stephani) 4: 469, 1911. 198

** Frullania hicksiae S.Hatt., Cryptog. Bryol. Lichénol. 5 (1/2): 182, 1984.

** Frullania higuchii Yuzawa, Koike et S.Hatt., J. Hattori Bot. Lab. 75: 194, 1994.

** Frullania hiroshii S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 6 (1): 34, 1980.

* Frullania hirtiflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 35, 1884.

*** Frullania howeana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 107, 1914.

** Frullania ignatovii Sofronova, Mamontov et Potemkin, Novosti Sist. Nizš. Rast. 47: 335, 2013.

** Frullania incisoduthiana S.Hatt., J. Hattori Bot. Lab. 46: 391, 1979.

* Frullania incisoduthiana var. parva S.Hatt., J. Hattori Bot. Lab. 46: 391, 1979.

* Frullania incisostipula Steph., Sp. Hepat. (Stephani) 6: 541, 1924.

** Frullania inflexiloba S.Hatt., J. Hattori Bot. Lab. 57: 415, 1984.

** Frullania inouei S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 6 (1): 36, 1980.

*** Frullania irregularis S.Hatt. et Piippo, Acta Bot. Fenn. 133: 46, 1986.

*** Frullania jackii Gottsche, Hepat. Eur., Leberm. 29-30: no. 294, 1863.

** Frullania jacobsii S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12 (4): 129, 1986.

* Frullania jacquinotii Gottsche, J. Bot. (Morot) 12: 138, 1898.

** Frullania jovetiana von Konrat et Hentschel, Phytotaxa 220 (2): 135, 2015. Based on: Frullania pseudericoides S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12 (4): 132, 1986, nom. illeg.

** Frullania kagoshimensis Steph., Sp. Hepat. (Stephani) 4: 353, 1910.

** Frullania kagoshimensis subsp. hunanensis (S.Hatt.) S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 137, 1985. Bas.: Frullania hunanensis S.Hatt., J. Hattori Bot. Lab. 49: 152, 1981.

** Frullania kalimantanensis S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12 (4): 127, 1986.

** Frullania kashyapii Verd., Ann. Bryol. 5: 162, 1932.

** Frullania kitagawana S.Hatt., J. Hattori Bot. Lab. 57: 417, 1984.

** Frullania laeviperiantha X.L.Bai et C.Gao, Phytotaxa 220 (2): 136, 2015. Based on: Frullania laeviperiantha X.L.Bai et C.Gao, Nova Hedwigia 70 (1/2): 135, 2000, nom. inval.

* Frullania lancistyla Steph., Sp. Hepat. (Stephani) 4: 389, 1910.

* Frullania latiflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 34, 1884.

** Frullania latogaleata Herzog, Trans. Brit. Bryol. Soc. 1 (3): 189, 1949.

** Frullania lepida S.Hatt. et Piippo, Acta Bot. Fenn. 133: 48, 1986.

* Frullania levieri Steph., Sp. Hepat. (Stephani) 4: 388, 1910.

** Frullania linii S.Hatt., J. Hattori Bot. Lab. 49: 155, 1981.

* Frullania longistyla Yuzawa et S.Hatt., J. Jap. Bot. 63 (11): 361, 1988. 199

* Frullania ludoviciae Steph., Rev. Bryol. 35 (2): 29, 1908. 200

** Frullania lushanensis S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 137, 1985.

* Frullania macularis Taylor, London J. Bot. 5: 403, 1846.

** Frullania maymyoensis Svihla, Bryologist 61 (4): 376, 1958 [1959].

** Frullania microauriculata Verd., Ann. Bryol. 2: 126, 1929.

** Frullania microauriculata var. rotundior Verd., Ann. Jard. Bot. Buitenzorg 40: 141, 1929.

** Frullania microrhyncha L.Clark et Svihla, Bryologist 53 (1): 63, 1950.

** Frullania mizoramensis Sushil K.Singh et Barbhuiya, Taiwania 57 (2): 106, 2012.

* Frullania montana Steph., Sp. Hepat. (Stephani) 4: 455, 1911. 201

** Frullania multituberculata Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015. Nom. nov. pro Frullania kalimantanensis Piippo et S.Hatt., J. Hattori Bot. Lab. 72: 117, 1992, nom. illeg.

** Frullania mutilata Steph., Sp. Hepat. (Stephani) 4: 673, 1911.

** Frullania nadeaudii Steph., Sp. Hepat. (Stephani) 4: 465, 1911.

*** Frullania nepalensis (Spreng.) Lehm. et Lindenb., Syn. Hepat. 3: 422, 1845. Bas.: Jungermannia nepalensis Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 324, 1827. 202

*** Frullania nicholsonii E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 368, 1949. Nom. nov. pro Frullania berggrenii W.E.Nicholson, Bryologist 28 (2): 17, 1925, nom. illeg.

** Frullania nigricaulis (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 457, 1845. Bas.: Jungermannia nigricaulis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 225, 1824 [1825].

* Frullania nigricaulis var. elongata Verd., Ann. Bryol., Suppl. 1: 155, 1930.

** Frullania nivimontana S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 8 (3): 95, 1982.

*** Frullania nobilis Steph., Hedwigia 33 (3): 154, 1894.

** Frullania nobilis var. cochleata (Steph.) S.Hatt., J. Hattori Bot. Lab. 59: 111, 1985. Bas.: Frullania cochleata Steph., Sp. Hepat. (Stephani) 4: 681, 1911.

** Frullania novocurvirostris S.Hatt., J. Hattori Bot. Lab. 49: 370, 1981. Nom. nov. pro Frullania curvirostris J.B.Jack et Steph., Hedwigia 33 (3): 143, 1894, nom. illeg.

** Frullania oahuensis Hampe ex Gottsche, Lindenb. et Nees, Observ. bot.: 471, 1843.

** Frullania obovata S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 8 (3): 97, 1982.

** Frullania obtusangula Hentschel et von Konrat, Phytotaxa 220 (2): 137, 2015.

** Frullania okinawensis Kamim., Misc. Bryol. Lichenol. 9 (4): 90, 1982.

** Frullania orbicularis Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 227, 1869.

*** Frullania orientalis Sande Lac., Plagiochila Sandei: 10, 1856.

** Frullania orinocensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 30, 1884.

*** Frullania ornithocephala (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 425, 1845. Bas.: Jungermannia ornithocephala Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 216, 1824 [1825].

* Frullania ornithocephala var. major (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 336, 1898. Bas.: Jungermannia ornithocephala β major Nees, Enum. Pl. Crypt. Javae: 47, 1830.

** Frullania ornithocephala var. pilosa Verd., Ann. Bryol. 2: 129, 1929.

** Frullania ornithocephala var. tuberculosa S.Hatt., J. Hattori Bot. Lab. 38: 179, 1974.

** Frullania pachyderma S.Hatt., J. Hattori Bot. Lab. 44: 525, 1978.

** Frullania pallidevirens Steph., Sp. Hepat. (Stephani) 4: 454, 1911.

** Frullania pariharii S.Hatt. et Thaithong, J. Jap. Bot. 53 (5): 130, 1978.

* Frullania parvifolia Steph., Sp. Hepat. (Stephani) 4: 354, 1910.

** Frullania pauciramea Steph., Sp. Hepat. (Stephani) 4: 458, 1911.

** Frullania pauciramea var. pauciramella S.Hatt. et Piippo, Acta Bot. Fenn. 133: 50, 1986.

** Frullania paucirameoides S.Hatt. et Piippo, Acta Bot. Fenn. 133: 51, 1986.

** Frullania pedicellata Steph., Bull. Herb. Boissier 5 (2): 90, 1897.

*** Frullania physantha Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 121, 1860 [1861].

** Frullania piptophylla S.Hatt., J. Hattori Bot. Lab. 47: 87, 1980.

** Frullania piptophylla var. minor S.Hatt., J. Hattori Bot. Lab. 60: 247, 1986.

** Frullania piptophylloides S.Hatt., J. Hattori Bot. Lab. 47: 90, 1980.

* Frullania plicata Hentschel et von Konrat, Phytotaxa 220 (2): 137, 2015. Nom. nov. pro Frullania acutiloba Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 477, 1947, nom. illeg.

** Frullania pocsantha Thaithong et S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 3 (4): 149, 1977.

** Frullania pran-nathii M.Dey et D.K.Singh, J. Jap. Bot. 83 (5): 281, 2008.

** Frullania pringlei Fulford et Sharp, Mem. New York Bot. Gard. 63: 45, 1990. Nom. nov. pro Frullania spicata Steph., Sp. Hepat. (Stephani) 4: 392, 1910, nom. illeg.

** Frullania prominula S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 112, 1985.

** Frullania propaginea S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 114, 1985.

* Frullania pseudericoides S.Hatt., J. Hattori Bot. Lab. 51: 256, 1982. Nom. nov. pro Frullania sharpii subsp. subrostrata S.Hatt., J. Hattori Bot. Lab. 38: 265, 1974.

** Frullania pseudoschensiana S.Hatt., J. Hattori Bot. Lab. 47: 101, 1980.

** Frullania pseudoschensiana var. darjeelingensis S.Hatt., J. Hattori Bot. Lab. 49: 149, 1981.

** Frullania pullei Verd., Nova Guinea 14: 542, 1930.

** Frullania pusilla Mitt., Fl. vit.: 417, 1871 [1873].

*** Frullania pycnantha (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 411, 1845. Bas.: Jungermannia pycnantha Hook.f. et Taylor, London J. Bot. 3: 566, 1844.

** Frullania queenslandica Steph., Sp. Hepat. (Stephani) 4: 424, 1910.

** Frullania recurvistipula S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 114, 1975.

*** Frullania reflexistipula Sande Lac., Ned. Kruidk. Arch. 3: 422, 1854 [1855].

** Frullania reflexistipula var. squarrosa S.Hatt. et Piippo, Acta Bot. Fenn. 133: 54, 1986.

** Frullania remotidens S.Hatt., J. Hattori Bot. Lab. 51: 257, 1982.

* Frullania reptans Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 161, 1855.

*** Frullania retusa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 119, 1860 [1861].

** Frullania retusa var. gymnantha S.Hatt. et Thaithong, J. Jap. Bot. 53 (5): 131, 1978.

** Frullania retusa var. hirsuta S.Hatt. et Thaithong, J. Hattori Bot. Lab. 44: 191, 1978.

*** Frullania rhystocolea Herzog, Symb. Sin. 5: 39, 1930.

** Frullania rhytidantha S.Hatt., J. Hattori Bot. Lab. 47: 97, 1980.

** Frullania rigida Steph., Sp. Hepat. (Stephani) 4: 371, 1910.

*** Frullania riparia Hampe, Nov. Stirp. Pug. 7: 14, 1838.

** Frullania rizalii Piippo et S.Hatt., J. Hattori Bot. Lab. 72: 119, 1992.

*** Frullania rostellata Mitt., Handb. N. Zeal. fl. 2: 755, 1867.

** Frullania rubella Gottsche, Hedwigia 28 (3): 159, 1889.

** Frullania rubella var. elongata (Steph.) S.Hatt., J. Hattori Bot. Lab. 54: 166, 1983. Bas.: Frullania elongata Steph., Sp. Hepat. (Stephani) 4: 423, 1910.

** Frullania rupicola Steph., Sp. Hepat. (Stephani) 6: 534, 1924.

** Frullania saipanensis S.Hatt. et Koike, J. Hattori Bot. Lab. 75: 188, 1994.

*** Frullania schensiana C.Massal., Hepat. Shen-si: 40, 1897.

** Frullania schusteri S.Hatt., Beih. Nova Hedwigia 90: 154, 1988.

** Frullania scottiana S.Hatt., Mem. New York Bot. Gard. 45: 551, 1987.

** Frullania setchellii Pearson, Univ. Calif. Publ. Bot. 10 (4): 326, 1923.

** Frullania shanensis Svihla, Bryologist 60 (4): 359, 1957.

** Frullania sharpantha Udar et Ad.Kumar, Misc. Bryol. Lichenol. 9 (9): 192, 1983.

** Frullania sharpii S.Hatt., J. Hattori Bot. Lab. 38: 180, 1974.

** Frullania sinensis Steph., Nuovo Giorn. Bot. Ital. (n.ser.) 13 (4): 349, 1906.

** Frullania sinosphaerantha S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 144, 1985.

** Frullania sphaerantha S.Hatt., J. Hattori Bot. Lab. 47: 99, 1980.

** Frullania sphaerolobulata S.H.Lin, Tunghai Journal 38: 104, 1997.

*** Frullania spinifera Taylor, London J. Bot. 5: 407, 1846.

** Frullania spinigastria S.Hatt., J. Hattori Bot. Lab. 45: 358, 1979.

** Frullania spiniplica S.Hatt., J. Hattori Bot. Lab. 36: 428, 1972 [1973].

*** Frullania spongiosa Steph., Hedwigia 33 (3): 147, 1894.

** Frullania squamuligera Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 33, 1884.

*** Frullania squarrosula (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 412, 1845. Bas.: Jungermannia squarrosula Hook.f. et Taylor, London J. Bot. 4: 88, 1845.

** Frullania subcaduca S.Hatt., J. Hattori Bot. Lab. 38: 267, 1974.

** Frullania subclavata Steph., Sp. Hepat. (Stephani) 4: 354, 1910.

** Frullania subnigricaulis S.Hatt., J. Hattori Bot. Lab. 37: 89, 1973.

** Frullania subnigricaulis var. subtruncata S.Hatt., J. Hattori Bot. Lab. 39: 308, 1975.

* Frullania subpedicellata S.Hatt., J. Hattori Bot. Lab. 47: 93, 1980. 203

** Frullania subsquarrosa S.Hatt., J. Hattori Bot. Lab. 36: 429, 1972 [1973].

** Frullania subvalida S.Hatt. et Thaithong, J. Jap. Bot. 53 (6): 173, 1978.

*** Frullania svihlana S.Hatt., J. Hattori Bot. Lab. 54: 180, 1983.

* Frullania taiheizana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 241, 1934.

** Frullania tamsuina Steph., Sp. Hepat. (Stephani) 4: 444, 1910.

*** Frullania taradakensis Steph., Sp. Hepat. (Stephani) 4: 352, 1910.

** Frullania tenuirostris Steph., Sp. Hepat. (Stephani) 4: 462, 1911.

** Frullania togashiana S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 118, 1975.

** Frullania tubercularis S.Hatt. et P.J.Lin, J. Jap. Bot. 60 (4): 107, 1985.

*** Frullania usamiensis Steph., Bull. Herb. Boissier 5 (2): 91, 1897.

** Frullania valdiviensis J.B.Jack et Steph., Hedwigia 33 (3): 149, 1894.

** Frullania valida Steph., Sp. Hepat. (Stephani) 4: 402, 1910.

** Frullania variegata Steph., Hedwigia 33 (3): 149, 1894.

* Frullania victoriensis Steph., Sp. Hepat. (Stephani) 4: 418, 1910.

*** Frullania vittiana S.Hatt., Bryologist 90 (4): 368, 1987 [1988].

** Frullania wangii S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 146, 1985.

*** Frullania yuennanensis Steph., Hedwigia 33 (3): 161, 1894.

** Frullania yuennanensis var. siamensis (N.Kitag., Thaithong et S.Hatt.) S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 148, 1985. Bas.: Frullania siamensis N.Kitag., Thaithong et S.Hatt., J. Hattori Bot. Lab. 43: 452, 1977 [1978].

** Frullania yuzawana S.Hatt., J. Hattori Bot. Lab. 49: 157, 1981.

** Frullania zangii S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 149, 1985.

** Frullania zennoskeana S.Hatt., J. Jap. Bot. 59 (10): 308, 1984.

** sect. Acutilobae Verd., Ann. Bryol., Suppl. 1: 44, 1930.

** Frullania allanii E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 371, 1949.

*** Frullania clavata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 428, 1845. Bas.: Jungermannia clavata Hook.f. et Taylor, London J. Bot. 4: 88, 1845.

** Frullania hamaticoma Steph., Hedwigia 28 (3): 158, 1889.

*** Frullania monocera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 418, 1845. Bas.: Jungermannia monocera Hook.f. et Taylor, London J. Bot. 4: 89, 1845.

** Frullania monocera var. acutiloba (Mitt.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015. Bas.: Frullania acutiloba Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 120, 1860 [1861].

** Frullania monocera var. depauperata S.Hatt., J. Hattori Bot. Lab. 57: 419, 1984.

** Frullania monocera var. schiffneri (Verd.) S.Hatt., J. Hattori Bot. Lab. 46: 120, 1979. Bas.: Frullania acutiloba var. schiffneri Verd., Ann. Bryol. 2: 123, 1929.

*** Frullania monocera var. subhampeana (E.A.Hodgs.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015. Bas.: Frullania subhampeana E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 370, 1949.

*** Frullania monocera var. undulata (Kamim.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015. Bas.: Frullania undulata Kamim., J. Hattori Bot. Lab. 24: 50, 1961.

** Frullania osumiensis (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 16: 87, 1956. Bas.: Frullania hampeana var. osumiensis S.Hatt., Bull. Tokyo Sci. Mus. 11: 144, 1944.

* Frullania pallidula S.Hatt., Beih. Nova Hedwigia 90: 152, 1988.

** Frullania pseudomonocera S.Hatt., J. Hattori Bot. Lab. 60: 216, 1986.

** Frullania seriata Gottsche, Hedwigia 28 (3): 160, 1889.

** Frullania spinistipula Steph., Sp. Hepat. (Stephani) 4: 463, 1911.

** Frullania streimannii S.Hatt., J. Hattori Bot. Lab. 54: 176, 1983.

*** sect. Australes Verd., Ann. Bryol., Suppl. 1: 58, 1930.

*** Frullania anomala E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 374, 1949.

* Frullania baileyana Steph., Sp. Hepat. (Stephani) 4: 417, 1910.

*** Frullania baladina Gottsche, Hedwigia 33 (3): 140, 1894. 204

* Frullania belmorensis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 106, 1914.

*** Frullania campanulata Sande Lac., Ned. Kruidk. Arch. 3: 422, 1854 [1855].

** Frullania campanulata var. caduca Verd., Ann. Bryol., Suppl. 1: 41, 1930.

** Frullania campanulata var. malesiaca (Verd.) S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 158, 1975. Bas.: Frullania malesiaca Verd., Ann. Bryol., Suppl. 1: 59, 1930.

* Frullania cataractarum Steph., Sp. Hepat. (Stephani) 4: 657, 1911.

* Frullania crawfordii Steph., Hedwigia 33 (3): 143, 1894.

*** Frullania dentata S.Hatt., J. Hattori Bot. Lab. 38: 231, 1974.

** Frullania dentata var. secernens S.Hatt., J. Hattori Bot. Lab. 65: 422, 1988.

*** Frullania errans Verd., Ann. Bryol., Suppl. 1: 59, 1930.

** Frullania errans var. angulistipula S.Hatt., J. Hattori Bot. Lab. 36: 431, 1972 [1973].

*** Frullania fugax (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 445, 1845. Bas.: Jungermannia fugax Hook.f. et Taylor, London J. Bot. 4: 87, 1845. 205

** Frullania fulfordiae S.Hatt., Bryologist 90 (4): 365, 1987 [1988].

*** Frullania glomerata (Lehm. et Lindenb.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 46, 1838. Bas.: Jungermannia glomerata Lehm. et Lindenb., Nov. Stirp. Pug. 5: 21, 1833.

*** Frullania incumbens Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 162, 1855.

*** Frullania inflexa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 120, 1860 [1861].

** Frullania media (E.A.Hodgs.) S.Hatt., J. Hattori Bot. Lab. 54: 153, 1983. Bas.: Frullania fugax var. media E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 375, 1949.

** Frullania mizutanii Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 524, 1973.

*** Frullania obscurifolia Mitt., Philos. Trans. 168: 400, 1879.

*** Frullania patagonica Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 88, 1911.

** Frullania pentapleura Taylor, London J. Bot. 5: 402, 1846.

** Frullania polyptera Taylor, London J. Bot. 5: 401, 1846.

** Frullania polyptera var. angustata (Mitt.) S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (2): 74, 1975. Bas.: Frullania angustata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 122, 1860 [1861].

** Frullania probosciphora Taylor, London J. Bot. 5: 402, 1846.

** Frullania pulchella Herzog, Rev. Bryol. Lichénol. 23 (1/2): 60, 1954.

*** Frullania sinskeana J.J.Engel et B.C.Tan, J. Hattori Bot. Lab. 60: 335, 1986. Nom. nov. pro Frullania spathulistipa Steph., Sp. Hepat. (Stephani) 4: 415, 1910, nom. illeg.

** Frullania socotrana Mitt., Trans. Roy Soc. Edinburgh 31: 335, 1888.

* Frullania solanderiana Colenso, Trans. & Proc. New Zealand Inst. 21: 75, 1889. 206

** Frullania subincumbens S.Hatt., Bryologist 90 (4): 367, 1987 [1988].

* Frullania subtropica Steph., Sp. Hepat. (Stephani) 4: 416, 1910.

* Frullania tjibodensis S.Hatt. et Thaithong, J. Jap. Bot. 52 (10): 289, 1977. 207

** Frullania tuyamae S.Hatt. et Thaithong, J. Jap. Bot. 53 (6): 175, 1978.

*** sect. Frullania

*** Frullania appalachiana R.M.Schust., Phytologia 53 (5): 366, 1983.

*** Frullania azorica Sim-Sim, Sérgio, Mues et Kraut, Cryptog. Bryol. Lichénol. 16 (2): 112, 1995.

*** Frullania catalinae A.Evans, Trans. Connecticut Acad. Arts 10 (1): 11, 1899.

*** Frullania dilatata (L.) Dumort., Recueil Observ. Jungerm.: 13, 1835. Bas.: Jungermannia dilatata L., Sp. Pl. 1: 1133, 1753.

** Frullania dilatata subsp. asiatica S.Hatt., J. Jap. Bot. 57 (9): 258, 1982.

*** Frullania eboracensis Lehm., Nov. Stirp. Pug. 8: 14, 1844. 208

*** Frullania ericoides (Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 12: 51, 1839. Bas.: Jungermannia ericoides Nees, Fl. Bras. (Martius) 1 (1): 346, 1833. 209

* Frullania ericoides var. laxa (Gottsche, Lindenb. et Nees) Schiffn., Consp. Hepat. Arch. Ind.: 324, 1898. Bas.: Frullania squarrosa γ laxa Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 772, 1847.

* Frullania ericoides var. minor Kamim., Bull. Kochi Gakuen Jun. Coll. 2: 22, 1971.

** Frullania ericoides var. verrucosa (Kamim.) Hentschel et von Konrat, Phytotaxa 220 (2): 134, 2015. Bas.: Frullania squarrosa var. verrucosa Kamim., J. Hattori Bot. Lab. 24: 19, 1961.

*** Frullania fragilifolia (Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 437, 1845. Bas.: Jungermannia fragilifolia Taylor, Ann. Mag. Nat. Hist. 12 (76): 172, 1843.

** Frullania fuegiana Steph., Sp. Hepat. (Stephani) 4: 428, 1910.

** Frullania hattoriana J.D.Godfrey et G.Godfrey, J. Hattori Bot. Lab. 48: 321, 1980.

** Frullania koponenii S.Hatt., Ann. Bot. Fenn. 15 (2): 111, 1978.

*** Frullania muscicola Steph., Hedwigia 33 (3): 146, 1894.

*** Frullania oakesiana Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 225, 1869.

** Frullania oakesiana subsp. takayuensis (Steph.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 195, 1992. Bas.: Frullania takayuensis Steph., Sp. Hepat. (Stephani) 4: 399, 1910.

*** Frullania parvistipula Steph., Sp. Hepat. (Stephani) 4: 397, 1910.

** Frullania sabaliana R.M.Schust., Phytologia 53 (5): 365, 1983.

** Frullania semivillosa Lindenb. et Gottsche, Syn. Hepat. 5: 774, 1847.

** Frullania stylifera (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 210, 1992. Bas.: Frullania inflata var. stylifera R.M.Schust., Phytologia 53 (5): 366, 1983.

* Frullania subdilatata C.Massal., Nuovo Giorn. Bot. Ital. (n.ser.) 13 (4): 349, 1906.

*** Frullania virginica Lehm., Nov. Stirp. Pug. 8: 19, 1844.

** sect. Irregulares E.A.Hodgs. ex S.Hatt., J. Hattori Bot. Lab. 54: 143, 1983.

* Frullania astrolabea Steph., Sp. Hepat. (Stephani) 4: 460, 1910. 210

*** Frullania deplanata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 161, 1855.

*** Frullania morobensis S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 109, 1985.

*** Frullania patula Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 159, 1854.

*** Frullania scandens Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 258, 1843.

** sect. Planae R.M.Schust., Phytologia 57 (5): 372, 1985.

*** Frullania plana Sull., Mem. Amer. Acad. Arts (n.ser.) 4: 175, 1849.

*** subg. Homotropantha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 35, 1884.

*** Frullania deflexa Mitt., Bonplandia 10 (2): 19, 1862.

*** Frullania integristipula (Nees) Nees, Syn. Hepat. 3: 431, 1845. Bas.: Jungermannia integristipula Nees, Enum. Pl. Crypt. Javae: 54, 1830.

** Frullania integristipula var. emarginata Verd., Ann. Bryol. 2: 153, 1929.

** Frullania macrophylla S.Hatt., J. Hattori Bot. Lab. 47: 220, 1980.

** Frullania sabahana S.Hatt., J. Hattori Bot. Lab. 40: 493, 1976.

** Frullania sackawana Steph., Bull. Herb. Boissier 5 (2): 91, 1897.

** Frullania sarawakensis S.Hatt., J. Hattori Bot. Lab. 40: 496, 1976.

** Frullania umbonata Mitt., Sp. Hepat. (Stephani) 4: 579, 1911.

*** Frullania utriculata Steph., Hedwigia 33 (3): 152, 1894.

*** sect. Fallaces Verd., Rev. Bryol. Lichénol. 1: 112, 1928.

* Frullania fallax Gottsche, Syn. Hepat. 3: 432, 1845.

*** Frullania intermedia (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 434, 1845. Bas.: Jungermannia intermedia Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 218, 1824 [1825]. 211

** Frullania intermedia subsp. morokensis (Steph.) S.Hatt., J. Hattori Bot. Lab. 47: 194, 1980. Bas.: Frullania morokensis Steph., Sp. Hepat. (Stephani) 4: 578, 1911.

* Frullania intermedia var. non-apiculata S.Hatt., J. Hattori Bot. Lab. 39: 291, 1975. 212

** Frullania novoguineensis Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 37, 1890.

** Frullania regularis Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 38, 1890.

*** sect. Nodulosae Verd., Rev. Bryol. Lichénol. 1: 116, 1928.

* Frullania brotheri Steph., Hedwigia 33 (3): 150, 1894. 213

** Frullania hamata Steph., Sp. Hepat. (Stephani) 4: 582, 1911.

** Frullania leeuwenii Verd., Nova Guinea 14: 545, 1930.

*** Frullania nodulosa (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 433, 1845. Bas.: Jungermannia nodulosa Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 217, 1824 [1825].

** sect. Remotilobae Verd., Rev. Bryol. Lichénol. 1: 119, 1928.

*** Frullania heteromorpha Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 38, 1890.

** Frullania remotiloba Steph., Hedwigia 33 (3): 152, 1894.

* subg. Mammillosae S.Hatt., J. Hattori Bot. Lab. 60: 226, 1986.

** Frullania huerlimannii S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 2 (3): 84, 1976.

* Frullania huerlimannii var. dioica S.Hatt., J. Hattori Bot. Lab. 57: 413, 1984.

** Frullania involvens S.Hatt. et Kamim., J. Hattori Bot. Lab. 37: 526, 1973.

** Frullania iriomotensis S.Hatt., J. Jap. Bot. 55 (5): 133, 1980.

** Frullania mammillosa S.Hatt., J. Hattori Bot. Lab. 43: 424, 1977 [1978].

*** Frullania meijeri S.Hatt., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 17 (4): 307, 1974.

** Frullania notarisii Steph., Sp. Hepat. (Stephani) 4: 651, 1911.

** Frullania papulosa Steph., Sp. Hepat. (Stephani) 4: 654, 1911.

** Frullania rudolfiana S.Hatt., J. Hattori Bot. Lab. 36: 437, 1972 [1973].

*** Frullania thiersiae S.Hatt., Beih. Nova Hedwigia 90: 156, 1988.

** Frullania tixieri S.Hatt., J. Jap. Bot. 51 (7): 193, 1976.

*** subg. Meteoriopsis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 37, 1884.

* Frullania caldensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 89, 1876 [1877].

** Frullania evoluta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 122, 1860 [1861].

*** Frullania tagawana (S.Hatt. et Thaithong) S.Hatt., J. Hattori Bot. Lab. 59: 160, 1985. Bas.: Frullania evoluta var. tagawana S.Hatt. et Thaithong, J. Hattori Bot. Lab. 43: 441, 1977 [1978].

*** sect. Intumescentes R.M.Schust., Phytologia 57 (5): 370, 1985.

*** Frullania aculeata Taylor, London J. Bot. 5: 407, 1846.

** Frullania ambronnii Steph., Biblioth. Bot. 87 (2): 242, 1916.

*** Frullania atrata (Sw.) Nees ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 12: 51, 1839. Bas.: Jungermannia atrata Sw., Prodr. (Swartz): 144, 1788.

*** Frullania beyrichiana (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 460, 1845. Bas.: Jungermannia beyrichiana Lehm. et Lindenb., Nov. Stirp. Pug. 5: 25, 1833.

*** Frullania bicornistipula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 46, 1884.

*** Frullania brasiliensis Raddi, Critt. Brasil.: 12, 1822. 214

** Frullania brasiliensis var. elegantula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 50, 1884.

* Frullania breuteliana Gottsche, Syn. Hepat. 3: 461, 1845. 215

* Frullania compacta Gottsche, Sp. Hepat. (Stephani) 4: 493, 1911. 216

* Frullania crenulifolia J.B.Jack et Steph., Hedwigia 31 (1): 14, 1892.

** Frullania crispiloba Steph., Hedwigia 33 (3): 156, 1894.

** Frullania curviramea Steph., Sp. Hepat. (Stephani) 4: 684, 1911.

*** Frullania ecuadorensis Steph., Sp. Hepat. (Stephani) 4: 526, 1911.

** Frullania formosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 46, 1884.

* Frullania granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 173 (79), 1864. 217

*** Frullania griffithsiana Gottsche, Syn. Hepat. 4: 466, 1846.

* Frullania guadalupensis Gottsche, Sp. Hepat. (Stephani) 4: 496, 1911.

* Frullania gualaquizana Steph., Sp. Hepat. (Stephani) 4: 531, 1911.

** Frullania hamiflora Herzog et L.Clark, Bryologist 56 (3): 180, 1953.

* Frullania humilis Spruce, Mem. Torrey Bot. Club 1 (3): 119, 1890.

*** Frullania intumescens (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 460, 1845. Bas.: Jungermannia intumescens Lehm. et Lindenb., Nov. Stirp. Pug. 6: 52, 1834.

* Frullania laticaulis Spruce, Mem. Torrey Bot. Club 1 (3): 120, 1890.

*** Frullania lobatohastata Steph., Sp. Hepat. (Stephani) 4: 499, 1911.

* Frullania longistipula var. apiculata Demaret et Vanden Berghen, Bull. Jard. Bot. État Bruxelles 20 (1): 4, 1950.

*** Frullania macrocephala (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 460, 1845. Bas.: Jungermannia macrocephala Lehm. et Lindenb., Nov. Stirp. Pug. 5: 20, 1833.

** Frullania madothecoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 47, 1884.

** Frullania meridana Steph., Sp. Hepat. (Stephani) 4: 500, 1911.

* Frullania microcephala Gottsche, Mexik. Leverm.: 251, 1863.

*** Frullania montagnei Gottsche, Syn. Hepat. 3: 456, 1845.

** Frullania moritziana Lindenb. et Gottsche, Syn. Hepat. 5: 782, 1847.

** Frullania osculatiana De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 236, 1857.

*** Frullania paradoxa Lehm. et Lindenb., Syn. Hepat. 3: 462, 1845.

** Frullania pearceana Steph., Sp. Hepat. (Stephani) 4: 515, 1911.

*** Frullania pittieri Steph., Primit. fl. costar.: 113, 1892 [1893].

** Frullania rigescens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 52, 1884.

*** Frullania setigera Steph., Hedwigia 33 (3): 159, 1894.

** Frullania speciosa Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 59, 1950.

** Frullania supradecomposita (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 431, 1845. Bas.: Jungermannia supradecomposita Lehm. et Lindenb., Nov. Stirp. Pug. 5: 23, 1833.

* Frullania trianae Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 173, 1864.

** Frullania triquetra Lindenb. et Gottsche, Syn. Hepat. 5: 780, 1847.

* Frullania trollii Herzog, Beih. Bot. Centralbl. 61B (3): 575, 1942.

*** Frullania uleana Steph., Hedwigia 33 (3): 155, 1894.

** sect. Meteoriopsis Uribe, von Konrat et Hentschel, Phytotaxa 220 (2): 132, 2015.

*** Frullania convoluta Lindenb. et Hampe, Linnaea 24 (3): 303, 1851 [1852].

* Frullania convoluta var. ampliata Herzog, Rev. Bryol. Lichénol. 20 (1/2): 129, 1951 [1952].

*** Frullania darwinii Gradst. et Uribe, Cryptog. Bryol. 25 (4): 296, 2004.

*** Frullania dulimensis Uribe, Cryptog. Bryol. 27 (3): 309, 2006.

*** Frullania grandifolia Steph., Sp. Hepat. (Stephani) 4: 684, 1911.

*** Frullania peruviana Gottsche, Syn. Hepat. 4: 465, 1846.

*** Frullania phalangiflora Steph., Biblioth. Bot. 87 (2): 247, 1916.

*** Frullania weberbaueri Steph., Sp. Hepat. (Stephani) 4: 510, 1911.

*** sect. Obtusilobae Verd., Ann. Bryol., Suppl. 1: 81, 1930.

** Frullania angulata Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 169, 1863.

** Frullania angulata var. laciniata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 72, 1976.

** Frullania apicalis Mitt., Philos. Trans. 168: 401, 1879.

** Frullania apicalis var. camerunensis Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 53, 1976.

* Frullania borbonica Lindenb., Syn. Hepat. 3: 455, 1845. 218

*** Frullania capensis Gottsche, Syn. Hepat. 3: 449, 1845.

** Frullania donnellii Austin, Bull. Torrey Bot. Club 6 (52): 301, 1879.

** Frullania eplicata Steph., Sp. Hepat. (Stephani) 4: 679, 1911.

** Frullania imerinensis Steph., Sp. Hepat. (Stephani) 4: 484, 1911.

*** Frullania kunzei (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 449, 1845. Bas.: Jungermannia kunzei Lehm. et Lindenb., Nov. Stirp. Pug. 6: 50, 1834.

** Frullania kunzei var. maritima R.M.Schust., Phytotaxa 220 (2): 135, 2015. Based on: Frullania kunzei var. maritima R.M.Schust., J. Hattori Bot. Lab. 70: 145, 1991, nom. inval.

** Frullania longistipula Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 199, 1891 [1892].

*** Frullania meyeniana Lindenb., Syn. Hepat. 3: 455, 1845.

** Frullania meyeniana var. dioica S.Hatt., J. Hattori Bot. Lab. 43: 426, 1977 [1978].

** Frullania onraedtii Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 60, 1976.

** Frullania papuana Verd., Ann. Bryol., Suppl. 1: 82, 1930.

*** Frullania schimperi Nees, Syn. Hepat. 3: 454, 1845. 219

** Frullania schimperi var. laciniata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 59, 1976.

*** subg. Microfrullania (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 34, 1992. Bas.: Neohattoria subg. Microfrullania R.M.Schust., J. Hattori Bot. Lab. 33: 280, 1970.

*** Frullania fertilis De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 235, 1857.

*** Frullania knightbridgei von Konrat et de Lange, PhytoKeys 8: 28, 2012.

*** Frullania magellanica F.Weber et Nees, Syn. Hepat. 3: 446, 1845. Nom. nov. pro Jungermannia magellanica Spreng. Ann. Wetterauischen Ges. Gesammte Naturk. 1: 25, 1809, nom. illeg.

** Frullania magellanica subsp. tristaniana (S.W.Arnell) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 59, 2013. Bas.: Frullania tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 9, 1958.

* Frullania matafaoica H.A.Mill., Phytologia 47 (4): 322, 1981. Nom. nov. pro Frullania minutissima Pearson, Amer. Samoa: 140, 1924, nom. illeg.

*** Frullania toropuku von Konrat, de Lange et Larraín, Polish Bot. J. 58 (2): 439, 2013.

*** sect. Amphijubula (R.M.Schust.) von Konrat, Hentschel, Heinrichs et Braggins, Bryologist 114 (1): 53, 2011. Bas.: Amphijubula R.M.Schust., J. Hattori Bot. Lab. 33: 298, 1970.

*** Frullania lobulata (Hook.) Hook. et Nees, Syn. Hepat. 3: 445, 1845. Bas.: Jungermannia lobulata Hook., Musci Exot. 2: tab. 119, 1820.

*** Frullania microcaulis Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 29 (1/4): 172, 1922 [1923].

*** Frullania truncatistyla von Konrat, Hentschel, Heinrichs et Braggins, Bryologist 114 (1): 63, 2011.

*** sect. Microfrullania (R.M.Schust.) von Konrat et Hentschel, Phytotaxa 220 (2): 133, 2015. Bas.: Neohattoria sect. Microfrullania R.M.Schust., J. Hattori Bot. Lab. 33: 288, 1970.

*** Frullania chevalieri (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 34, 1992. Bas.: Neohattoria chevalieri R.M.Schust., J. Hattori Bot. Lab. 33: 289, 1970. 220

*** Frullania microscopica Pearson, J. Linn. Soc., Bot. 46 (305): 33, 1922.

** Frullania neocaledonica J.J.Engel, Novon 9 (3): 344, 1999. Nom. nov. pro Neohattoria caledonica R.M.Schust., J. Hattori Bot. Lab. 33: 291, 1970.

*** Frullania parhamii (R.M.Schust.) R.M.Schust. ex von Konrat, L.Söderstr. et A.Hagborg, Telopea 13 (3): 407, 2011. Bas.: Neohattoria parhamii R.M.Schust., J. Hattori Bot. Lab. 26: 243, 1963.

*** sect. Regulares Verd., Ann. Bryol., Suppl. 1: 133, 1930.

*** Frullania junghuhniana Gottsche, Syn. Hepat. 3: 444, 1845. 221

** Frullania junghuhniana var. bisexualis S.Hatt., J. Hattori Bot. Lab. 40: 485, 1976.

** Frullania junghuhniana var. tenella (Sande Lac.) Grolle et S.Hatt., Misc. Bryol. Lichenol. 9 (6): 123, 1982. Bas.: Frullania tenella Sande Lac., Ned. Kruidk. Arch. 3: 423, 1854 [1855].

** Frullania mcveanii S.Hatt., J. Hattori Bot. Lab. 37: 55, 1973.

** Frullania pseudomeyeniana S.Hatt., J. Hattori Bot. Lab. 60: 231, 1986.

*** Frullania rostrata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 445, 1845. Bas.: Jungermannia rostrata Hook.f. et Taylor, London J. Bot. 4: 87, 1845. 222

** Frullania scalaris S.Hatt., J. Hattori Bot. Lab. 43: 432, 1977 [1978].

** subg. Saccophora Verd., Ann. Bryol. 2: 121, 1929.

*** Frullania gaudichaudii (Nees et Mont.) Nees et Mont., Syn. Hepat. 3: 435, 1845. Bas.: Jubula gaudichaudii Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 64, 1836.

** Frullania gaudichaudii var. ceylanica (Nees) S.Hatt., J. Hattori Bot. Lab. 47: 104, 1980. Bas.: Frullania ceylanica Nees, Syn. Hepat. 3: 436, 1845.

*** Frullania hedrantha S.Hatt. et Kamim., J. Hattori Bot. Lab. 37: 519, 1973.

** Frullania immersa Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 315, 1896.

** Frullania pancheri Gottsche, Hedwigia 33 (3): 159, 1894.

** Frullania papillilobula S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 141, 1975.

** Frullania sublignosa Steph., Hedwigia 33 (3): 148, 1894.

* subg. Steerea (S.Hatt. et Kamim.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 32, 1992. Bas.: Steerea S.Hatt. et Kamim., J. Hattori Bot. Lab. 34: 429, 1971.

*** Frullania clemensiana Verd., Ned. Kruidk. Arch. (ser. 3) 42 (2): 493, 1932.

*** subg. Thyopsiella Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 41, 1884.

*** Frullania acicularis Hentschel et von Konrat, Phytotaxa 220 (2): 134, 2015. Nom. nov. pro Frullania tamarisci var. azorica J.-P.Frahm, Trop. Bryol. 27: 102, 2006. 223

** Frullania alstonii Verd., Ann. Bryol., Suppl. 1: 76, 1930.

** Frullania aoshimensis Horik., Sci. Rep. Tôhoku Imp. Univ., Ser. 4, Biol. 4 (1): 64, 1929.

*** Frullania appendiculata Steph., Bull. Herb. Boissier 5 (2): 88, 1897. 224

*** Frullania asagrayana Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 14, 1842. 225

*** Frullania calcarifera Steph., Bol. Soc. Brot. 4: 241, 1886 [1887]. 226

*** Frullania californica (M.Howe) A.Evans, Trans. Connecticut Acad. Arts 10 (1): 25, 1899. Bas.: Frullania asagrayana var. californica M.Howe, Erythea 2 (6): 98, 1894.

*** Frullania densiloba Steph. ex A.Evans, Proc. Wash. Acad. Sci. 8: 157, 1906.

*** Frullania franciscana M.Howe, Erythea 2 (6): 99, 1894.

** Frullania iwatsukii S.Hatt., J. Hattori Bot. Lab. 35: 240, 1972.

*** Frullania microphylla (Gottsche) Pearson, J. Bot. 32: 328, 1894. Bas.: Frullania tamarisci var. microphylla Gottsche, Hepat. Eur., Leberm. 21-22: no. 109 [209], 1862.

*** Frullania moniliata (Reinw., Blume et Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 13, 1842. Bas.: Jungermannia moniliata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 224, 1824 [1825]. 227

*** Frullania nisquallensis Sull., Mem. Amer. Acad. Arts (n.ser.) 4: 175, 1849.

*** Frullania polysticta Lindenb., Syn. Hepat. 3: 440, 1845.

** Frullania pseudoalstonii Tsudo et J.Haseg., Bryol. Res. 9 (3): 44, 2006.

** Frullania punctata Reimers, Hedwigia 71 (1/2): 36, 1931.

** Frullania schaefer-verwimpii Yuzawa et S.Hatt., J. Jap. Bot. 64 (2): 37, 1989.

** Frullania selwyniana Pearson, List. Canad. Hepat.: 1, 1890.

*** Frullania sergiae Sim-Sim, Fontinha, Mues et Lion, Nova Hedwigia 71 (1/2): 186, 2000.

*** Frullania subarctica Vilnet, Borovich. et Bakalin, Phytotaxa 173 (1): 67, 2014. 228

*** Frullania tamarisci (L.) Dumort., Recueil Observ. Jungerm.: 13, 1835. Bas.: Jungermannia tamarisci L., Sp. Pl. 1: 1134, 1753.

*** Frullania teneriffae (F.Weber) Nees, Naturgesch. Eur. Leberm. 3: 239, 1838. Bas.: Jungermannia teneriffae F.Weber, Hist. Musc. Hepat. Prodr.: 23, 1815.

** Frullania trigona L.Clark, Jovet-Ast et Frye, Bryologist 50 (1): 52, 1947.

Incertae sedis

* Frullania affinis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 257, 1843. 229

* Frullania allionii Steph., Sp. Hepat. (Stephani) 4: 394, 1910.

* Frullania alpina Steph., Sp. Hepat. (Stephani) 4: 533, 1911.

** Frullania alternans Nees, Syn. Hepat. 3: 430, 1845.

* Frullania apertilobula Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 478, 1947.

** Frullania armata Herzog et L.Clark, Bryologist 57 (1): 36, 1954.

*** Frullania bolanderi Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 226, 1869.

*** Frullania boveana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 244, 1885.

* Frullania caespitans Beauverd, Sp. Hepat. (Stephani) 6: 537, 1924. Nom. nov. pro Frullania campanulata Steph., Biblioth. Bot. 87 (2): 242, 1916, nom. illeg.

* Frullania canaliculata Gottsche, Sp. Hepat. (Stephani) 4: 391, 1910.

** Frullania capillaris Steph., Sp. Hepat. (Stephani) 4: 616, 1911.

* Frullania cavallii Gola, Ann. Bot. (Rome) 6 (2): 275, 1907.

* Frullania chiapasana Steph., Sp. Hepat. (Stephani) 4: 392, 1910.

* Frullania chilcootiensis Steph., Bot. Jahrb. Syst. 8 (2): 98, 1886.

* Frullania chiovendae Gola, Ann. Bot. (Rome) 13 (1): 71, 1914.

** Frullania ciliata Lindenb. et Gottsche, Syn. Hepat. 5: 775, 1847.

** Frullania cinchonae Gottsche, Syn. Hepat. 3: 455, 1845.

* Frullania complicata Steph., Sp. Hepat. (Stephani) 4: 671, 1911.

* Frullania cordaeana Lindenb., Syn. Hepat. 3: 463, 1845.

** Frullania cuneatistipula Steph., Sp. Hepat. (Stephani) 6: 538, 1924.

* Frullania cuspiloba Steph., Sp. Hepat. (Stephani) 4: 394, 1910.

* Frullania dispar Nees, Syn. Hepat. 3: 429, 1845.

* Frullania guatemalensis Steph., Sp. Hepat. (Stephani) 4: 497, 1911.

*** Frullania inflata Gottsche, Syn. Hepat. 3: 424, 1845.

** Frullania inflata var. communis R.M.Schust., Phytologia 57 (5): 372, 1985.

** Frullania inflata var. dioica S.Hatt. et Thaithong, J. Hattori Bot. Lab. 44: 185, 1978.

** Frullania inflata var. mayebarae (S.Hatt.) K.Yamada, Misc. Bryol. Lichenol. 6 (9): 163, 1974. Bas.: Frullania mayebarae S.Hatt., Bot. Mag. (Tokyo) 65 (763/764): 13, 1952.

* Frullania laetevirens Hampe ex Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 420, 1845.

* Frullania larjiana Sushil K.Singh et D.K.Singh, J. Bryol. 27 (2): 105, 2005.

* Frullania larjiana var. didyhatii S.N.Srivast. et M.Rai, Geophytology 41 (1/2): 109, 2011.

* Frullania leana Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 227, 1869.

* Frullania madagascariensis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 364, 1882.

* Frullania mauritiana Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 227, 1869.

*** Frullania mirabilis J.B.Jack et Steph., Hedwigia 31 (1): 15, 1892.

** Frullania monoica Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 67, 1900.

* Frullania ocanniensis Steph., Sp. Hepat. (Stephani) 6: 542, 1924.

*** Frullania platycalyx Herzog, Feddes Repert. Spec. Nov. Regni Veg. 55 (1): 10, 1952.

* Frullania pyricalycina Steph., Sp. Hepat. (Stephani) 4: 394, 1910.

** Frullania quillotensis (Nees et Mont.) Nees et Mont., Syn. Hepat. 3: 427, 1845. Bas.: Jubula quillotensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 64, 1836.

** Frullania reicheana Steph., Sp. Hepat. (Stephani) 4: 427, 1910.

* Frullania sabanetica Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 170, 1864.

* Frullania semienana Gola, Ann. Bot. (Rome) 13 (1): 71, 1914.

** Frullania serrifolia Steph., Sp. Hepat. (Stephani) 4: 526, 1911.

** Frullania subpyricalycina Herzog, Rev. Bryol. Lichénol. 23 (1/2): 57, 1954.

** Frullania subtruncata Steph., Sp. Hepat. (Stephani) 4: 389, 1910.

** Frullania turfosa Lindenb. et Gottsche, Syn. Hepat. 5: 779, 1847.

** Frullania udarii V.Nath et Ajit P.Singh, Curr. Sci. 91 (6): 744, 2006.

** Frullania valparaisiana Lehm., Nov. Stirp. Pug. 10: 17, 1857.

** Frullania wagneri Steph., Sp. Hepat. (Stephani) 4: 392, 1910.

*** Jubulaceae H.Klinggr.

by J. Hentschel and M. von Konrat

The systematic placement of Neohattoria herzogii has been contentious since its description six decades ago. It has been interpreted as either a member of the genus Frullania or segregated into its own genus, Neohattoria, due to morphological similarities with both Frullania and Jubula. We follow Larrain et al. (2015) that provided molecular evidence supporting the recognition of the genus Neohattoria and its inclusion within the Jubulaceae, together with Jubula and Nipponolejeunea.

*** Jubula Dumort., Commentat. Bot. (Dumortier): 112, 1822 nom. conserv.

*** Jubula blepharophylla Grolle, J. Hattori Bot. Lab. 28: 47, 1965.

** Jubula hattorii Udar et V.Nath, Misc. Bryol. Lichenol. 8 (3): 49, 1978.

** Jubula hattorii var. muthukuzhiana A.E.D.Daniels et P.Daniel, Bryofl. Southernm. W Ghats: 181, 2013.

** Jubula himalayensis S.C.Srivast. et D.Sharma, Proc. Indian Acad. Sci. Pl. Sci. 100 (2): 85, 1990.

*** Jubula hutchinsiae (Hook.) Dumort., Syll. Jungerm. Europ.: 36, 1831. Bas.: Jungermannia hutchinsiae Hook., Brit. Jungermann.: tab. 1, 1812. 230

** Jubula hutchinsiae subsp. australiae Pócs et A.Cairns, Nova Hedwigia 86 (1/2): 232, 2008.

** Jubula hutchinsiae subsp. bogotensis (Steph.) Verd., Ann. Cryptog. Exot. 1 (2): 215, 1928. Bas.: Jubula bogotensis Steph., Sp. Hepat. (Stephani) 4: 687, 1911. 231

*** Jubula hutchinsiae subsp. caucasica Konstant. et Vilnet, Arctoa 20: 234, 2011.

*** Jubula hutchinsiae subsp. japonica (Steph.) Horik. et Ando, J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 6: 300, 1954. Bas.: Jubula japonica Steph., Bull. Herb. Boissier 5 (2): 92, 1897.

*** Jubula hutchinsiae subsp. javanica (Steph.) Verd., Ann. Cryptog. Exot. 1 (2): 216, 1928. Bas.: Jubula javanica Steph., Sp. Hepat. (Stephani) 4: 688, 1911. 232

*** Jubula hutchinsiae subsp. pennsylvanica (Steph.) Verd., Ann. Cryptog. Exot. 1 (2): 215, 1928. Bas.: Frullania pennsylvanica Steph., Hedwigia 22 (10): 147, 1883.

** Jubula kwangsiensis C.Gao et K.C.Chang, Bull. Bot. Res., Harbin 4 (3): 89, 1984.

*** Neohattoria Kamim., J. Jap. Bot. 37 (7): 218, 1962. Nom. nov. pro Hattoria Kamim., J. Hattori Bot. Lab. 24: 93, 1961.

*** Neohattoria herzogii (S.Hatt.) Kamim., J. Jap. Bot. 37 (7): 218, 1962. Bas.: Frullania herzogii S.Hatt., Feddes Repert. Spec. Nov. Regni Veg. 58: 53, 1955.

*** Nipponolejeunea S.Hatt., Bull. Tokyo Sci. Mus. 11: 124, 1944.

*** Nipponolejeunea pilifera (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 125, 1944. Bas.: Pycnolejeunea pilifera Steph., Sp. Hepat. (Stephani) 5: 624, 1914.

*** Nipponolejeunea subalpina (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 125, 1944. Bas.: Pycnolejeunea subalpina Horik., J. Jap. Bot. 15: 360, 1939.

*** Lejeuneaceae Cavers

by S.R. Gradstein, T. Pócs and R.-L. Zhu with contributions by G. Dauphin (Ceratolejeunea), X. He (Pycnolejeunea), A.-L. Ilkiu-Borges (Prionolejeunea), E. Reiner-Drehwald (Rectolejeunea, Lejeunea), A. Sass-Gyarmati (Lopholejeunea), A. Schäfer-Verwimp (Diplasiolejeunea) and P. Sukkharak (Thysananthus)

The subdivision of Lejeuneaceae follows with updates from, ) and. Some nomenclatural and taxonomic notes can also be found in,,, ), and.

*** subtrib. Brachiolejeuneinae Gradst.

** Acanthocoleus R.M.Schust., Bull. Torrey Bot. Club 97 (6): 339, 1970 [1971].

*** Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt, Bryophyt. Biblioth. 36: 62, 1988. Bas.: Lejeunea aberrans Lindenb. et Gottsche, Syn. Hepat. 5: 751, 1847.

*** Acanthocoleus aberrans var. laevis Gradst., Fl. Neotrop. Monogr. 62: 193, 1994.

*** Acanthocoleus chrysophyllus (Lehm.) Kruijt, Bryophyt. Biblioth. 36: 72, 1988. Bas.: Jungermannia chrysophylla Lehm., Linnaea 9 (4): 423, 1835.

** Acanthocoleus elgonensis Gyarmati et Pócs, Cryptog. Bryol. 35 (2): 120, 2014.

*** Acanthocoleus gilvus (Gottsche) Kruijt, Bryophyt. Biblioth. 36: 79, 1988. Bas.: Lejeunea gilva Gottsche, Abh. Naturwiss. Vereins Bremen 7: 353, 1882.

*** Acanthocoleus javanicus (Steph.) Kruijt, Bryophyt. Biblioth. 36: 85, 1988. Bas.: Dicranolejeunea javanica Steph., Sp. Hepat. (Stephani) 5: 169, 1912.

*** Acanthocoleus juddii Kruijt, Bryophyt. Biblioth. 36: 93, 1988.

** Acanthocoleus madagascariensis (Steph.) Kruijt, Bryophyt. Biblioth. 36: 98, 1988. Bas.: Dicranolejeunea madagascariensis Steph., Sp. Hepat. (Stephani) 5: 158, 1912.

*** Acanthocoleus trigonus (Nees et Mont.) Gradst., Contr. Univ. Michigan Herb. 18: 101, 1992. Bas.: Lejeunea trigona Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 61, 1836.

** Acanthocoleus yoshinaganus (S.Hatt.) Kruijt, Bryophyt. Biblioth. 36: 105, 1988. Bas.: Lopholejeunea subfusca var. yoshinagana S.Hatt., Bot. Mag. (Tokyo) 58 (686): 38, 1944.

*** Blepharolejeunea S.W.Arnell, Svensk Bot. Tidskr. 56 (2): 335, 1962.

*** Blepharolejeunea chimantaensis van Slageren et Kruijt, Beih. Nova Hedwigia 80: 126, 1985.

*** Blepharolejeunea fuegiana (Besch. et C.Massal.) Gradst., Beih. Nova Hedwigia 80: 130, 1985. Bas.: Lejeunea fuegiana Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 638, 1886.

*** Blepharolejeunea incongrua (Lindenb. et Gottsche) van Slageren et Kruijt, Beih. Nova Hedwigia 80: 133, 1985. Bas.: Lejeunea incongrua Lindenb. et Gottsche, Syn. Hepat. 5: 750, 1847.

*** Blepharolejeunea saccata (Steph.) van Slageren et Kruijt, Beih. Nova Hedwigia 80: 138, 1985. Bas.: Dicranolejeunea saccata Steph., Hedwigia 35 (3): 78, 1896.

*** Blepharolejeunea securifolia (Steph.) R.M.Schust., Phytologia 45 (5): 424, 1980. Bas.: Brachiolejeunea securifolia Steph., Sp. Hepat. (Stephani) 5: 128, 1912.

*** Brachiolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Lejeunea subg. Brachiolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 129, 1884.

*** Brachiolejeunea conduplicata (Steph.) Gradst., Fl. Neotrop. Monogr. 62: 175, 1994. Bas.: Archilejeunea conduplicata Steph., Sp. Hepat. (Stephani) 4: 712, 1911.

*** Brachiolejeunea fernandeziana S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 18, 1957.

*** Brachiolejeunea laxifolia (Taylor) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Phragmicoma laxifolia Taylor, London J. Bot. 6: 341, 1847.

*** Brachiolejeunea leiboldiana (Gottsche et Lindenb.) Schiffn., Hedwigia 33 (4): 182, 1894. Bas.: Phragmicoma leiboldiana Gottsche et Lindenb., Syn. Hepat. 2: 296, 1845.

*** Brachiolejeunea phyllorhiza (Nees) Kruijt et Gradst., Nova Hedwigia 43 (3/4): 299, 1986. Bas.: Jungermannia phyllorhiza Nees, Fl. Bras. (Martius) 1 (1): 348, 1833.

*** Brachiolejeunea spruceana (C.Massal.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Lejeunea spruceana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 246, 1885.

*** Dicranolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Lejeunea subg. Dicranolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 138, 1884.

*** Dicranolejeunea axillaris (Nees et Mont.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Lejeunea axillaris Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 59, 1836.

* Dicranolejeunea bovonei Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 27 (2/4): 248, 1920.

*** Lindigianthus Kruijt et Gradst., Beih. Nova Hedwigia 80: 165, 1985.

*** Lindigianthus cipaconeus (Gottsche) Kruijt et Gradst., Beih. Nova Hedwigia 80: 166, 1985. Bas.: Lejeunea cipaconea Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 150, 1864.

*** Odontolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893. Bas.: Lejeunea subg. Odontolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 142, 1884.

*** Odontolejeunea decemdentata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 175, 1912. Bas.: Lejeunea decemdentata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 148, 1884.

*** Odontolejeunea lunulata (F.Weber) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Jungermannia lunulata F.Weber, Hist. Musc. Hepat. Prodr.: 33, 1815.

*** Odontolejeunea rhomalea (Spruce) Steph., Sp. Hepat. (Stephani) 5: 176, 1912. Bas.: Lejeunea rhomalea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 147, 1884.

*** subtrib. Stictolejeuneinae Gradst.

*** Neurolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893. Bas.: Lejeunea subg. Neurolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 84, 1884.

*** Neurolejeunea breutelii (Gottsche) A.Evans, Bull. Torrey Bot. Club 34 (1): 13, 1907. Bas.: Lejeunea breutelii Gottsche, Syn. Hepat. 3: 324, 1845.

** Neurolejeunea breutelii var. africana Pócs, Herzogia 28 (1): 63, 2015.

*** Neurolejeunea catenulata (Nees) Schiffn., Hepat. (Engl.-Prantl): 131, 1893. Bas.: Lejeunea catenulata Nees, Syn. Hepat. 3: 323, 1845.

*** Neurolejeunea sastreana Gradst., Bryologist 92 (3): 345, 1989.

*** Neurolejeunea seminervis (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893. Bas.: Lejeunea seminervis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 84, 1884.

*** Stictolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893. Bas.: Lejeunea subg. Stictolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 81, 1884.

*** subg. Leptostictolejeunea R.M.Schust., Phytologia 56 (2): 70, 1984.

*** Stictolejeunea balfourii (Mitt.) E.W.Jones, J. Bryol. 9 (1): 50, 1976. Bas.: Lejeunea balfourii Mitt., Philos. Trans. 168: 398, 1879.

*** Stictolejeunea balfourii var. bekkeri Gradst., Beih. Nova Hedwigia 80: 214, 1985.

*** Stictolejeunea iwatsukii Mizut., J. Hattori Bot. Lab. 44: 134, 1978.

*** subg. Stictolejeunea

*** Stictolejeunea squamata (Willd.) Schiffn., Hepat. (Engl.-Prantl): 131, 1893. Bas.: Jungermannia squamata Willd. ex F.Weber, Hist. Musc. Hepat. Prodr.: 33, 1815.

*** trib. Lejeuneeae Dumort.

** Dactylophorella R.M.Schust., Phytologia 45 (5): 427, 1980.

*** Dactylophorella muricata (Gottsche) R.M.Schust., Phytologia 45 (5): 427, 1980. Bas.: Lejeunea muricata Gottsche, Syn. Hepat. 3: 348, 1845.

*** Metalejeunea Grolle, Bryophyt. Biblioth. 48: 17, 1995.

** Metalejeunea crassitexta (J.B.Jack et Steph.) Pócs, Telopea 13 (3): 456, 2011. Bas.: Microlejeunea crassitexta J.B.Jack et Steph., Bot. Centralbl. 60 (4): 106, 1894.

*** Metalejeunea cucullata (Reinw., Blume et Nees) Grolle, Bryophyt. Biblioth. 48: 100, 1995. Bas.: Jungermannia cucullata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 227, 1824 [1825].

*** Metalejeunea winkleri R.L.Zhu et Grolle, Nova Hedwigia 74 (3/4): 498, 2002.

*** Pictolejeunea Grolle, Feddes Repert. 88 (4): 248, 1977.

*** Pictolejeunea levis Grolle et M.E.Reiner, J. Bryol. 27 (3): 281, 2005.

*** Pictolejeunea mizutanii Grolle, Feddes Repert. 88 (4): 255, 1977. Nom. nov. pro Cheilolejeunea picta Mizut., J. Hattori Bot. Lab. 33: 226, 1970.

** Pictolejeunea piconii Pócs, Acta Bot. Hung. 49 (1/2): 110, 2007.

*** Pictolejeunea picta (Steph.) Grolle, Feddes Repert. 88 (4): 252, 1977. Bas.: Prionolejeunea picta Steph., Sp. Hepat. (Stephani) 5: 223, 1913.

*** Pictolejeunea reginae Ilk.-Borg., Brittonia 54 (4): 318, 2002 [2003].

*** Pictolejeunea sprucei Grolle, Feddes Repert. 88 (4): 249, 1977.

** subtrib. Ceratolejeuneinae Gradst.

*** Ceratolejeunea (Spruce) J.B.Jack et Steph., Hedwigia 31 (1): 16, 1892. Bas.: Lejeunea subg. Ceratolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 198, 1884. 233

*** subg. Ceratolejeunea

*** Ceratolejeunea andringitrae Pócs, Polish Bot. J. 56 (2): 144, 2011.

** Ceratolejeunea atlantica Alvarenga et Ilk.-Borg., Nova Hedwigia 86 (1/2): 238, 2008 (Ilkiu-Borges and Alvarenga 2008).

** Ceratolejeunea belangeriana (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 396, 1913. Bas.: Lejeunea belangeriana Gottsche, Syn. Hepat. 3: 398, 1845.

** Ceratolejeunea beninensis E.W.Jones et Vanden Berghen, Bull. Jard. Bot. État Bruxelles 21 (1/2): 63, 1951.

*** Ceratolejeunea brevinervis (Spruce) A.Evans, Bull. Torrey Bot. Club 32 (6): 282, 1905. Bas.: Lejeunea brevinervis Spruce, J. Linn. Soc., Bot. 30 (210): 342, 1895.

*** Ceratolejeunea ceratantha (Nees et Mont.) Schiffn., Bot. Jahrb. Syst. 23 (5): 582, 1897. Bas.: Lejeunea ceratantha Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 335, 1840.

** Ceratolejeunea coalita (Ångstr.) Steph., Sp. Hepat. (Stephani) 5: 402, 1913. Bas.: Lejeunea coalita Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 135, 1873.

*** Ceratolejeunea coarina (Gottsche) Schiffn., Hepat. (Engl.-Prantl): 125, 1893. Bas.: Lejeunea coarina Gottsche, Syn. Hepat. 3: 395, 1845.

*** Ceratolejeunea confusa R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (2): 313, 1956.

*** Ceratolejeunea cornuta (Lindenb.) Steph., Pflanzenw. Ost-Afrikas C: 65, 1895. Bas.: Jungermannia cornuta Lindenb., Syn. hepat. eur: 23, 1829.

*** Ceratolejeunea cubensis (Mont.) Schiffn., Hepat. (Engl.-Prantl): 125, 1893. Bas.: Lejeunea cubensis Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 481, 1842.

*** Ceratolejeunea dentistipula Steph., Sp. Hepat. (Stephani) 5: 407, 1913.

*** Ceratolejeunea fallax (Lehm. et Lindenb.) Bonner, Candollea 14: 189, 1953. Bas.: Jungermannia fallax Lehm. et Lindenb., Nov. Stirp. Pug. 5: 17, 1833.

*** Ceratolejeunea filaria (Taylor) Steph., Sp. Hepat. (Stephani) 5: 412, 1913. Bas.: Lejeunea filaria Taylor, Nov. Stirp. Pug. 8: 28, 1844.

** Ceratolejeunea floribunda Steph., Sp. Hepat. (Stephani) 5: 412, 1913.

*** Ceratolejeunea guianensis (Nees et Mont.) Steph., Sp. Hepat. (Stephani) 5: 416, 1913. Bas.: Lejeunea guianensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 335, 1840.

* Ceratolejeunea karstenii Steph., Sp. Hepat. (Stephani) 5: 420, 1913.

* Ceratolejeunea kuerschneri Eb.Fisch. et Vanderp., Beih. Nova Hedwigia 138: 87, 2010. 234

*** Ceratolejeunea laetefusca (Austin) R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (2): 306, 1956. Bas.: Lejeunea laetefusca Austin, Bot. Bull. (Hanover) 1 (8): 36, 1876.

* Ceratolejeunea ledermannii Steph., Sp. Hepat. (Stephani) 6: 399, 1923.

*** Ceratolejeunea malleigera (Spruce) Steph., Sp. Hepat. (Stephani) 5: 422, 1913. Bas.: Lejeunea malleigera Spruce, Mem. Torrey Bot. Club 1 (3): 123, 1890.

** Ceratolejeunea maranhensis Silva Brito et Ilk.-Borg., Nova Hedwigia 95 (3/4): 424, 2012.

** Ceratolejeunea minor Mizut., J. Hattori Bot. Lab. 49: 311, 1981.

*** Ceratolejeunea minuta G.Dauphin, Fl. Neotrop. Monogr. 90: 66, 2003.

** Ceratolejeunea moniliata Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 205, 1931.

** Ceratolejeunea oculata (Gottsche) Steph., Bull. Herb. Boissier 5 (10): 842, 1897. Bas.: Lejeunea oculata Gottsche, Syn. Hepat. 3: 357, 1845.

*** Ceratolejeunea oxygonia Steph., Sp. Hepat. (Stephani) 5: 429, 1913.

*** Ceratolejeunea papuliflora Steph., Sp. Hepat. (Stephani) 5: 430, 1913.

*** Ceratolejeunea patentissima (Hampe et Gottsche) A.Evans, Bull. Torrey Bot. Club 32 (6): 286, 1905. Bas.: Lejeunea patentissima Hampe et Gottsche, Linnaea 25 (3): 355, 1852 [1853].

*** Ceratolejeunea pungens Steph., Sp. Hepat. (Stephani) 5: 434, 1913.

*** Ceratolejeunea rubiginosa Steph., Hedwigia 34 (5): 237, 1895.

*** Ceratolejeunea saroltae Pócs, Polish Bot. J. 56 (2): 150, 2011.

* Ceratolejeunea sinensis P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 232, 1964. 235

** Ceratolejeunea singapurensis (Lindenb.) Schiffn., Consp. Hepat. Arch. Ind.: 273, 1898. Bas.: Lejeunea singapurensis Lindenb., Syn. Hepat. 3: 397, 1845.

*** Ceratolejeunea spinosa (Gottsche) Steph., Hedwigia 34 (5): 238, 1895. Bas.: Lejeunea spinosa Gottsche, Syn. Hepat. 3: 402, 1845.

*** Ceratolejeunea temnantha (Spruce) M.E.Reiner, Cryptog. Bryol. 32 (2): 95, 2011. Bas.: Lejeunea temnantha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 250, 1884.

*** Ceratolejeunea umbonata Steph., Sp. Hepat. (Stephani) 5: 446, 1913.

** Ceratolejeunea vitiensis Steph., Sp. Hepat. (Stephani) 5: 448, 1913.

** Ceratolejeunea zenkeri Steph., Sp. Hepat. (Stephani) 5: 449, 1914.

*** subg. Ceratophora R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (2): 294, 1956.

*** Ceratolejeunea desciscens (Sande Lac.) Schiffn., Hepat. (Engl.-Prantl): 126, 1893. Bas.: Lejeunea desciscens Sande Lac., Syn. hepat. jav.: 107, 1856 [1857].

*** Ceratolejeunea globulifera Herzog, Rev. Bryol. Lichénol. 13: 23, 1942.

*** Ceratolejeunea grandiloba J.B.Jack et Steph., Hedwigia 31 (1): 16, 1892.

** Ceratolejeunea grandiloba subsp. inflata (Mizut.) Gradst., Phytotaxa 81 (1): 5, 2013. Bas.: Ceratolejeunea inflata Mizut., J. Hattori Bot. Lab. 49: 313, 1981.

*** Ceratolejeunea szyszylowiczii (Loitl.) Steph., Sp. Hepat. (Stephani) 5: 443, 1913. Bas.: Lejeunea szyszylowiczii Loitl., Diagn. pl. nov.: 19, 1894.

Incertae sedis

** Ceratolejeunea aliena Herzog, Trans. Brit. Bryol. Soc. 2 (1): 71, 1952.

*** Luteolejeunea Piippo, Acta Bot. Fenn. 132: 56, 1986.

*** Luteolejeunea herzogii (Buchloh) Piippo, Acta Bot. Fenn. 132: 57, 1986. Bas.: Stictolejeunea herzogii Buchloh, Nova Hedwigia 3 (4): 515, 1961.

*** Otigoniolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 125, 1893. Bas.: Lejeunea subg. Otigoniolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 226, 1884.

* Otigoniolejeunea crenulata Steph., Sp. Hepat. (Stephani) 6: 408, 1923. 236

*** Otigoniolejeunea huctumalcensis (Lindenb. et Gottsche) Y.M.Wei, R.L.Zhu et Gradst., Phytotaxa 162 (4): 237, 2014. Bas.: Lejeunea huctumalcensis Lindenb. et Gottsche, Syn. Hepat. 5: 762, 1847.

* Otigoniolejeunea ledermannii Steph., Sp. Hepat. (Stephani) 6: 409, 1923. 237

*** Otigoniolejeunea portoricensis (Hampe et Gottsche) Y.M.Wei, R.L.Zhu et Gradst., Phytotaxa 162 (4): 237, 2014. Bas.: Lejeunea portoricensis Hampe et Gottsche, Linnaea 25 (3): 352, 1852 [1853].

*** subtrib. Cheilolejeuneinae Gradst.

* Aureolejeunea R.M.Schust., Phytologia 39 (6): 428, 1978. 238

*** Aureolejeunea aurifera R.M.Schust., Phytologia 39 (6): 429, 1978.

*** Aureolejeunea lumae (Herzog) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 121, 1985. Bas.: Brachiolejeunea lumae Herzog, Beih. Bot. Centralbl. 60B (1/2): 15, 1939.

*** Aureolejeunea paramicola (Herzog) R.M.Schust., Phytologia 61 (7): 446, 1987. Bas.: Brachiolejeunea paramicola Herzog, Hedwigia 74 (2): 95, 1934.

*** Aureolejeunea quinquecarinata R.M.Schust., Phytologia 39 (6): 429, 1978.

*** Aureolejeunea tonduzana (Steph.) Gradst., Phytotaxa 76 (3): 46, 2013. Bas.: Archilejeunea tonduzana Steph., Sp. Hepat. (Stephani) 4: 721, 1911.

*** Cheilolejeunea (Spruce) Steph., Bot. Gaz. 15 (11): 284, 1890. Bas.: Lejeunea subg. Cheilolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 251, 1884. 239

** Cheilolejeunea sandvicensis (Prantl) Steph., Bull. Herb. Boissier 5 (10): 842, 1897. Bas.: Lejeunea sandvicensis Prantl, Hedwigia 29: xvii, 1890.

** subg. Cheilolejeunea

*** Cheilolejeunea acanthina (Spruce) Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 62, 2009. Bas.: Lejeunea acanthina Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 182, 1884.

*** Cheilolejeunea adnata (Kunze ex Lehm.) Grolle, J. Bryol. 9 (4): 529, 1977 [1978]. Bas.: Jungermannia adnata Kunze ex Lehm., Nov. Stirp. Pug. 6: 46, 1834.

** Cheilolejeunea adnata var. autoica Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 64, 2009.

** Cheilolejeunea albovirens (Hook.f. et Taylor) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 127, 1962. Bas.: Jungermannia albovirens Hook.f. et Taylor, London J. Bot. 3: 397, 1844.

*** Cheilolejeunea aneogyna (Spruce) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 440, 1900. Bas.: Lejeunea aneogyna Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 254, 1884.

** Cheilolejeunea ascensionis (Hook.f. et Taylor) Grolle, Haussknechtia 4: 45, 1988. Bas.: Jungermannia ascensionis Hook.f. et Taylor, London J. Bot. 4: 91, 1845.

*** Cheilolejeunea asperiflora (Spruce) Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 62, 2009. Bas.: Lejeunea asperiflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 183, 1884.

** Cheilolejeunea australis Solari, Comun. Mus. Argent. Ci. Nat. “Bernardino Rivadavia,” Ci. Bot. 2 (11): 70, 1981.

** Cheilolejeunea baumannii Hürl., Bauhinia 11 (3): 160, 1995.

*** Cheilolejeunea beyrichii (Lindenb.) M.E.Reiner, Nova Hedwigia 83 (3/4): 474, 2006. Bas.: Lejeunea beyrichii Lindenb., Syn. Hepat. 3: 371, 1845.

** Cheilolejeunea boninensis Mizut., J. Hattori Bot. Lab. 51: 153, 1982.

*** Cheilolejeunea caducifolia (Gradst. et Schäf.-Verw.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010. Bas.: Leucolejeunea caducifolia Gradst. et Schäf.-Verw., J. Hattori Bot. Lab. 74: 64, 1993.

** Cheilolejeunea camerunensis S.W.Arnell, Svensk Bot. Tidskr. 52 (1): 63, 1958.

** Cheilolejeunea celebensis (Steph.) Mizut., J. Hattori Bot. Lab. 36: 157, 1972 [1973]. Bas.: Trachylejeunea celebensis Steph., Sp. Hepat. (Stephani) 5: 312, 1913.

** Cheilolejeunea chenii R.L.Zhu et M.L.So, Taxon 48 (4): 663, 1999. Nom. nov. pro Neurolejeunea fukiensis P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 227, 1964.

*** Cheilolejeunea clypeata (Schwein.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010. Bas.: Jungermannia clypeata Schwein., Spec. Fl. Amer. Crypt.: 12, 1821.

** Cheilolejeunea compressa (Herzog) Grolle, Bryophyt. Biblioth. 48: 38, 1995. Bas.: Strepsilejeunea compressa Herzog, Bot. Not. 100 (4): 325, 1947.

*** Cheilolejeunea conchifolia (A.Evans) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010. Bas.: Archilejeunea conchifolia A.Evans, Mem. Torrey Bot. Club 8 (2): 128, 1902.

*** Cheilolejeunea cordigera (Steph.) Grolle, J. Bryol. 9 (4): 530, 1977 [1978]. Bas.: Hygrolejeunea cordigera Steph., Hedwigia 35 (3): 100, 1896.

** Cheilolejeunea coronalis (Gottsche) R.M.Schust., Phytologia 45 (5): 431, 1980. Bas.: Lejeunea coronalis Gottsche, Syn. Hepat. 3: 361, 1845.

* Cheilolejeunea curvatilobula (Herzog) Grolle, J. Bryol. 21 (1): 41, 1999. Bas.: Harpalejeunea curvatilobula Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 184, 1955.

*** Cheilolejeunea decursiva (Sande Lac.) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963. Bas.: Lejeunea decursiva Sande Lac., Ned. Kruidk. Arch. 3: 522, 1855.

*** Cheilolejeunea discoidea (Lehm. et Lindenb.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961. Bas.: Jungermannia discoidea Lehm. et Lindenb., Nov. Stirp. Pug. 6: 47, 1834.

** Cheilolejeunea diversifolia Augier, Ann. Fac. Sci. Univ. Féd. Cameroun 11: 66, 1972.

** Cheilolejeunea ecarinata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (1/2): 11, 1984.

** Cheilolejeunea erostrata R.M.Schust., Phytologia 39 (6): 427, 1978.

*** Cheilolejeunea exinnovata E.W.Jones, J. Bryol. 12 (1): 37, 1982.

*** Cheilolejeunea fragrantissima (Spruce) R.M.Schust., Phytologia 45 (5): 431, 1980. Bas.: Lejeunea fragrantissima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 243, 1884.

* Cheilolejeunea fukiensis (P.C.Chen et P.C.Wu) Piippo, J. Hattori Bot. Lab. 68: 133, 1990. Bas.: Euosmolejeunea fukiensis P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 232, 1964. 240

* Cheilolejeunea galliotii Steph., Sp. Hepat. (Stephani) 5: 656, 1914.

** Cheilolejeunea germanii (Besch. et Spruce) Grolle, Acta Bot. Fenn. 125: 64, 1984. Bas.: Lejeunea germanii Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxvii, 1889 [1890].

** Cheilolejeunea ghatensis G.Asthana, S.C.Srivast. et A.K.Asthana, Lindbergia 20 (2/3): 132, 1995 [1996].

*** Cheilolejeunea gradsteinii (Grolle et Piippo) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010. Bas.: Leucolejeunea gradsteinii Grolle et Piippo, Ann. Bot. Fenn. 27 (2): 122, 1990.

** Cheilolejeunea grandibracteata Steph., Sp. Hepat. (Stephani) 5: 657, 1914.

** Cheilolejeunea hamlinii Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 212, 1982. Nom. nov. pro Strepsilejeunea curnowii Steph., Hedwigia 35 (3): 129, 1896.

** Cheilolejeunea hawaica Steph., Bull. Herb. Boissier 5 (10): 847, 1897.

* Cheilolejeunea herzogiana Steph., Biblioth. Bot. 87 (2): 267, 1916.

** Cheilolejeunea implexicaulis (Hook.f. et Taylor) R.M.Schust., J. Hattori Bot. Lab. 26: 245, 1963. Bas.: Jungermannia implexicaulis Hook.f. et Taylor, London J. Bot. 3: 397, 1844.

*** Cheilolejeunea insecta Grolle et Gradst., Taxon 50 (4): 1071, 2001 [2002].

** Cheilolejeunea insignis Jovet-Ast et Tixier, Rev. Bryol. Lichénol. 31 (1/2): 25, 1962.

** Cheilolejeunea intricata (Steph.) J.J.Engel, Bryologist 79 (4): 514, 1976 [1977]. Bas.: Harpalejeunea intricata Steph., Sp. Hepat. (Stephani) 5: 269, 1913.

** Cheilolejeunea invaginata R.M.Schust., Phytologia 39 (6): 427, 1978.

** Cheilolejeunea jamaicensis Steph., Hedwigia 34 (5): 241, 1895.

** Cheilolejeunea japonica (Horik.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010. Bas.: Archilejeunea japonica Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 84, 1932.

** Cheilolejeunea kitagawae W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010. Nom. nov. pro Leucolejeunea paroica N.Kitag., Acta Phytotax. Geobot. 18 (7): 190, 1960.

*** Cheilolejeunea lacerata C.J.Bastos et Gradst., J. Bryol. 28 (2): 133, 2006.

** Cheilolejeunea larsenii Mizut., Dansk Bot. Ark. 27 (1): 95, 1969.

* Cheilolejeunea laurentii Steph., Sp. Hepat. (Stephani) 5: 647, 1914. 241

** Cheilolejeunea leptophylla (Ångstr.) Steph., Sp. Hepat. (Stephani) 5: 657, 1914. Bas.: Lejeunea leptophylla Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 86, 1876 [1877].

* Cheilolejeunea longiflora (Taylor) R.M.Schust., Phytologia 45 (5): 431, 1980. Bas.: Lejeunea longiflora Taylor, London J. Bot. 5: 396, 1846. 242

** Cheilolejeunea longispina (Herzog) R.M.Schust., Beih. Nova Hedwigia 9: 111, 1963. Bas.: Harpalejeunea longispina Herzog, Rev. Bryol. Lichénol. 23 (1/2): 61, 1954.

** Cheilolejeunea loriana (Steph.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010. Bas.: Symbiezidium lorianum Steph., Sp. Hepat. (Stephani) 5: 106, 1912.

** Cheilolejeunea ludoviciae Steph., Sp. Hepat. (Stephani) 5: 668, 1914.

** Cheilolejeunea lurida (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 658, 1914. Bas.: Lejeunea lurida Lindenb., Syn. Hepat. 3: 379, 1845.

** Cheilolejeunea macroloba (Herzog) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 212, 1982. Bas.: Strepsilejeunea macroloba Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 742, 1942.

*** Cheilolejeunea malaccensis (G.Hoffm.) Xiao L.He, Ann. Bot. Fenn. 33 (1): 59, 1996. Bas.: Pycnolejeunea malaccensis G.Hoffm., Ann. Bryol. 8: 118, 1935.

** Cheilolejeunea mammifera R.M.Schust., Phytologia 45 (5): 429, 1980.

*** Cheilolejeunea mariana (Gottsche) B.M.Thiers et Gradst., Mem. New York Bot. Gard. 52: 75, 1989. Bas.: Lejeunea mariana Gottsche, Syn. Hepat. 3: 337, 1845.

** Cheilolejeunea mexicana Steph., Sp. Hepat. (Stephani) 6: 417, 1923.

* Cheilolejeunea micholitzii (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961. Bas.: Pycnolejeunea micholitzii Steph., Sp. Hepat. (Stephani) 5: 627, 1914. 243

** Cheilolejeunea microscypha (Hook.f. et Taylor) M.Wigginton, J. Bryol. 34 (4): 269, 2012. Bas.: Jungermannia microscypha Hook.f. et Taylor, London J. Bot. 4: 90, 1845.

*** Cheilolejeunea mizutanii W.Ye et R.L.Zhu, J. Bryol. 32 (4): 281, 2010. Nom. nov. pro Hygrolejeunea decurrens Steph., Hedwigia 35 (3): 101, 1896.

*** Cheilolejeunea neblinensis Ilk.-Borg. et Gradst., Nova Hedwigia 87 (3/4): 522, 2008.

** Cheilolejeunea nietneri (Steph.) Mizut., J. Hattori Bot. Lab. 37: 191, 1973. Bas.: Harpalejeunea nietneri Steph., Sp. Hepat. (Stephani) 6: 392, 1923.

*** Cheilolejeunea norisiae G.Dauphin et Gradst., J. Bryol. 25 (4): 259, 2003.

** Cheilolejeunea novaezelandiae R.M.Schust., Phytologia 56 (7): 459, 1985.

*** Cheilolejeunea obcordata Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 209, 1931.

** Cheilolejeunea obtruncata (Mont.) Solari, J. Hattori Bot. Lab. 54: 539, 1983. Bas.: Lejeunea obtruncata Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 354, 1845.

** Cheilolejeunea obtusa (Herzog) Solari, J. Hattori Bot. Lab. 54: 537, 1983. Bas.: Harpalejeunea obtusa Herzog, Rev. Bryol. Lichénol. 29 (3/4): 193, 1960 [1961].

** Cheilolejeunea obtusifolia (Steph.) S.Hatt., J. Hattori Bot. Lab. 18: 116, 1957. Bas.: Harpalejeunea obtusifolia Steph., Sp. Hepat. (Stephani) 5: 265, 1913.

** Cheilolejeunea occlusa (Herzog) T.Kodama et N.Kitag., Bull. Osaka Mus. Nat. Hist. 28: 40, 1974. Bas.: Strepsilejeunea occlusa Herzog, Trans. Brit. Bryol. Soc. 1 (4): 320, 1950.

** Cheilolejeunea orientalis (Gottsche) Mizut., J. Hattori Bot. Lab. 35: 399, 1972. Bas.: Lejeunea orientalis Gottsche, Syn. Hepat. 3: 371, 1845.

** Cheilolejeunea ovistipula Steph., Hedwigia 34 (5): 244, 1895.

** Cheilolejeunea panurensis (Spruce) Steph., Sp. Hepat. (Stephani) 5: 660, 1914. Bas.: Lejeunea panurensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 255, 1884.

** Cheilolejeunea papillata Solari, Comun. Mus. Argent. Ci. Nat. “Bernardino Rivadavia,” Ci. Bot. 2 (11): 72, 1981.

** Cheilolejeunea paroica Mizut., J. Hattori Bot. Lab. 46: 364, 1979.

** Cheilolejeunea piriflora Schiffn., Bot. Jahrb. Syst. 23 (5): 592, 1897. Based on: Lejeunea piriflora Gottsche ex Pol., J. Bot. 15: 227, 1877, nom. inval.

*** Cheilolejeunea polystachya (Spruce) Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 62, 2009. Bas.: Lejeunea polystachya Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 250, 1884.

** Cheilolejeunea renneri (G.Hoffm.) Xiao L.He, Ann. Bot. Fenn. 33 (1): 62, 1996. Bas.: Pycnolejeunea renneri G.Hoffm., Ann. Bryol. 8: 117, 1935.

*** Cheilolejeunea rotundistipula (Lindenb. ex Lehm.) Malombe, Acta Bot. Hung. 51 (3/4): 322, 2009. Bas.: Jungermannia rotundistipula Lindenb. ex Lehm., Linnaea 4: 360, 1829.

** Cheilolejeunea rufescens (Lindenb.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 212, 1982. Bas.: Lejeunea rufescens Lindenb., Syn. Hepat. 3: 366, 1845.

** Cheilolejeunea stenoschiza (Ångstr.) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 436, 1900. Bas.: Lejeunea stenoschiza Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 26, 1872.

** Cheilolejeunea subcrenulata (Spruce) R.M.Schust., Phytologia 45 (5): 431, 1980. Bas.: Lejeunea subcrenulata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 245, 1884.

* Cheilolejeunea suborbicularis (Herzog) H.A.Mill., Bonner et Bischl., Nova Hedwigia 4: 559, 1962 [1963]. Bas.: Microlejeunea suborbicularis Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 65, 1950 [1951].

*** Cheilolejeunea suprema (Grolle et Piippo) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 281, 2010. Bas.: Leucolejeunea suprema Grolle et Piippo, Ann. Bot. Fenn. 27 (2): 123, 1990.

** Cheilolejeunea surrepens (Mitt.) E.W.Jones, J. Bryol. 9 (1): 49, 1976. Bas.: Lejeunea surrepens Mitt., Philos. Trans. 168: 399, 1879.

*** Cheilolejeunea turgida (Mitt.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 281, 2010. Bas.: Lejeunea turgida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 110, 1860 [1861].

* Cheilolejeunea ulugurica Malombe, Eb.Fisch. et Pócs, Acta Biol. Pl. Agr. 1: 24, 2010 [2011]. 244

*** Cheilolejeunea unciloba (Lindenb.) Malombe, Acta Bot. Hung. 51 (3/4): 325, 2009. Bas.: Lejeunea unciloba Lindenb., Syn. Hepat. 3: 331, 1845.

** Cheilolejeunea upoluensis S.W.Arnell, Svensk Bot. Tidskr. 50 (3): 516, 1956.

** Cheilolejeunea valenciae (Steph.) Xiao L.He, Ann. Bot. Fenn. 33 (1): 55, 1996. Bas.: Pycnolejeunea valenciae Steph., Sp. Hepat. (Stephani) 5: 605, 1914.

** Cheilolejeunea virescens (Lehm. et Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 662, 1914. Bas.: Lejeunea virescens Lehm. et Lindenb., Nov. Stirp. Pug. 7: 21, 1838.

* Cheilolejeunea viridis Steph., Sp. Hepat. (Stephani) 5: 673, 1914.

** Cheilolejeunea warnstorffii (Steph.) Solari, Darwiniana 20 (3/4): 387, 1976. Bas.: Strepsilejeunea warnstorffii Steph., Hedwigia 35 (3): 131, 1896.

* Cheilolejeunea wrightii Steph., Sp. Hepat. (Stephani) 5: 662, 1914.

*** Cheilolejeunea xanthocarpa (Lehm. et Lindenb.) Malombe, Acta Bot. Hung. 51 (3/4): 326, 2009. Bas.: Jungermannia xanthocarpa Lehm. et Lindenb., Nov. Stirp. Pug. 5: 8, 1833.

** Cheilolejeunea xanthophylla (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 663, 1914. Bas.: Lejeunea xanthophylla Lindenb., Syn. Hepat. 3: 370, 1845.

** subg. Euosmolejeunea (Spruce) Kachroo, Ceylon J. Sci., Biol. Sci. 8 (1): 6, 1968. Bas.: Lejeunea subg. Euosmolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 241, 1884.

** Cheilolejeunea aciculifera R.M.Schust., Phytologia 45 (5): 428, 1980.

** Cheilolejeunea acutangula (Nees) Grolle, J. Hattori Bot. Lab. 45: 173, 1979. Bas.: Jungermannia acutangula Nees, Fl. Bras. (Martius) 1 (1): 357, 1833.

*** Cheilolejeunea asperrima (Steph.) Grolle, J. Hattori Bot. Lab. 58: 197, 1985. Bas.: Taxilejeunea asperrima Steph., Biblioth. Bot. 87 (2): 259, 1916.

** Cheilolejeunea birmensis (Steph.) Mizut., J. Hattori Bot. Lab. 27: 139, 1964. Bas.: Strepsilejeunea birmensis Steph., Sp. Hepat. (Stephani) 5: 286, 1913.

** Cheilolejeunea campbelliensis (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 245, 1963. Bas.: Strepsilejeunea campbelliensis Steph., Hedwigia 35 (3): 128, 1896.

** Cheilolejeunea cedercreutzii (H.Buch et Perss.) Grolle, Feddes Repert. 87 (3/4): 188, 1976. Bas.: Euosmolejeunea cedercreutzii H.Buch et Perss., Bryophyt. Azoren Madeira: 9, 1941.

** Cheilolejeunea choachina (Gottsche) Gradst., Mem. New York Bot. Gard. 84: 69, 1999. Bas.: Lejeunea choachina Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 156, 1864.

*** Cheilolejeunea clausa (Nees et Mont.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 863, 1980. Bas.: Lejeunea clausa Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 337, 1840.

** Cheilolejeunea clavata Mizut., J. Hattori Bot. Lab. 46: 361, 1979.

*** Cheilolejeunea comans (Spruce) R.M.Schust., Phytologia 45 (5): 431, 1980. Bas.: Lejeunea comans Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 246, 1884.

** Cheilolejeunea convexa (S.W.Arnell) S.W.Arnell, Hepat. South Africa: 209, 1963. Bas.: Lejeunea convexa S.W.Arnell, Bot. Not. 106: 272, 1953.

** Cheilolejeunea cookiensis (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961. Bas.: Pycnolejeunea cookiensis Steph., Sp. Hepat. (Stephani) 5: 617, 1914.

** Cheilolejeunea cordistipula (Steph.) Grolle ex E.W.Jones, J. Bryol. 13 (3): 395, 1985. Bas.: Strepsilejeunea cordistipula Steph., Sp. Hepat. (Stephani) 5: 276, 1913.

** Cheilolejeunea fischeri Malombe, Acta Bot. Hung. 51 (3/4): 319, 2009.

** Cheilolejeunea gaoi R.L.Zhu, M.L.So et Grolle, Bryologist 103 (3): 499, 2000.

** Cheilolejeunea hallingii B.M.Thiers, Brittonia 44 (2): 160, 1992.

*** Cheilolejeunea inflexa (Hampe) Grolle, J. Hattori Bot. Lab. 45: 174, 1979. Bas.: Lejeunea inflexa Hampe, Nov. Stirp. Pug. 7: 22, 1838.

*** Cheilolejeunea intertexta (Lindenb.) Steph., Bull. Herb. Boissier 5 (2): 79, 1897. Bas.: Lejeunea intertexta Lindenb., Syn. Hepat. 3: 379, 1845.

*** Cheilolejeunea krakakammae (Lindenb.) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963. Bas.: Lejeunea krakakammae Lindenb., Syn. Hepat. 3: 353, 1845.

*** Cheilolejeunea laciniata D.F.Peralta et M.E.Reiner, Bryologist 116 (1): 54, 2013.

*** Cheilolejeunea laevicalyx (J.B.Jack et Steph.) Grolle, J. Hattori Bot. Lab. 65: 403, 1988. Bas.: Strepsilejeunea laevicalyx J.B.Jack et Steph., Sp. Hepat. (Stephani) 5: 283, 1913.

** Cheilolejeunea laeviuscula (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 668, 1914. Bas.: Lejeunea laeviuscula Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 114, 1860 [1861].

*** Cheilolejeunea lindenbergii (Gottsche) Mizut., J. Hattori Bot. Lab. 33: 226, 1970. Bas.: Lejeunea lindenbergii Gottsche, Syn. Hepat. 3: 336, 1845.

** Cheilolejeunea mimosa (Hook.f. et Taylor) R.M.Schust., J. Hattori Bot. Lab. 26: 245, 1963. Bas.: Jungermannia mimosa Hook.f. et Taylor, London J. Bot. 3: 398, 1844.

** Cheilolejeunea nana R.M.Schust., Phytologia 39 (6): 426, 1978.

** Cheilolejeunea ngongensis Malombe et Pócs, Acta Bot. Hung. 51 (3/4): 317, 2009.

** Cheilolejeunea nipponica (S.Hatt.) S.Hatt., Misc. Bryol. Lichenol. 1 (14): 1, 1957. Bas.: Strepsilejeunea nipponica S.Hatt., Bull. Tokyo Sci. Mus. 11: 134, 1944.

*** Cheilolejeunea omphalogastria Pócs, Trop. Bryol. 9: 131, 1994.

*** Cheilolejeunea oncophylla (Ångstr.) Grolle et M.E.Reiner, J. Bryol. 19 (4): 781, 1997. Bas.: Lejeunea oncophylla Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 86, 1876 [1877].

** Cheilolejeunea ornata C.J.Bastos, J. Bryol. 33 (1): 86, 2011.

** Cheilolejeunea osumiensis (S.Hatt.) Mizut., Misc. Bryol. Lichenol. 8 (7): 148, 1980. Bas.: Euosmolejeunea osumiensis S.Hatt., Bull. Tokyo Sci. Mus. 11: 105, 1944.

** Cheilolejeunea pluriplicata (Pearson) R.M.Schust., Phytologia 45 (5): 430, 1980. Bas.: Lejeunea pluriplicata Pearson, Forh. Vidensk.-Selsk. Kristiania 1887 (9): 5, 1887.

** Cheilolejeunea pocsii E.W.Jones, J. Bryol. 15 (1): 156, 1988.

** Cheilolejeunea polyantha A.Evans, Mem. Torrey Bot. Club 8 (2): 141, 1902.

** Cheilolejeunea polyantha var. caduciloba R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 887, 1980.

*** Cheilolejeunea revoluta (Herzog) Gradst. et Grolle, J. Hattori Bot. Lab. 74: 59, 1993. Bas.: Pycnolejeunea revoluta Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 193, 1955.

*** Cheilolejeunea rigidula (Nees ex Mont.) R.M.Schust., Castanea 36 (2): 102, 1971. Bas.: Lejeunea rigidula Nees ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 336, 1840.

** Cheilolejeunea ruwenzorensis (S.W.Arnell) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963. Bas.: Euosmolejeunea ruwenzorensis S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 529, 1956.

** Cheilolejeunea ryukyuensis Mizut., J. Hattori Bot. Lab. 51: 162, 1982.

** Cheilolejeunea subopaca (Mitt.) Mizut., J. Hattori Bot. Lab. 26: 183, 1963. Bas.: Lejeunea subopaca Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 116, 1860 [1861].

*** Cheilolejeunea trifaria (Reinw., Blume et Nees) Mizut., J. Hattori Bot. Lab. 27: 132, 1964. Bas.: Jungermannia trifaria Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 226, 1824 [1825].

* Cheilolejeunea udarii G.Asthana, S.C.Srivast. et A.K.Asthana, Lindbergia 20 (2/3): 142, 1995 [1996]. 245

** Cheilolejeunea usambarana (Steph.) Grolle, J. Hattori Bot. Lab. 46: 344, 1979. Bas.: Strepsilejeunea usambarana Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 731, 1913.

** Cheilolejeunea verrucosa Steph., Sp. Hepat. (Stephani) 5: 673, 1914.

** subg. Renilejeunea R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963.

*** Cheilolejeunea montagnei (Gottsche ex Mont.) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963. Bas.: Lejeunea montagnei Gottsche ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 261, 1843.

** subg. Xenolejeunea Kachroo et R.M.Schust., J. Linn. Soc., Bot. 56 (368): 496, 1961.

** Cheilolejeunea ceylanica (Gottsche) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961. Bas.: Lejeunea ceylanica Gottsche, Syn. Hepat. 3: 359, 1845.

** Cheilolejeunea eximia (Jovet-Ast et Tixier) R.L.Zhu et M.L.So, Beih. Nova Hedwigia 121: 114, 2001. Bas.: Pycnolejeunea eximia Jovet-Ast et Tixier, Rev. Bryol. Lichénol. 31 (1/2): 31, 1962.

*** Cheilolejeunea falsinervis (Sande Lac.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961. Bas.: Lejeunea falsinervis Sande Lac., Ned. Kruidk. Arch. 3: 421, 1854 [1855].

** Cheilolejeunea gardneri (Mitt.) Mizut., J. Hattori Bot. Lab. 26: 181, 1963. Bas.: Lejeunea gardneri Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 115, 1860 [1861].

** Cheilolejeunea gigantea (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961. Bas.: Pycnolejeunea gigantea Steph., Hedwigia 35 (3): 125, 1896.

** Cheilolejeunea huerlimannii Tixier, Misc. Bryol. Lichenol. 9 (9): 184, 1983.

** Cheilolejeunea hyalomarginata R.L.Zhu et Frank Müll., Bryologist 115 (2): 218, 2012.

*** Cheilolejeunea incisa (Gottsche) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961. Bas.: Lejeunea incisa Gottsche, Syn. Hepat. 3: 360, 1845.

** Cheilolejeunea incisa var. teretiflora B.M.Thiers, J. Hattori Bot. Lab. 82: 321, 1997.

** Cheilolejeunea longidens (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961. Bas.: Pycnolejeunea longidens Steph., Sp. Hepat. (Stephani) 5: 634, 1914.

** Cheilolejeunea meyeniana (Nees, Lindenb. et Gottsche) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961. Bas.: Lejeunea meyeniana Nees, Lindenb. et Gottsche, Observ. bot.: 472, 1843.

** Cheilolejeunea obtusilobula (S.Hatt.) S.Hatt., Misc. Bryol. Lichenol. 1 (14): 2, 1957. Bas.: Pycnolejeunea obtusilobula S.Hatt., J. Hattori Bot. Lab. 3: 44, 1948 [1950].

** Cheilolejeunea parvidens B.M.Thiers, J. Hattori Bot. Lab. 82: 323, 1997.

** Cheilolejeunea streimannii Pócs et Ninh, Acta Bot. Hung. 47 (1/2): 162, 2005.

*** Cheilolejeunea trapezia (Nees) Kachroo et R.M.Schust., J. Linn. Soc., Bot. 56 (368): 509, 1961. Bas.: Jungermannia trapezia Nees, Enum. Pl. Crypt. Javae: 41, 1830.

** Cheilolejeunea ventricosa (Schiffn. ex P.Syd.) Xiao L.He, Acta Bot. Fenn. 163: 60, 1999. Bas.: Pycnolejeunea ventricosa Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894.

*** Cheilolejeunea vittata (Steph. ex G.Hoffm.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961. Bas.: Pycnolejeunea vittata Steph. ex G.Hoffm., Ann. Bryol. 8: 115, 1935.

Incertae sedis

** Cheilolejeunea gottscheana C.J.Bastos, J. Bryol. 34 (4): 316, 2012. Nom. nov. pro Strepsilejeunea lindenbergii Steph., Hedwigia 35 (3): 130, 1896.

* Cheilolejeunea heteroclada (Spruce) Schiffn., Hepat. (Engl.-Prantl): 124, 1893. Bas.: Lejeunea heteroclada Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 256, 1884. 246

** Cheilolejeunea minutilobula Amakawa, J. Jap. Bot. 35 (12): 365, 1960.

** Cheilolejeunea norrisii (Grolle) M.A.M.Renner, Bryologist 115 (4): 550, 2012. Bas.: Lejeunea norrisii Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 215, 1982.

** Cheilolejeunea oscilla M.A.M.Renner, Bryologist 115 (4): 551, 2012.

** Cheilolejeunea papulosa Schiffn., Hepat. (Engl.-Prantl): 124, 1893. Nom. nov. pro Lejeunea papulosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 258, 1884, nom. illeg.

*** Cheilolejeunea riparia (Steph.) M.E.Reiner, Nova Hedwigia 95 (3/4): 467, 2012. Bas.: Potamolejeunea riparia Steph., Sp. Hepat. (Stephani) 5: 639, 1914.

*** Cheilolejeunea rotalis (Hook.f. et Taylor) M.Wigginton, J. Bryol. 34 (4): 270, 2012. Bas.: Jungermannia rotalis Hook.f. et Taylor, London J. Bot. 4: 89, 1845.

** Cheilolejeunea tenerrima (Steph.) C.J.Bastos, J. Bryol. 34 (4): 317, 2012. Bas.: Strepsilejeunea tenerrima Steph., Sp. Hepat. (Stephani) 5: 286, 1913.

** Cheilolejeunea urubuensis (Zartman et I.L.Ackerman) R.L.Zhu et Y.M.Wei, Phytotaxa 152 (1): 50, 2013. Bas.: Vitalianthus urubuensis Zartman et I.L.Ackerman, Bryologist 105 (2): 267, 2002.

* Cyrtolejeunea A.Evans, Bull. Torrey Bot. Club 30 (10): 553, 1903.

*** Cyrtolejeunea holostipa (Spruce) A.Evans, Bull. Torrey Bot. Club. 30 (10): 553, 1903. Bas.: Lejeunea holostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 171.

** Cystolejeunea A.Evans, Bull. Torrey Bot. Club 33 (1): 16, 1906)

*** Cystolejeunea lineata (Lehm. et Lindenb.) A.Evans, Bull. Torrey Bot. Club. 33 (1): 17, 1906. Bas.: Jungermannia lineata Lehm. et Lindenb., Nov. Stirp. Pug. 4: 53, 1832.

* Omphalanthus Lindenb. et Nees, Syn. Hepat. 2: 303, 1845. 247

** Omphalanthus baracoensis Mustelier, M.E.Reiner et Gradst., J. Bryol. 29 (2): 95, 2007.

*** Omphalanthus filiformis (Sw.) Nees, Syn. Hepat. 2: 304, 1845. Bas.: Jungermannia filiformis Sw., Prodr. (Swartz): 144, 1788.

** Omphalanthus filiformis var. platycoleus (Herzog) Gradst., Phytotaxa 76 (3): 46, 2013. Bas.: Omphalanthus platycoleus Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 171, 1955.

** Omphalanthus filiformis var. wallisii (Prantl) Gradst., Phytotaxa 76 (3): 46, 2013. Bas.: Lejeunea wallisii Prantl, Hedwigia 31: xvii, 1892.

** Omphalanthus huanucensis (Gottsche) Gradst., Beih. Nova Hedwigia 80: 109, 1985. Bas.: Lejeunea huanucensis Gottsche, Syn. Hepat. 3: 335, 1845.

*** Omphalanthus jackii (Prantl) Gradst., Proc. Kon. Ned. Akad. Wetensch. C 80: 410, 1977. Bas.: Lejeunea jackii Prantl, Hedwigia 31: xvii, 1892.

*** Omphalanthus ovalis (Lindenb. et Gottsche) Gradst., Proc. Kon. Ned. Akad. Wetensch. C 80: 411, 1977. Bas.: Lejeunea ovalis Lindenb. et Gottsche, Syn. Hepat. 5: 754, 1847.

*** Omphalanthus roccatii (Gola) R.M.Schust., Beih. Nova Hedwigia 9: 96, 1963. Bas.: Acrolejeunea roccatii Gola, Ann. Bot. (Rome) 6 (2): 275, 1907.

*** subtrib. Cololejeuneinae Gradst.

** Aphanotropis Herzog, Trans. Brit. Bryol. Soc. 2 (1): 63, 1952.

*** Aphanotropis saxicola Herzog, Trans. Brit. Bryol. Soc. 2 (1): 63, 1952.

** Calatholejeunea K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 8, 1928.

*** Calatholejeunea lamii Mizut., J. Hattori Bot. Lab. 56: 334, 1984.

*** Calatholejeunea paradoxa (Schiffn.) K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 8, 1928. Bas.: Lejeunea paradoxa Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 243, 1893.

*** Cololejeunea (Spruce) Steph., Hedwigia 30 (5): 208, 1891. Bas.: Lejeunea subg. Cololejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 291, 1884. 248

** Cololejeunea micrandroecia (Spruce) M.Menzel, Willdenowia 14: 492, 1984 [1985]. Bas.: Lejeunea micrandroecia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 298, 1884.

* subg. Aphanolejeunea (A.Evans) Pócs, Phytotaxa 202 (1): 64, 2015. Bas.: Aphanolejeunea A.Evans, Bull. Torrey Bot. Club 38 (6): 272, 1911.

*** Cololejeunea berneckerae Pócs, Polish Bot. J. 54 (1): 4, 2009. Nom. nov. pro Aphanolejeunea pocsii Bern.-Lück., Nova Hedwigia 66 (1/2): 168, 1998.

*** Cololejeunea cingens (Herzog) Bernecker et Pócs, Polish Bot. J. 54 (1): 4, 2009. Bas.: Aphanolejeunea cingens Herzog, Svensk Bot. Tidskr. 46 (1): 104, 1952.

*** Cololejeunea cornutissima (R.M.Schust.) Stotler et Crand.-Stotl., Bryologist 80 (3): 411, 1977. Bas.: Aphanolejeunea cornutissima R.M.Schust., Bryologist 59 (3): 217, 1956.

*** Cololejeunea costaricensis (Bern.-Lück.) Bernecker et Pócs, Polish Bot. J. 54 (1): 5, 2009. Bas.: Aphanolejeunea costaricensis Bern.-Lück., Nova Hedwigia 66 (1/2): 164, 1998.

*** Cololejeunea cubensis Pócs, Polish Bot. J. 54 (1): 5, 2009. Nom. nov. pro Aphanolejeunea evansii Herzog, Beih. Bot. Centralbl. 61B (3): 583, 1942.

*** Cololejeunea gracilis (Jovet-Ast) Pócs, Cryptog. Bryol. 29 (3): 233, 2008. Bas.: Aphanolejeunea gracilis Jovet-Ast, Rev. Bryol. Lichénol. 16 (1/2): 21, 1947 [1948].

** Cololejeunea gracilis var. linearifolia (R.M.Schust.) Pócs, Acta Bot. Hung. 56 (1/2): 189, 2014. Bas.: Aphanolejeunea gracilis var. linearifolia R.M.Schust., Phytologia 45 (5): 434, 1980.

*** Cololejeunea grossepapillosa (Horik.) N.Kitag., Hikobia, Suppl. 1: 68, 1981. Bas.: Aphanolejeunea grossepapillosa Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 92, 1932.

** Cololejeunea iwatsukiana (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009. Bas.: Aphanolejeunea iwatsukiana Pócs, Hikobia 11: 457, 1994.

*** Cololejeunea jovetastiana (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009. Bas.: Aphanolejeunea jovetastiana Pócs, Cryptog. Bryol. Lichénol. 5 (3): 251, 1984.

** Cololejeunea lisowskii (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009. Bas.: Aphanolejeunea lisowskii Pócs, Cryptog. Bryol. Lichénol. 5 (3): 259, 1984.

*** Cololejeunea madeirensis Schiffn., Hedwigia 41 (5): 279, 1902.

*** Cololejeunea microscopica (Taylor) Schiffn., Hepat. (Engl.-Prantl): 122, 1893. Bas.: Jungermannia microscopica Taylor, Mackay, Fl. Hibern. 2: 59, 1836.

*** Cololejeunea microscopica var. africana (Pócs) Pócs et Bernecker, Cryptog. Bryol. 29 (3): 234, 2008. Bas.: Aphanolejeunea exigua var. africana Pócs, Cryptog. Bryol. Lichénol. 5 (3): 247, 1984.

*** Cololejeunea microscopica var. exigua (A.Evans) Pócs, Mem. New York Bot. Gard. 76 (4): 73, 2009. Bas.: Aphanolejeunea exigua A.Evans, Bull. Torrey Bot. Club 38 (6): 273, 1911.

*** Cololejeunea minuscula Pócs, Polish Bot. J. 54 (1): 7, 2009. Nom. nov. pro Aphanolejeunea minuta R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 1310, 1980.

** Cololejeunea moramangae Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 241, 1977 [1979].

*** Cololejeunea norrisii (Pócs) Pócs, Polish Bot. J. 54 (1): 7, 2009. Bas.: Aphanolejeunea norrisii Pócs, Acta Bot. Fenn. 165: 95, 1999.

*** Cololejeunea papillosa (K.I.Goebel) Mizut., J. Hattori Bot. Lab. 29: 156, 1966. Bas.: Physocolea papillosa K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 41, 1928.

*** Cololejeunea sicifolia (Gottsche ex A.Evans) Pócs et Bernecker, Polish Bot. J. 54 (1): 8, 2009. Bas.: Aphanolejeunea sicifolia Gottsche ex A.Evans, Bull. Torrey Bot. Club 38 (6): 277, 1911.

*** Cololejeunea sicifolia subsp. jamaicensis (R.M.Schust.) Bernecker et Pócs, Polish Bot. J. 54 (1): 8, 2009. Bas.: Aphanolejeunea jamaicensis R.M.Schust., Phytologia 45 (5): 434, 1980.

*** Cololejeunea sintenisii (Steph.) Pócs, Cryptog. Bryol. 29 (3): 235, 2008. Bas.: Aphanolejeunea sintenisii Steph., Sp. Hepat. (Stephani) 5: 861, 1916.

*** Cololejeunea subsphaeroidea (R.M.Schust.) Pócs, Cryptog. Bryol. 29 (3): 235, 2008. Bas.: Aphanolejeunea subsphaeroidea R.M.Schust., Phytologia 39 (6): 431, 1978.

** Cololejeunea thiersiae (Pócs) Pócs, Polish Bot. J. 54 (1): 9, 2009. Bas.: Aphanolejeunea thiersiae Pócs, Hikobia 11: 459, 1994.

** Cololejeunea veillonii Tixier, Nova Hedwigia 31: 757, 1979.

*** Cololejeunea winkleri (M.I.Morales et A.Lücking) Pócs, Mem. New York Bot. Gard. 76 (4): 78, 2009. Bas.: Aphanolejeunea winkleri M.I.Morales et A.Lücking, Nova Hedwigia 60 (1/2): 120, 1995.

* subg. Austrocololejeunea Tixier, Nova Hedwigia 31: 776, 1979.

** Cololejeunea australis Tixier, Nova Hedwigia 31: 781, 1979.

** Cololejeunea caledonica Gottsche, Hedwigia 34 (5): 246, 1895.

** Cololejeunea sophiana Tixier, Bot. Not. 128: 428, 1975 [1976].

** Cololejeunea virotana Tixier, Nova Hedwigia 31: 777, 1979.

* subg. Chlorocolea R.M.Schust., Beih. Nova Hedwigia 9: 178, 1963.

*** Cololejeunea ceratilobula (P.C.Chen) R.M.Schust., Beih. Nova Hedwigia 9: 179, 1963. Bas.: Leptocolea ceratilobula P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 49, 1955.

*** Cololejeunea desciscens Steph., Hedwigia 34 (5): 248, 1895.

*** Cololejeunea linopteroides H.Rob., Bryologist 67 (4): 457, 1964.

* Cololejeunea rotundilobula (P.C.Wu et P.J.Lin) Piippo, J. Hattori Bot. Lab. 68: 134, 1990. Bas.: Pedinolejeunea rotundilobula P.C.Wu et P.J.Lin, Acta Phytotax. Sin. 16 (2): 69, 1978. 249

*** Cololejeunea sigmoidea Jovet-Ast et Tixier, Rev. Bryol. Lichénol. 31 (1/2): 27, 1962.

** Cololejeunea sigmoidea var. dubia Tixier, Bryophyt. Biblioth. 27: 114, 1985.

*** Cololejeunea standleyi Herzog, Rev. Bryol. Lichénol. 20 (1/2): 172, 1951 [1952].

*** Cololejeunea stylilobula Tixier, Phytotaxa 202 (1): 65, 2015. Based on: Cololejeunea stylilobula Tixier, Bryophyt. Biblioth. 27: 119, 1985, nom. inval.

*** Cololejeunea zangii R.L.Zhu et M.L.So, Syst. Bot. 24 (4): 501, 1999 [2000].

*** subg. Chlorolejeunea Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 81, 1953. 250

** Cololejeunea lacinulata Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 82, 1953.

*** Cololejeunea madothecoides (Steph.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 81, 1953. Bas.: Physocolea madothecoides Steph., Sp. Hepat. (Stephani) 5: 898, 1916.

** Cololejeunea ombrophila Tixier, Nat. Hist. Bull. Siam Soc. 23 (4): 550, 1970.

*** Cololejeunea stotleriana Gradst., Ilk.-Borg. et Vanderp., Bryologist 114 (1): 13, 2011.

*** subg. Chondriolejeunea Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 75, 1953. 251

*** Cololejeunea chinii Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 445, 1973.

*** Cololejeunea pseudostipulata Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894. Based on: Cololejeunea pseudostipulata Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 33, 1890, nom. inval.

*** Cololejeunea shimizui N.Kitag., Acta Phytotax. Geobot. 23 (5/6): 185, 1969.

*** Cololejeunea shimizui var. phangngana N.Kitag., Acta Phytotax. Geobot. 23 (5/6): 187, 1969.

*** subg. Cololejeunea, 252

** Cololejeunea albodentata P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 252, 1964.

** Cololejeunea armata Tixier, Gard. Bull. Singapore 26 (1): 149, 1972.

*** Cololejeunea bhutanica Grolle et Mizut., J. Bryol. 15 (2): 281, 1989.

** Cololejeunea biddlecomiae (Austin) A.Evans, Mem. Torrey Bot. Club 8 (2): 168, 1902. Bas.: Lejeunea biddlecomiae Austin, List. Canad. Hepat.: 5, 1890.

* Cololejeunea caihuaella But et P.C.Wu, Hepat. Fl. Hong Kong: 132, 2009.

*** Cololejeunea calcarea (Lib.) Steph., Bot. Gaz. 17 (6): 171, 1892. Bas.: Lejeunea calcarea Lib., Ann. Gen. Sci. Phys. 6: 373, 1820.

** Cololejeunea capuronii Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 241, 1977 [1979].

** Cololejeunea dinghuiana R.L.Zhu et Y.F.Wang, J. E. China Norm. Univ., Nat. Sci. Ed. 2: 91, 1992.

** Cololejeunea dolichodonta Tixier, Bryophyt. Biblioth. 27: 215, 1985.

*** Cololejeunea dozyana (Sande Lac.) Schiffn., Hedwigia 39 (4): 199, 1900. Bas.: Lejeunea dozyana Sande Lac., Ned. Kruidk. Arch. 3: 522, 1855.

*** Cololejeunea elegans Steph., Hedwigia 30 (5): 208, 1891.

** Cololejeunea falcidentata R.M.Schust., Nova Hedwigia 15: 507, 1968.

** Cololejeunea filidens Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 49, 1953.

* Cololejeunea frahmii Tixier, Trop. Bryol. 11: 63, 1995. 253

*** Cololejeunea grolleana Pócs, J. Hattori Bot. Lab. 48: 312, 1980.

*** Cololejeunea haskarliana (Lehm.) Schiffn., Consp. Hepat. Arch. Ind.: 244, 1898. Bas.: Lejeunea haskarliana Lehm., Nov. Stirp. Pug. 8: 26, 1844.

*** Cololejeunea hyalina G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 25, 2003.

* Cololejeunea kahuziensis Tixier, Trop. Bryol. 11: 63, 1995. 254

*** Cololejeunea karnatakensis G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 26, 2003.

** Cololejeunea kodamae Kamim., Feddes Repert. Spec. Nov. Regni Veg. 58: 55, 1955.

*** Cololejeunea kolombangarae Pócs, Acta Bryolichenol. Asiat. 4: 67, 2011.

** Cololejeunea kolombangarae subsp. sepikensis Pócs, Acta Bryolichenol. Asiat. 4: 68, 2011.

*** Cololejeunea konratii Pócs, Acta Bot. Hung. 54 (1/2): 156, 2012.

*** Cololejeunea kuciana Pócs et Schäf.-Verw., Polish Bot. J. 57 (1): 51, 2012.

*** Cololejeunea longiana Grolle et Mizut., J. Bryol. 15 (2): 284, 1989.

*** Cololejeunea macounii (Spruce) A.Evans, Mem. Torrey Bot. Club 8 (2): 171, 1902. Bas.: Lejeunea macounii Spruce, Bull. Torrey Bot. Club 17 (10): 259, 1890.

** Cololejeunea magillii Pócs, J. Hattori Bot. Lab. 74: 49, 1993.

*** Cololejeunea malanjae Steph., Bot. Jahrb. Syst. 30 (2): 261, 1901.

** Cololejeunea mamillata (Ångstr.) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (11): 184, 1967. Bas.: Lejeunea mamillata Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 14, 1872.

** Cololejeunea mizutaniana Udar et G.Srivast., Misc. Bryol. Lichenol. 9 (7): 138, 1983.

*** Cololejeunea mocambiquensis S.W.Arnell, Mitt. Thüring. Bot. Ges. 1 (1): 7, 1955 [1956].

** Cololejeunea nanhutashanensis J.D.Yang et S.H.Lin, Phytotaxa 177 (1): 56, 2014.

*** Cololejeunea nilgiriensis G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 27, 2003.

*** Cololejeunea ornata A.Evans, Bryologist 41 (4): 73, 1938.

** Cololejeunea parva Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 239, 1977.

*** Cololejeunea planiflora Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 60, 1953.

*** Cololejeunea platyneura (Spruce) A.Evans, Mem. Torrey Bot. Club 8 (2): 172, 1902. Bas.: Lejeunea platyneura Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 299, 1884.

*** Cololejeunea pluridentata P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 105, 1978.

** Cololejeunea pretiosa Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 62, 1953.

*** Cololejeunea pseudocristallina P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 257, 1964.

** Cololejeunea pseudoplagiophylla P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 106, 1978.

** Cololejeunea pseudoschmidtii Tixier, Gard. Bull. Singapore 26 (1): 145, 1972.

** Cololejeunea ramromensis Pócs, Biodiv., biogeogr., nat. conserv. Wallacea N. Guinea II: 113, 2014.

*** Cololejeunea rosellata Mizut., J. Hattori Bot. Lab. 29: 159, 1966.

*** Cololejeunea rossettiana (C.Massal.) Schiffn., Hepat. (Engl.-Prantl): 122, 1893. Bas.: Lejeunea rossettiana C.Massal., Nuovo Giorn. Bot. Ital. 21 (3): 487, 1889.

*** Cololejeunea runssorensis (Steph.) Pócs, Acta Bot. Acad. Sci. Hung. 21 (3/4): 371, 1975. Bas.: Aphanolejeunea runssorensis Steph., Sp. Hepat. (Stephani) 5: 858, 1916.

** Cololejeunea schaeferi Grolle, J. Bryol. 13 (4): 488, 1986.

*** Cololejeunea schmidtii Steph., Bot. Tidsskr. 24 (3): 278, 1902.

** Cololejeunea schmidtii var. acutepapillosa Pócs, Acta Bot. Hung. 54 (1/2): 165, 2012.

** Cololejeunea selaginellicola Tixier, Gard. Bull. Singapore 26 (1): 149, 1972.

*** Cololejeunea serrata (Steph.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 53, 1953. Bas.: Physocolea serrata Steph., Sp. Hepat. (Stephani) 5: 905, 1916.

** Cololejeunea shikokiana (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 101, 1944. Bas.: Physocolea shikokiana Horik., Bot. Mag. (Tokyo) 46 (544): 182, 1932.

** Cololejeunea spinosa (Horik.) Pandé et R.N.Misra, J. Indian Bot. Soc. 22 (2/4): 166, 1943. Bas.: Physocolea spinosa Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 70, 1931.

*** Cololejeunea stellaris Pócs, Acta Bryolichenol. Asiat. 4: 70, 2011.

** Cololejeunea subkodamae Mizut., J. Hattori Bot. Lab. 60: 448, 1986.

** Cololejeunea tamasii Schäf.-Verw., Phytotaxa 60: 9, 2012.

** Cololejeunea tanneri Pócs, Acta Bot. Hung. 31 (1/4): 126, 1985.

*** Cololejeunea tenella Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 55, 1953.

** Cololejeunea tenella var. dentiloba Onr., Bull. Jard. Bot. Natl. Belg. 59 (3/4): 436, 1989.

** Cololejeunea thiersiana Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995. Based on: Cololejeunea thiersiana Tixier, Candollea 46 (2): 291, 1991, nom. inval.

** Cololejeunea tuiwawana Pócs, Acta Bot. Hung. 54 (1/2): 165, 2012.

*** Cololejeunea verrucosa Steph., Hedwigia 34 (5): 253, 1895.

** Cololejeunea verrucosa var. rectispina (Herzog) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 55, 1953. Bas.: Physocolea verrucosa var. rectispina Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 214, 1931.

*** Cololejeunea zenkeri (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 420, 1954. Bas.: Aphanolejeunea zenkeri Steph., Sp. Hepat. (Stephani) 5: 858, 1916.

* subg. Cryptolejeunea Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 77, 1953.

*** Cololejeunea angustiflora (Steph.) Mizut., J. Hattori Bot. Lab. 28: 113, 1965. Bas.: Leptocolea angustiflora Steph., Sp. Hepat. (Stephani) 5: 848, 1916.

** Cololejeunea drepanolejeuneoides (Horik.) R.M.Schust., Beih. Nova Hedwigia 9: 174, 1963. Bas.: Boninoleptocolea drepanolejeuneoides Horik., Bot. Mag. (Tokyo) 50 (598): 558, 1936. 255

** Cololejeunea hattoriana Mizut. et Pócs, Ann. Bot. Fenn. 31 (3): 188, 1994. Nom. nov. pro Campylolejeunea pusilla Mizut., J. Hattori Bot. Lab. 29: 154, 1966.

*** Cololejeunea inflectens (Mitt.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 79, 1953. Bas.: Lejeunea inflectens Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 117, 1860 [1861].

** Cololejeunea mouensis (Tixier) H.A.Mill., Phytologia 47 (4): 322, 1981. Bas.: Campylolejeunea mouensis Tixier, Nova Hedwigia 31: 727, 1979.

*** Cololejeunea vesicaria (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 247, 1898. Bas.: Lejeunea vesicaria Sande Lac., Syn. hepat. jav.: 74, 1856 [1857].

* subg. Diaphanae R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (1): 103, 1956.

*** Cololejeunea amphibola B.M.Thiers, Beih. Nova Hedwigia 90: 130, 1988.

** Cololejeunea antillana Pócs, Polish Bot. J. 54 (1): 3, 2009. Nom. nov. pro Aphanolejeunea longifolia Jovet-Ast, Rev. Bryol. Lichénol. 16 (1/2): 23, 1947 [1948].

* Cololejeunea augieri Tixier, Trop. Bryol. 11: 48, 1995.

*** Cololejeunea azorica V.Allorge et Jovet-Ast, Mitt. Thüring. Bot. Ges. 1 (2/3): 17, 1955.

*** Cololejeunea camillii (Lehm.) A.Evans, Bryologist 15 (4): 59, 1912. Bas.: Lejeunea camillii Lehm., Nov. Stirp. Pug. 10: 15, 1857.

*** Cololejeunea contractiloba A.Evans, Amer. J. Bot. 5 (3): 131, 1918.

*** Cololejeunea cornuta E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 436, 1954.

** Cololejeunea crenata (A.Evans) Pócs, Polish Bot. J. 54 (1): 5, 2009. Bas.: Aphanolejeunea crenata A.Evans, Bull. Torrey Bot. Club 38 (6): 276, 1911.

*** Cololejeunea diaphana A.Evans, Bull. Torrey Bot. Club 32 (4): 184, 1905.

*** Cololejeunea erostrata (Herzog) Bernecker et Pócs, Polish Bot. J. 54 (1): 5, 2009. Bas.: Physocolea erostrata Herzog, Beih. Bot. Centralbl. 61B (3): 582, 1942.

** Cololejeunea gradsteinii M.J.Lai et R.L.Zhu, Ann. Bot. Fenn. 45 (5): 334, 2008. Nom. nov. pro Cololejeunea pusilla Tixier, Nat. Hist. Bull. Siam Soc. 23 (4): 552, 1970, nom. illeg.

** Cololejeunea koponenii (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009. Bas.: Aphanolejeunea koponenii Pócs, Acta Bot. Fenn. 165: 91, 1999.

*** Cololejeunea lanceolata E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 428, 1954.

*** Cololejeunea moralesiae (Bern.-Lück.) Bernecker et Pócs, Polish Bot. J. 54 (1): 7, 2009. Bas.: Aphanolejeunea moralesiae Bern.-Lück., Nova Hedwigia 66 (1/2): 166, 1998.

*** Cololejeunea morobensis (Pócs) Pócs, Polish Bot. J. 54 (1): 7, 2009. Bas.: Aphanolejeunea morobensis Pócs, Ann. Bot. Fenn. 31 (3): 180, 1994.

*** Cololejeunea obtusifolia (E.W.Jones) Tixier, Trop. Bryol. 11: 42, 1995. Bas.: Cololejeunea pusilla var. obtusifolia E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 427, 1954.

* Cololejeunea obtusifolia var. madecassa (Tixier) Pócs, Phytotaxa 202 (1): 64, 2015. Bas.: Cololejeunea androphylla var. madecassa Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 216, 1977 [1979].

*** Cololejeunea papilliloba (Steph.) Steph., Hedwigia 34 (5): 250, 1895. Bas.: Lejeunea papilliloba Steph., Hedwigia 29 (2): 73, 1890.

** Cololejeunea papulosa R.M.Schust., Phytologia 45 (5): 433, 1980.

*** Cololejeunea paucifolia (Spruce) Bernecker et Pócs, Polish Bot. J. 54 (1): 8, 2009. Bas.: Lejeunea paucifolia Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxciv, 1889 [1890].

*** Cololejeunea peponiformis Mizut., J. Hattori Bot. Lab. 33: 258, 1970.

** Cololejeunea pseudocuspidata Tixier, Bauhinia 8 (4): 228, 1987.

*** Cololejeunea pterocolea Herzog, Svensk Bot. Tidskr. 46 (1): 103, 1952.

*** Cololejeunea pusilla Steph., Hedwigia 34 (5): 251, 1895.

** Cololejeunea spathulata Jovet-Ast, Rev. Bryol. Lichénol. 29 (1/2): 36, 1960.

** Cololejeunea taurifolia Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 152, 1965.

*** Cololejeunea wightii Steph., Hedwigia 34 (5): 253, 1895.

*** Cololejeunea yelitzae Pócs et Bernecker, Acta Bot. Hung. 55 (3/4): 386, 2013.

* subg. Leptocolea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 122, 1893. Bas.: Lejeunea sect. Leptocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 294, 1884.

** Cololejeunea acuminata Mizut., J. Hattori Bot. Lab. 33: 261, 1970.

*** Cololejeunea aequabilis (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 242, 1898. Bas.: Lejeunea aequabilis Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 310, 1864.

** Cololejeunea altimontana Pócs, Acta Bryolichenol. Asiat. 4: 78, 2011.

*** Cololejeunea amaniensis Pócs, Acta Bot. Hung. 31 (1/4): 120, 1985.

*** Cololejeunea angustibracteata Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894. Based on: Cololejeunea angustibracteata Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 34, 1890, nom. inval.

*** Cololejeunea apiculata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 174, 1963. Bas.: Leptocolea apiculata E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 415, 1954.

* Cololejeunea arrectifolia (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896. Bas.: Lejeunea arrectifolia Mitt., Fl. vit.: 415, 1871 [1873].

** Cololejeunea aurantia (Tixier) Thouvenot, Cryptog. Bryol. 32 (4): 290, 2011. Bas.: Jovetastella aurantia Tixier, Cryptog. Bryol. Lichénol. 3 (1): 29, 1982.

** Cololejeunea bandamiae Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 220, 1977 [1979].

** Cololejeunea bebourensis Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 229, 1977 [1979].

** Cololejeunea bergmansiana Tixier, Bull. Jard. Bot. Natl. Belg. 59 (3/4): 440, 1989.

** Cololejeunea bidentula (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 423, 1954. Bas.: Physocolea bidentula Steph., Sp. Hepat. (Stephani) 5: 868, 1916.

** Cololejeunea bifalcata Pócs, Acta Bot. Hung. 54 (1/2): 146, 2012.

*** Cololejeunea blepharophylla Pócs, Acta Bot. Hung. 54 (1/2): 149, 2012.

** Cololejeunea borhidiana Pócs, J. Hattori Bot. Lab. 48: 305, 1980.

** Cololejeunea bosseriana Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 235, 1977 [1979].

** Cololejeunea calcarata E.W.Jones, Bull. Brit. Mus. (Nat. Hist.), Bot. 11 (3): 235, 1983.

* Cololejeunea camusii Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 227, 1977 [1979].

*** Cololejeunea ceatocarpa (Ångstr.) Steph., Bull. Herb. Boissier 5 (10): 842, 1897. Bas.: Lejeunea ceatocarpa Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 27, 1872.

*** Cololejeunea ceylanica Onr., Acta Bot. Acad. Sci. Hung. 25 (1/2): 107, 1979.

* Cololejeunea chamlongiana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 181, 1970. 256

*** Cololejeunea chenii Tixier, Bryophyt. Biblioth. 27: 219, 1985. Nom. nov. pro Cololejeunea plagiophylla var. grossipapillosa P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 254, 1964.

*** Cololejeunea ciliata Pócs, Ann. Bot. Fenn. 31 (3): 182, 1994.

** Cololejeunea comptonii (Pearson) H.A.Mill., Phytologia 47 (4): 321, 1981. Bas.: Leptocolea comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 40, 1922.

** Cololejeunea cookei A.Evans, Trans. Connecticut Acad. Arts 10 (8): 447, 1900.

** Cololejeunea cordiflora Steph., Hedwigia 34 (5): 246, 1895.

** Cololejeunea crateris Pócs, Acta Bot. Hung. 54 (1/2): 153, 2012.

* Cololejeunea crenulata (Pearson) H.A.Mill., Phytologia 47 (4): 321, 1981. Bas.: Leptocolea crenulata Pearson, J. Linn. Soc., Bot. 46 (305): 41, 1922. 257

** Cololejeunea cucullifolia (Herzog) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 127, 1962. Bas.: Physocolea cucullifolia Herzog, Trans. & Proc. Roy. Soc. New Zealand 68 (1): 45, 1938.

** Cololejeunea cuneifolia Steph., Hedwigia 31 (4): 166, 1892.

* Cololejeunea dadeuniana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 186, 1970.

** Cololejeunea decemplicata (Steph.) Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 213, 1977 [1979]. Bas.: Physocolea decemplicata Steph., Sp. Hepat. (Stephani) 5: 869, 1916.

** Cololejeunea decliviloba Steph., Hedwigia 34 (5): 247, 1895.

** Cololejeunea dentata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 175, 1963. Bas.: Leptocolea dentata E.W.Jones, Trans. Brit. Bryol. Soc. 2 (2): 161, 1953.

** Cololejeunea denticulata (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 99, 1944. Bas.: Physocolea denticulata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 287, 1934.

* Cololejeunea dentilobula (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 241, 1963. Bas.: Physocolea dentilobula Steph., Sp. Hepat. (Stephani) 5: 892, 1916. 258

*** Cololejeunea dianae M.Wigginton, J. Bryol. 28 (4): 364, 2006.

** Cololejeunea dilatata (Steph.) Mizut., J. Hattori Bot. Lab. 28: 113, 1965. Bas.: Leptocolea dilatata Steph., Sp. Hepat. (Stephani) 5: 850, 1916.

*** Cololejeunea diplasiolejeunoides Tixier, Bryologist 82 (4): 602, 1979.

*** Cololejeunea distalopapillata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963. Bas.: Leptocolea distalopapillata E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 202, 1957.

* Cololejeunea effusa (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896. Bas.: Lejeunea effusa Mitt., Fl. vit.: 415, 1871 [1873]. 259

*** Cololejeunea elephantorum Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 185, 1970.

** Cololejeunea ellipsoidea R.M.Schust., Nova Hedwigia 15: 508, 1968.

*** Cololejeunea equialbi Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 178, 1970.

*** Cololejeunea filicis (Herzog) Piippo, J. Hattori Bot. Lab. 68: 133, 1990. Bas.: Leptocolea filicis Herzog, Symb. Sin. 5: 52, 1930.

* Cololejeunea fischeri Tixier, Trop. Bryol. 11: 59, 1995. 260

** Cololejeunea flavicans (Steph.) Mizut., J. Hattori Bot. Lab. 28: 115, 1965. Bas.: Physocolea flavicans Steph., Sp. Hepat. (Stephani) 5: 893, 1916.

* Cololejeunea fredericii Onr., Rev. Bryol. Lichénol. 44 (1): 80, 1978. 261

** Cololejeunea fusca (Steph.) Mizut., J. Hattori Bot. Lab. 28: 120, 1965. Bas.: Physocolea fusca Steph., Sp. Hepat. (Stephani) 5: 893, 1916.

** Cololejeunea gottschei (Steph.) Pandé, K.P.Srivast. et Ahmad, J. Indian Bot. Soc. 36 (3): 345, 1957. Bas.: Physocolea gottschei Steph., Sp. Hepat. (Stephani) 5: 894, 1916.

*** Cololejeunea grossestyla M.Wigginton, J. Bryol. 28 (4): 369, 2006.

*** Cololejeunea grushvitzkiana Pócs, Bot. Zhurn. (Moscow & Leningrad) 56 (5): 674, 1971.

*** Cololejeunea hainanensis R.L.Zhu, J. Hattori Bot. Lab. 78: 87, 1995.

*** Cololejeunea harrisii Pócs, Acta Bot. Acad. Sci. Hung. 21 (3/4): 357, 1975.

*** Cololejeunea hildebrandii (Austin) Steph., Bull. Herb. Boissier 5 (10): 842, 1897. Bas.: Lejeunea hildebrandii Austin, Bot. Bull. (Hanover) 1 (8): 35, 1876.

*** Cololejeunea hirta Steph., Bull. Misc. Inform. Kew 1899 (151/152): 125, 1899.

** Cololejeunea hodgsoniae (Herzog) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (11): 184, 1967. Bas.: Physocolea hodgsoniae Herzog, Trans. & Proc. Roy. Soc. New Zealand 68 (1): 46, 1938.

*** Cololejeunea horikawana (S.Hatt.) Mizut., J. Hattori Bot. Lab. 24: 254, 1961. Bas.: Leptocolea horikawana S.Hatt., J. Jap. Bot. 18 (11): 653, 1942.

*** Cololejeunea huerlimannii Tixier, Nova Hedwigia 31: 773, 1979.

** Cololejeunea inflexifolia R.M.Schust., Phytologia 56 (7): 458, 1985.

*** Cololejeunea iradieri M.Infante et Heras, Trop. Bryol. 17: 14, 1999.

** Cololejeunea irianensis Tixier, Bull. Jard. Bot. Natl. Belg. 59 (3/4): 440, 1989.

*** Cololejeunea johannis-winkleri (Herzog) R.L.Zhu, Nova Hedwigia 79 (3/4): 528, 2004. Bas.: Leptocolea johannis-winkleri Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 214, 1931.

** Cololejeunea kegelii Steph., Hedwigia 34 (5): 249, 1895.

** Cololejeunea khanii Tixier, Dacca Univ. Stud., B 15: 9, 1967.

* Cololejeunea kohkongensis Tixier, Bryophyt. Biblioth. 27: 302, 1985. 262

*** Cololejeunea lichenyae R.D.Porley, N.G.Hodgetts et M.Wigginton, J. Bryol. 29 (1): 7, 2007.

* Cololejeunea lobulilineata Tixier, Trop. Bryol. 11: 42, 1995. 263

*** Cololejeunea longifolia (Mitt.) Benedix ex Mizut., J. Hattori Bot. Lab. 26: 184, 1963. Bas.: Lejeunea longifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 117, 1860 [1861].

*** Cololejeunea magna (Tixier) M.Infante et Heras, Trop. Bryol. 17: 17, 1999. Bas.: Cololejeunea harrisii var. magna Tixier, Trop. Bryol. 11: 56, 1995.

** Cololejeunea magnifica Pócs, Acta Bryolichenol. Asiat. 4: 88, 2011.

** Cololejeunea magnilobula (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 99, 1944. Bas.: Physocolea magnilobula Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 288, 1934.

* Cololejeunea micronesica H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 63, 1963. 264

** Cololejeunea mooreaensis Tixier, Bauhinia 8 (4): 226, 1987.

* Cololejeunea ninguana Tixier, Nova Hedwigia 31: 776, 1979. 265

** Cololejeunea obcordata (Austin) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 448, 1900. Bas.: Lejeunea obcordata Austin, Bot. Bull. (Hanover) 1 (8): 36, 1876.

*** Cololejeunea obliqua (Nees et Mont.) Schiffn., Bot. Jahrb. Syst. 23 (5): 586, 1897. Bas.: Lejeunea obliqua Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 264, 1843.

* Cololejeunea oblongiperianthia (P.C.Wu et J.S.Lou) Piippo, J. Hattori Bot. Lab. 68: 134, 1990. Bas.: Leptocolea oblongiperianthia P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 109, 1978. 266

** Cololejeunea oleana Sim, Trans. Roy. Soc. South Africa 15 (1): 49, 1926.

** Cololejeunea ovalifolia A.Evans, Trans. Connecticut Acad. Arts 10 (8): 450, 1900.

** Cololejeunea panchoana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 183, 1970.

*** Cololejeunea paniensis (Tixier) Grolle, J. Bryol. 8 (4): 485, 1975. Bas.: Jovetastella paniensis Tixier, Rev. Bryol. Lichénol. 39 (4): 662, 1973 [1974].

** Cololejeunea papuliflora Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 85: 199, 1910.

** Cololejeunea pentagona (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896. Bas.: Lejeunea pentagona Mitt., Fl. vit.: 416, 1871 [1873].

*** Cololejeunea plagiochiliana Tixier, Bot. Not. 128: 428, 1975 [1976].

** Cololejeunea planiuscula Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995. Based on: Cololejeunea planiuscula Tixier, Candollea 46 (2): 289, 1991, nom. inval.

*** Cololejeunea pseudoserrata Tixier, Nova Hedwigia 31: 770, 1979.

* Cololejeunea pteroporum Tixier, Bryophyt. Biblioth. 27: 271, 1985. 267

** Cololejeunea pulchella (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 241, 1963. Bas.: Lejeunea pulchella Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 157, 1854.

** Cololejeunea pulchella var. stylifera R.M.Schust., Phytologia 56 (7): 458, 1985.

*** Cololejeunea quadridentata (S.Hatt.) Grolle, Acta Bot. Fenn. 125: 65, 1984. Bas.: Leptocolea quadridentata S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 117, 1951.

** Cololejeunea retusula (Mitt.) H.A.Mill., Phytologia 47 (4): 322, 1981. Bas.: Lejeunea retusula Mitt., Fl. vit.: 416, 1871 [1873].

** Cololejeunea salgadoi Onr., Bull. Jard. Bot. Natl. Belg. 59 (3/4): 433, 1989.

*** Cololejeunea sanctae-helenae M.Wigginton, J. Bryol. 28 (4): 366, 2006.

** Cololejeunea serrulata Steph., Hedwigia 34 (5): 252, 1895.

*** Cololejeunea setiloba A.Evans, Bryologist 16 (4): 51, 1913.

*** Cololejeunea siangensis G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 57, 2003.

*** Cololejeunea skottsbergii Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 749, 1942.

*** Cololejeunea societatis Tixier, Bauhinia 8 (4): 230, 1987.

* Cololejeunea spathulifolia (Steph.) H.A.Mill., Phytologia 47 (4): 322, 1981. Bas.: Leptocolea spathulifolia Steph., Sp. Hepat. (Stephani) 5: 855, 1916. 268

** Cololejeunea spruceana Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995. Based on: Cololejeunea spruceana Tixier, Candollea 46 (2): 286, 1991, nom. inval.

** Cololejeunea stenophylla Herzog, Bot. Not. 100 (4): 330, 1947.

*** Cololejeunea streimannii Pócs, Acta Bryolichenol. Asiat. 4: 96, 2011.

** Cololejeunea streimannii subsp. solomonensis Pócs, Acta Bryolichenol. Asiat. 4: 96, 2011.

*** Cololejeunea subalpina Pócs, Acta Bryolichenol. Asiat. 4: 98, 2011.

** Cololejeunea subcristata A.Evans, Bryologist 20 (2): 24, 1917.

* Cololejeunea takamakae Tixier, Bryophyt. Biblioth. 27: 319, 1985. 269

*** Cololejeunea tanzaniae Pócs, J. Hattori Bot. Lab. 48: 312, 1980.

** Cololejeunea tenuiparietata Tixier, Trop. Bryol. 11: 56, 1995.

** Cololejeunea teurnoumensis Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 183, 1970.

*** Cololejeunea timoi Pócs, Acta Bryolichenol. Asiat. 4: 98, 2011.

** Cololejeunea touwii Pócs, Acta Bryolichenol. Asiat. 4: 101, 2011.

*** Cololejeunea tranninhiana Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 97, 1974.

*** Cololejeunea trichomanis (Gottsche) Besch., Rev. Bryol. 19 (1): 14, 1892. Bas.: Lejeunea trichomanis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 362, 1882.

* Cololejeunea tuksapiana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 186, 1970. 270

** Cololejeunea vulcania Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 181, 1970.

*** Cololejeunea yoshinagana (S.Hatt.) Mizut., J. Hattori Bot. Lab. 24: 250, 1961. Bas.: Leptocolea yoshinagana S.Hatt., Bull. Tokyo Sci. Mus. 11: 115, 1944.

* subg. Metzgeriopsis (K.I.Goebel) Pócs, Acta Bryolichenol. Asiat. 4: 106, 2011. Bas.: Metzgeriopsis K.I.Goebel, Ann. Jard. Bot. Buitenzorg 7 (1): 54, 1888.

*** Cololejeunea metzgeriopsis (K.I.Goebel) Gradst., R.Wilson, Ilk.-Borg. et Heinrichs, Bot. J. Linn. Soc. 151 (3): 306, 2006. Bas.: Lejeunea metzgeriopsis K.I.Goebel, Flora 72 (1): 2, 1889.

*** subg. Pedinolejeunea Benedix ex Mizut., J. Hattori Bot. Lab. 24: 240, 1961.

** Cololejeunea abnormis Mizut., J. Hattori Bot. Lab. 33: 260, 1970.

*** Cololejeunea adhaesiva (Mitt.) R.M.Schust., Beih. Nova Hedwigia 9: 177, 1963. Bas.: Lejeunea adhaesiva Mitt., J. Linn. Soc., Bot. 22 (146): 325, 1886.

** Cololejeunea adnata Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 194, 1977 [1979].

*** Cololejeunea africana (Steph.) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963. Bas.: Physocolea africana Steph., Sp. Hepat. (Stephani) 5: 867, 1916.

** Cololejeunea ambeliensis Tixier, Bryophyt. Biblioth. 27: 142, 1985.

** Cololejeunea amieuensis Tixier, Nova Hedwigia 31: 748, 1979.

** Cololejeunea andapania Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 178, 1977 [1979].

*** Cololejeunea angulata (Steph.) Mizut., J. Hattori Bot. Lab. 28: 108, 1965. Bas.: Leptocolea angulata Steph., Sp. Hepat. (Stephani) 5: 847, 1916.

*** Cololejeunea ankaiana Tixier, Bryophyt. Biblioth. 27: 62, 1985.

** Cololejeunea attilana Pócs, Magyar Bot. Kut. Ezredf. Tanul. Borhidi: 186, 2002.

*** Cololejeunea auriculata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 177, 1963. Bas.: Leptocolea auriculata E.W.Jones, Trans. Brit. Bryol. Soc. 2 (2): 152, 1953.

** Cololejeunea autoica (Steph.) Grolle, Bryophyt. Biblioth. 48: 43, 1995. Bas.: Physocolea autoica Steph., Sp. Hepat. (Stephani) 5: 867, 1916.

*** Cololejeunea bekkeri Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995. Based on: Cololejeunea bekkeri Tixier, Candollea 46 (2): 269, 1991, nom. inval.

*** Cololejeunea bischleriana Tixier, Bradea 3 (6): 36, 1980.

** Cololejeunea bolovenensis Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 442, 1973.

*** Cololejeunea borbonica Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 188, 1977 [1979].

* Cololejeunea brunelii Tixier, Dacca Univ. Stud., B 15: 10, 1967. 271

*** Cololejeunea cardiocarpa (Mont.) A.Evans, Mem. Torrey Bot. Club 8 (2): 172, 1902. Bas.: Lejeunea cardiocarpa Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 476, 1842.

* Cololejeunea chittagongensis Tixier, Bryophyt. Biblioth. 27: 97, 1985. 272

*** Cololejeunea cocoscola Tixier, Cryptog. Bryol. Lichénol. 14 (3): 353, 1993.

** Cololejeunea cremersii Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995. Based on: Cololejeunea cremersii Tixier, Candollea 46 (2): 271, 1991, nom. inval.

** Cololejeunea cristata (Steph.) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963. Bas.: Physocolea cristata Steph., Sp. Hepat. (Stephani) 5: 869, 1916.

** Cololejeunea cuneata (Lehm. et Lindenb.) Herzog, Bot. Not. 100 (4): 320, 1947. Bas.: Jungermannia cuneata Lehm. et Lindenb., Nov. Stirp. Pug. 4: 56, 1832.

* Cololejeunea deroinii Tixier, Cryptog. Bryol. Lichénol. 14 (3): 355, 1993.

* Cololejeunea deslooveri Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 227, 1977. 273

* Cololejeunea dzumacensis Tixier, Nova Hedwigia 31: 754, 1979. 274

*** Cololejeunea ecuadoriensis Pócs, Acta Bot. Hung. 44 (3/4): 372, 2002.

* Cololejeunea epiphylla G.Asthana et A.Shukla, Cryptog. Bryol. 31 (3): 218, 2010. 275

** Cololejeunea fissilobula Herzog, Trans. Brit. Bryol. Soc. 1 (4): 323, 1950.

** Cololejeunea florencei Tixier, Cryptog. Bryol. Lichénol. 14 (3): 355, 1993.

** Cololejeunea foliicola S.C.Srivast. et G.Srivast., Proc. Indian Acad. Sci. Pl. Sci. 99 (2): 86, 1989.

** Cololejeunea fructumarginata Tixier, Bryophyt. Biblioth. 27: 58, 1985.

*** Cololejeunea furcilobulata (Berrie et E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 178, 1963. Bas.: Leptocolea furcilobulata Berrie et E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 417, 1954.

*** Cololejeunea geissleriana Tixier, Bradea 3 (6): 37, 1980.

** Cololejeunea georgiana Tixier, Bryophyt. Biblioth. 27: 145, 1985.

*** Cololejeunea guadelupensis Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995. Based on: Cololejeunea guadelupensis Tixier, Candollea 46 (2): 276, 1991, nom. inval.

*** Cololejeunea hebridensis Tixier, Bot. Not. 128: 425, 1975 [1976].

*** Cololejeunea hinidumae Onr., Acta Bot. Acad. Sci. Hung. 25 (1/2): 109, 1979.

* Cololejeunea hoabinhiana Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 91, 1974. 276

** Cololejeunea hoeana Tixier, Bryophyt. Biblioth. 27: 56, 1985.

** Cololejeunea hungii Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 91, 1974.

*** Cololejeunea indosinica Tixier, Bryophyt. Biblioth. 27: 63, 1985.

** Cololejeunea inoueana Mizut., J. Hattori Bot. Lab. 57: 440, 1984.

** Cololejeunea japonica (Schiffn.) Mizut., J. Hattori Bot. Lab. 24: 241, 1961. Bas.: Leptocolea japonica Schiffn., Ann. Bryol. 2: 92, 1929.

*** Cololejeunea jonesii Pócs, Acta Bot. Acad. Sci. Hung. 21 (3/4): 361, 1975.

*** Cololejeunea kapingaensis H.A.Mill., Bryologist 59 (3): 170, 1956.

** Cololejeunea kiriromensis Tixier, Bryophyt. Biblioth. 27: 147, 1985.

*** Cololejeunea kulenensis Tixier, Bryophyt. Biblioth. 27: 71, 1985. Nom. nov. pro Leptocolea verdoornii Herzog, Ann. Bryol. 5: 97, 1932.

** Cololejeunea laevigata (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 241, 1963. Bas.: Lejeunea laevigata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 157, 1854.

*** Cololejeunea lanciloba Steph., Hedwigia 34 (5): 250, 1895.

*** Cololejeunea latilobula (Herzog) Tixier, Bryophyt. Biblioth. 27: 156, 1985. Bas.: Leptocolea latilobula Herzog, Symb. Sin. 5: 54, 1930.

*** Cololejeunea latistyla R.L.Zhu, Hikobia 11: 544, 1994.

*** Cololejeunea leloutrei (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963. Bas.: Leptocolea leloutrei E.W.Jones, Trans. Brit. Bryol. Soc. 2 (2): 146, 1953.

** Cololejeunea leloutrei var. microlobulata Tixier, Bryophyt. Biblioth. 27: 78, 1985.

** Cololejeunea leloutrei var. ulugurica Pócs ex Tixier, Bryophyt. Biblioth. 27: 74, 1985.

*** Cololejeunea lemuriana Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 191, 1977 [1979].

** Cololejeunea littoralis Tixier, Bryophyt. Biblioth. 27: 163, 1985.

** Cololejeunea longistylis A.Evans, Trans. Connecticut Acad. Arts 10 (8): 453, 1900.

*** Cololejeunea magnistyla (Horik.) Mizut., J. Hattori Bot. Lab. 24: 243, 1961. Bas.: Leptocolea magnistyla Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 131, 1932.

* Cololejeunea malaccensis Tixier, Bryophyt. Biblioth. 27: 42, 1985. 277

** Cololejeunea malayana Tixier, Bryophyt. Biblioth. 27: 154, 1985.

*** Cololejeunea marginata (Lehm. et Lindenb.) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892. Bas.: Jungermannia marginata Lehm. et Lindenb., Nov. Stirp. Pug. 5: 11, 1833.

* Cololejeunea maritima Tixier, Nova Hedwigia 31: 752, 1979.

*** Cololejeunea minutilobula Herzog, Rev. Bryol. Lichénol. 20 (1/2): 171, 1951 [1952].

*** Cololejeunea nigerica (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 177, 1963. Bas.: Leptocolea nigerica E.W.Jones, Trans. Brit. Bryol. Soc. 2 (2): 150, 1953.

** Cololejeunea occidentalis (E.W.Jones) Vanden Berghen, Rev. Bryol. Lichénol. 44 (4): 449, 1978. Bas.: Leptocolea cristata var. occidentalis E.W.Jones, Trans. Brit. Bryol. Soc. 2 (2): 149, 1953.

** Cololejeunea onraedtii Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 202, 1977 [1979].

*** Cololejeunea pacifica Pócs, Acta Bot. Hung. 54 (1/2): 158, 2012.

** Cololejeunea panamensis G.Dauphin et Pócs, Trop. Bryol. 27: 76, 2006.

*** Cololejeunea paucimarginata Tixier, Bryophyt. Biblioth. 27: 100, 1985.

** Cololejeunea perakensis Tixier, Bryophyt. Biblioth. 27: 95, 1985.

*** Cololejeunea plagiophylla Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 73, 1953.

*** Cololejeunea planissima (Mitt.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 73, 1959. Bas.: Lejeunea planissima Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 117, 1860 [1861].

** Cololejeunea planissima var. chagosensis Pócs, J. Bryol. 28 (1): 14, 2006.

** Cololejeunea praeruptorum Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 205, 1977 [1979].

** Cololejeunea producta (Mitt.) S.Hatt., Fl. E. Himalaya: 533, 1966. Bas.: Lejeunea producta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 117, 1860 [1861].

* Cololejeunea punctata (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 9, 1893. Bas.: Lejeunea punctata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 361, 1882. 278

*** Cololejeunea raduliloba Steph., Hedwigia 34 (5): 251, 1895.

* Cololejeunea reineckeana Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896.

** Cololejeunea saltuum Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 188, 1977 [1979].

** Cololejeunea sambiroana Tixier, Bryologist 82 (4): 608, 1979.

*** Cololejeunea saroltae Pócs, Acta Bot. Hung. 54 (1/2): 160, 2012.

*** Cololejeunea schusteri Pócs, Acta Bot. Hung. 44 (3/4): 376, 2002.

** Cololejeunea schwabei Herzog, J. Hattori Bot. Lab. 14: 54, 1955.

*** Cololejeunea selangorensis Tixier, Bryophyt. Biblioth. 27: 166, 1985.

*** Cololejeunea shibiensis Mizut., J. Hattori Bot. Lab. 57: 437, 1984.

*** Cololejeunea smitinandii Tixier, Bryophyt. Biblioth. 27: 131, 1985. Based on: Cololejeunea smitinandii Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 439, 1973, nom. inval.

** Cololejeunea stoniana Tixier, Bot. Not. 128: 429, 1975 [1976].

*** Cololejeunea stylosa Steph., Trans. Connecticut Acad. Arts 10 (8): 454, 1900. Based on: Cololejeunea stylosa Steph., Hedwigia 27 (11/12): 289, 1888, nom. inval.

*** Cololejeunea subcardiocarpa Tixier, Bradea 3 (6): 39, 1980.

** Cololejeunea subinflata Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 182, 1977 [1979].

*** Cololejeunea submarginata Tixier, Bradea 3 (6): 40, 1980.

** Cololejeunea subminutilobula Mizut., J. Hattori Bot. Lab. 24: 282, 1961. Nom. nov. pro Leptocolea minutilobula Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 16, 1931.

** Cololejeunea subscariosa (Spruce) Pócs, Acta Bot. Hung. 56 (1/2): 197, 2014. Bas.: Lejeunea subscariosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 300, 1884.

** Cololejeunea subtriapiculata Tixier, Nova Hedwigia 31: 744, 1979.

*** Cololejeunea succinea Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 194, 1977 [1979].

*** Cololejeunea surinamensis Tixier, Bradea 3 (6): 42, 1980.

** Cololejeunea tahitensis Tixier, Cryptog. Bryol. Lichénol. 14 (3): 359, 1993.

** Cololejeunea tamatavensis Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 191, 1977 [1979].

* Cololejeunea taprobanea Tixier, Bryophyt. Biblioth. 27: 158, 1985.

* Cololejeunea thailandensis Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 441, 1973.

** Cololejeunea triapiculata (Herzog) Tixier, Gard. Bull. Singapore 25 (3): 344, 1971. Bas.: Leptocolea triapiculata Herzog, Ann. Bryol. 5: 95, 1932.

* Cololejeunea tribracteata Tixier, Trop. Bryol. 11: 46, 1995. 279

* Cololejeunea tridentata Tixier, Bryophyt. Biblioth. 27: 83, 1985.

* Cololejeunea uchimae Amakawa, J. Jap. Bot. 33 (5): 142, 1958. 280

*** Cololejeunea verwimpii Tixier, Cryptog. Bryol. Lichénol. 16 (3): 230, 1995.

*** Cololejeunea vidaliana Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 444, 1973.

** Cololejeunea vietnamensis Tixier, Bryophyt. Biblioth. 27: 127, 1985.

** Cololejeunea vitaliana Tixier, Cryptog. Bryol. Lichénol. 16 (3): 230, 1995.

*** Cololejeunea yakusimensis (S.Hatt.) Mizut., J. Hattori Bot. Lab. 57: 430, 1984. Bas.: Leptocolea lanciloba var. yakusimensis S.Hatt., J. Jap. Bot. 18 (11): 655, 1942.

* subg. Protocolea R.M.Schust., Beih. Nova Hedwigia 9: 171, 1963.

** Cololejeunea chuahiana Pócs, Polish Bot. J. 47 (1): 11, 2002. 281

** Cololejeunea dauphinii R.L.Zhu, J. Bryol. 28 (3): 277, 2006. Nom. nov. pro Cololejeunea tixieri M.I.Morales et G.Dauphin, Trop. Bryol. 14: 133, 1998, nom. illeg. 282

** Cololejeunea disciflora Tixier, Bryologist 82 (4): 604, 1979. 283

*** subg. Taeniolejeunea (Zwickel) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 21, 1953. Bas.: Taeniolejeunea Zwickel, Ann. Bryol. 6: 106, 1933.

* Cololejeunea amoena Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 25, 1953. 284

*** Cololejeunea appressa (A.Evans) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 31, 1953. Bas.: Leptocolea appressa A.Evans, Bull. Torrey Bot. Club 39 (12): 606, 1912 [1913].

* Cololejeunea bachmaensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 560, 1969 [1970].

* Cololejeunea bontocensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 583, 1969 [1970]. 285

* Cololejeunea crassipapillata Tixier, Rev. Bryol. Lichénol. 36 (3/4): 565, 1969 [1970].

** Cololejeunea eustacei Pócs, J. Bryol. 29 (2): 83, 2007.

*** Cololejeunea falcata (Horik.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 29, 1953. Bas.: Physocolea falcata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 22, 1931.

** Cololejeunea falcata var. madecassa Tixier, Bryologist 82 (4): 606, 1979.

* Cololejeunea flavida P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 103, 1978. 286

** Cololejeunea flavovittata Pócs, Acta Bryolichenol. Asiat. 4: 120, 2011.

*** Cololejeunea floccosa (Lehm. et Lindenb.) Schiffn., Consp. Hepat. Arch. Ind.: 243, 1898. Bas.: Jungermannia floccosa Lehm. et Lindenb., Nov. Stirp. Pug. 5: 26, 1833.

* Cololejeunea floccosa var. amoenoides Tixier, Cryptog. Bryol. Lichénol. 2 (1): 59, 1981.

** Cololejeunea floccosa var. angustibracteata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 65, 1981.

** Cololejeunea floccosa var. aurita Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 24, 1953.

** Cololejeunea floccosa var. conivens Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 24, 1953.

* Cololejeunea floccosa var. ocellata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 69, 1981.

** Cololejeunea floccosa var. plicata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 65, 1981.

** Cololejeunea floccosa var. trivittata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 62, 1981.

** Cololejeunea gresicola Tixier, Rev. Bryol. Lichénol. 36 (3/4): 569, 1969 [1970].

** Cololejeunea gynophthalma Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 32, 1953.

*** Cololejeunea inflata Steph., Hedwigia 34 (5): 249, 1895.

** Cololejeunea khiavensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 557, 1969 [1970].

** Cololejeunea koratensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 585, 1969 [1970].

* Cololejeunea manlinensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 563, 1969 [1970].

** Cololejeunea maquilingensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 579, 1969 [1970].

** Cololejeunea mutabilis Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 27, 1953.

** Cololejeunea nakajimae S.Hatt., J. Hattori Bot. Lab. 10: 57, 1953.

*** Cololejeunea ocellata (Horik.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 38, 1953. Bas.: Leptocolea ocellata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 86, 1932.

*** Cololejeunea ocelloides (Horik.) Mizut., J. Hattori Bot. Lab. 24: 277, 1961. Bas.: Leptocolea ocelloides Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 280, 1934.

*** Cololejeunea peraffinis (Schiffn.) Schiffn., Consp. Hepat. Arch. Ind.: 245, 1898. Bas.: Lejeunea peraffinis Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 242, 1893.

* Cololejeunea peraffinis var. ciconiae Tixier, Bull. Jard. Bot. Natl. Belg. 59 (3/4): 444, 1989.

** Cololejeunea peraffinis var. elegans Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 35, 1953.

** Cololejeunea peraffinis var. serrulata Schiffn. ex Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 35, 1953.

* Cololejeunea polisiana Tixier, Rev. Bryol. Lichénol. 36 (3/4): 550, 1969 [1970]. 287

*** Cololejeunea pseudofloccosa (Horik.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 36, 1953. Bas.: Leptocolea pseudofloccosa Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 87, 1932.

*** Cololejeunea pseudostephanii Tixier, Rev. Bryol. Lichénol. 36 (3/4): 571, 1969 [1970].

** Cololejeunea setosa Mizut., J. Hattori Bot. Lab. 29: 163, 1966.

*** Cololejeunea sharpii Mizut., J. Hattori Bot. Lab. 39: 258, 1975.

*** Cololejeunea siamensis Steph., Bot. Tidsskr. 24 (3): 279, 1902.

*** Cololejeunea sphaerodonta Mizut., J. Hattori Bot. Lab. 29: 165, 1966.

*** Cololejeunea stephanii Schiffn. ex Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 40, 1953.

** Cololejeunea subfloccosa Mizut., J. Hattori Bot. Lab. 57: 168, 1984.

** Cololejeunea subocelloides Mizut., J. Hattori Bot. Lab. 57: 163, 1984.

* Cololejeunea tamdaoensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 567, 1969 [1970].

** Cololejeunea verdoornii (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 17: 75, 1956 [1957]. Bas.: Taeniolejeunea verdoornii S.Hatt., J. Jap. Bot. 17: 459, 1941.

** Cololejeunea yipii R.L.Zhu, Beih. Nova Hedwigia 121: 346, 2001.

** Cololejeunea zantenorum Pócs, Acta Bryolichenol. Asiat. 4: 127, 2011.

Incertae sedis

** Cololejeunea conchifolia (Gottsche) Gradst., J. Hattori Bot. Lab. 45: 109, 1979. Bas.: Lejeunea conchifolia Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 163, 1864. 288

** Cololejeunea dankiaensis Tixier, Phytotaxa 220 (2): 199, 2015. Based on: Cololejeunea dankiaensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 581, 1969 [1970], nom. inval.

** Cololejeunea ensifera Tixier, Phytotaxa 220 (2): 199, 2015. Based on: Cololejeunea ensifera Tixier, Rev. Bryol. Lichénol. 36 (3/4): 562, 1969 [1970], nom. inval.

** Cololejeunea hamata Steph., Hedwigia 34 (5): 248, 1895.

** Cololejeunea herzogii K.I.Goebel, Biblioth. Bot. 87 (2): 269, 1916.

** Cololejeunea jamesii (Austin) M.E.Reiner et Pócs, Phytotaxa 208 (1): 98, 2015. Bas.: Lejeunea jamesii Austin, Bull. Torrey Bot. Club 6 (30): 158, 1877.

** Cololejeunea sublatistyla Jian Wang bis et R.L.Zhu, Phytotaxa 161 (2): 165, 2014.

** Cololejeunea tixieri Onr., Bull. Jard. Bot. Natl. Belg. 59 (3/4): 436, 1989.

* Cololejeunea variifolia (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896. Bas.: Lejeunea variifolia Mitt., Fl. vit.: 415, 1871 [1873].

*** Colura (Dumort.) Dumort., Recueil Observ. Jungerm.: 12, 1835. Bas.: Lejeunea sect. Colura Dumort., Syll. Jungerm. Europ.: 32, 1831.

** subg. Colura

** sect. Colura

*** Colura berghenii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 245, 1953 [1954].

*** Colura calyptrifolia (Hook.) Dumort., Recueil Observ. Jungerm.: 12, 1835. Bas.: Jungermannia calyptrifolia Hook., Brit. Jungermann.: tab. 43, 1813.

*** Colura hedbergiana Pócs, J. Bryol. 14 (3): 499, 1987.

*** Colura humbertii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 251, 1953 [1954].

*** Colura irrorata (Spruce) Heinrichs, Y.Yu, Schäf.-Verw. et Pócs, Phytotaxa 66: 58, 2012. Bas.: Myriocolea irrorata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 305, 1884.

** Colura junghuhniana (Prantl) Steph., Sp. Hepat. (Stephani) 5: 938, 1916. Bas.: Lejeunea junghuhniana Prantl, Hedwigia 29: xvi, 1890.

** Colura medusa J.Eggers et Pócs, Chenia 11: 22, 2013.

** Colura mizutanii Pócs, Chenia 11: 22, 2013.

*** Colura rhynchophora Jovet-Ast, Rev. Bryol. Lichénol. 17 (1/4): 27, 1948 [1949].

*** Colura tenuicornis (A.Evans) Steph., Sp. Hepat. (Stephani) 5: 942, 1916, nom. conserv. Bas.: Colurolejeunea tenuicornis A.Evans, Trans. Connecticut Acad. Arts 10 (8): 455, 1900.

** sect. Gamolepis Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 207, 1983.

*** Colura cristata Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 291, 1953 [1954].

*** Colura greig-smithii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 293, 1953 [1954].

*** Colura inflata K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 11, 1928.

*** Colura jovet-astiae Grolle, J. Hattori Bot. Lab. 28: 44, 1965. Nom. nov. pro Colura undulata Jovet-Ast, Rev. Bryol. Lichénol. 30 (1/2): 7, 1961, nom. illeg.

*** Colura meijeri Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 290, 1953 [1954].

*** Colura obvoluta Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 207, 1983.

*** Colura ornata K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 26, 1890 [1891].

*** Colura palawanensis Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 305, 1953 [1954].

*** Colura valida Jovet-Ast, Rev. Bryol. Lichénol. 30 (1/2): 6, 1961.

*** Colura verdoornii Herzog et Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 288, 1953 [1954].

** sect. Harmophyllum Grolle, J. Hattori Bot. Lab. 28: 44, 1965.

*** Colura andoi Gradst. et Jovet-Ast, Hikobia 9: 355, 1986.

*** Colura ari (Steph.) Steph., Sp. Hepat. (Stephani) 5: 936, 1916. Bas.: Colurolejeunea ari Steph., Hedwigia 35 (3): 73, 1896.

*** Colura australiensis Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 260, 1953 [1954].

*** Colura brevistyla Herzog, Beih. Bot. Centralbl. 38 (2): 331, 1921.

*** Colura calderae Pócs, J. Bryol. 29 (2): 84, 2007.

*** Colura clementis Grolle, J. Hattori Bot. Lab. 28: 45, 1965.

*** Colura conica (Sande Lac.) K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 3, 1928. Bas.: Lejeunea conica Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 311, 1864.

*** Colura corynophora (Nees, Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 402, 1877. Bas.: Lejeunea corynophora Nees, Lindenb. et Gottsche, Observ. bot.: 474, 1843.

*** Colura crenulata Grolle, J. Hattori Bot. Lab. 28: 46, 1965.

*** Colura crispiloba Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 205, 1983.

** Colura cylindrica Herzog, Svensk Bot. Tidskr. 46 (1): 106, 1952.

*** Colura cymbalifera Herzog et Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 268, 1953 [1954].

*** Colura digitalis (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 931, 1916. Bas.: Lejeunea digitalis Mitt., J. Linn. Soc., Bot. 22 (146): 325, 1886.

** Colura digitalis var. mucronata Pócs, Fragm. Florist. Geobot. 40 (1): 263, 1995.

*** Colura dusenii Steph., Sp. Hepat. (Stephani) 5: 931, 1916. Based on: Colurolejeunea dusenii Steph., Hedwigia 31 (4): 168, 1892.

*** Colura fastigiata Jovet-Ast, Rev. Bryol. Lichénol. 27 (1/2): 28, 1958.

*** Colura fistulosa Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 211, 1983.

*** Colura hattoriana Pócs, J. Hattori Bot. Lab. 74: 47, 1993.

*** Colura heimii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 275, 1953 [1954].

*** Colura hemisphaerica Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 267, 1953 [1954].

*** Colura herzogii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 261, 1953 [1954].

*** Colura inuii Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 68, 1931.

** Colura koponenii Pócs, Chenia 11: 29, 2013.

** Colura mauritiana Pócs, Cryptog. Bryol. Lichénol. 18 (3): 196, 1997.

*** Colura maxima Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 284, 1953 [1954].

*** Colura mosenii Steph., Sp. Hepat. (Stephani) 5: 940, 1916.

** Colura norrisii Pócs, Chenia 11: 29, 2013.

*** Colura obesa Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 273, 1953 [1954].

*** Colura pallida Steph., Sp. Hepat. (Stephani) 5: 941, 1916.

*** Colura pluridentata Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 265, 1953 [1954].

*** Colura speciosa Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 307, 1953 [1954].

** Colura streimannii Pócs, Polish Bot. J. 60 (1): 7, 2015.

*** Colura superba (Mont.) Steph., Sp. Hepat. (Stephani) 5: 941, 1916. Bas.: Lejeunea superba Mont., Ann. Sci. Nat. Bot. (sér. 3) 10: 115, 1848.

*** Colura thomeensis Pócs, Bryologist 114 (2): 363, 2011.

*** Colura tortifolia (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 402, 1877. Bas.: Lejeunea tortifolia Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 265, 1843.

*** Colura tutuilana (Pearson) H.A.Mill., Phytologia 47 (4): 322, 1981. Bas.: Colurolejeunea tutuilana Pearson, Amer. Samoa: 151, 1924.

*** Colura ulei Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 270, 1953 [1954].

*** Colura vitiensis Pócs et J.Eggers, Polish Bot. J. 52 (2): 88, 2007.

** sect. Oidocorys Jovet-Ast ex Grolle, J. Hattori Bot. Lab. 32: 140, 1969.

*** Colura itatyana Steph., Sp. Hepat. (Stephani) 5: 932, 1916.

*** Colura naumannii (Schiffn.) Steph., Sp. Hepat. (Stephani) 5: 935, 1916. Bas.: Colurolejeunea naumannii Schiffn., Hepat. (Engl.-Prantl): 121, 1893.

*** Colura ornithocephala Herzog, Svensk Bot. Tidskr. 46 (1): 107, 1952.

*** Colura pulcherrima Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 235, 1953 [1954].

** Colura pulcherrima var. bartlettii Jovet-Ast, Cryptog. Bryol. Lichénol. 1 (3): 278, 1980.

*** Colura saccophylla E.A.Hodgs. et Herzog, Trans. & Proc. Roy. Soc. New Zealand 77 (2): 253, 1949.

*** Colura schusteri Grolle, J. Hattori Bot. Lab. 32: 140, 1969.

** subg. Glotta Grolle et R.L.Zhu, J. Hattori Bot. Lab. 92: 187, 2002.

** sect. Glotta Grolle et R.L.Zhu, J. Hattori Bot. Lab. 92: 187, 2002.

*** Colura bicornis Jovet-Ast, Rev. Bryol. Lichénol. 25 (3/4): 272, 1956 [1957].

*** Colura bisvoluta Herzog et Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 228, 1953 [1954].

*** Colura inornata Jovet-Ast, Rev. Bryol. Lichénol. 25 (3/4): 275, 1956 [1957].

*** Colura karstenii K.I.Goebel, Pflanzenbiol. Schilderungen 2 (1): 153, 1891.

*** Colura queenslandica B.M.Thiers, Brittonia 39 (2): 175, 1987.

*** Colura saroltae Pócs, J. Bryol. 14 (3): 497, 1987.

*** Colura strophiolata Jovet-Ast, Rev. Bryol. Lichénol. 42 (4): 917, 1976.

*** Colura usambarica E.W.Jones, J. Bryol. 14 (3): 495, 1987.

** sect. Heterophyllum Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 213, 1983.

*** Colura acroloba (Prantl) Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 297, 1953 [1954]. Bas.: Lejeunea acroloba Prantl, Hedwigia 29: xiv, 1890.

*** Colura corniantha Grolle, J. Hattori Bot. Lab. 28: 44, 1965. Nom. nov. pro Colura cornuta Jovet-Ast, Rev. Bryol. Lichénol. 30 (1/2): 11, 1961, nom. illeg.

** Colura denticulata Jovet-Ast, Rev. Bryol. Lichénol. 23 (1/2): 2, 1954.

*** Colura galeata Jovet-Ast, Rev. Bryol. Lichénol. 35 (1/4): 146, 1967 [1968].

** Colura imperfecta Steph., Sp. Hepat. (Stephani) 5: 938, 1916.

*** Colura siamensis Jovet-Ast, Rev. Bryol. Lichénol. 35 (1/4): 139, 1967 [1968].

*** Colura tixieri Jovet-Ast, Rev. Bryol. Lichénol. 30 (1/2): 9, 1961.

*** Colura vietnamensis Jovet-Ast et Tixier, Rev. Bryol. Lichénol. 27 (3/4): 205, 1958 [1959].

Incertae sedis

** Colura amboinensis Steph., Sp. Hepat. (Stephani) 5: 935, 1916.

*** Colura hirta Steph., Sp. Hepat. (Stephani) 5: 932, 1916.

*** Diplasiolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 121, 1893. Bas.: Lejeunea subg. Diplasiolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 301, 1884. 289

*** Diplasiolejeunea utriculata Steph., Sp. Hepat. (Stephani) 5: 920, 1916.

** subg. Austrolejeuneopsis R.M.Schust., Bull. Torrey Bot. Club 97 (6): 349, 1970.

*** Diplasiolejeunea alata Jovet-Ast, Rev. Bryol. Lichénol. 17 (1/4): 31, 1948 [1949].

*** Diplasiolejeunea eggersii Pócs, Bryologist 109 (3): 408, 2006.

*** Diplasiolejeunea erostrata Schäf.-Verw., Cryptog. Bryol. 25 (1): 3, 2004.

* Diplasiolejeunea guadalupensis Steph., Sp. Hepat. (Stephani) 5: 924, 1916. 290

* Diplasiolejeunea heimii Jovet-Ast, Rev. Bryol. Lichénol. 29 (1/2): 39, 1960. 291

** Diplasiolejeunea involuta S.Winkl., Rev. Bryol. Lichénol. 35 (1/4): 320, 1967 [1968].

** Diplasiolejeunea involuta subsp. andicola Pócs, Cryptog. Bryol. Lichénol. 19 (1): 12, 1998.

*** Diplasiolejeunea johnsonii A.Evans, Bull. Torrey Bot. Club 39 (12): 603, 1912 [1913].

** Diplasiolejeunea johnsonii var. mexicana Jovet-Ast, Rev. Bryol. Lichénol. 29 (1/2): 39, 1960.

* Diplasiolejeunea montecristensis S.Winkl., Rev. Bryol. Lichénol. 35 (1/4): 321, 1967 [1968]. 292

*** Diplasiolejeunea papilionacea R.M.Schust., Phytologia 39 (6): 431, 1978.

*** Diplasiolejeunea pauckertii (Nees) Steph., Sp. Hepat. (Stephani) 5: 924, 1916. Bas.: Lejeunea pauckertii Nees, Syn. Hepat. 3: 392, 1845.

*** Diplasiolejeunea pluridentata Schäf.-Verw., Haussknechtia 8: 71, 2001.

*** Diplasiolejeunea replicata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 926, 1916. Bas.: Lejeunea replicata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 302, 1884.

** Diplasiolejeunea rudolphiana Steph., Hedwigia 35 (3): 79, 1896. 293

*** Diplasiolejeunea runssorensis Steph., Bot. Jahrb. Syst. 20 (3): 318, 1895.

*** Diplasiolejeunea unidentata (Lehm. et Lindenb.) Schiffn., Bot. Jahrb. Syst. 23 (5): 583, 1897. Bas.: Jungermannia unidentata Lehm. et Lindenb., Nov. Stirp. Pug. 6: 48, 1834.

** subg. Diplasiolejeunea

*** Diplasiolejeunea armatiloba Steph., Hedwigia 35 (3): 80, 1896.

*** Diplasiolejeunea borhidiana Reyes Montoya, Acta Bot. Acad. Sci. Hung. 28 (1/2): 177, 1982 [1983].

*** Diplasiolejeunea brunnea Steph., Sp. Hepat. (Stephani) 5: 922, 1916.

** Diplasiolejeunea caribea Tixier, Bryophyt. Biblioth. 27: 377, 1985.

*** Diplasiolejeunea cavifolia Steph., Sp. Hepat. (Stephani) 5: 918, 1916. Nom. nov. pro Lejeunea cavifolia Steph., Bot. Jahrb. Syst. 8 (2): 89, 1886, nom. illeg.

*** Diplasiolejeunea cobrensis Steph., Sp. Hepat. (Stephani) 5: 923, 1916.

** Diplasiolejeunea cobrensis subsp. antsirananae Pócs, Candollea 56 (1): 72, 2001.

** Diplasiolejeunea cogoensis M.Infante, Heras et Pócs, Trop. Bryol. 17: 9, 1999.

** Diplasiolejeunea cubensis Tixier, Bryophyt. Biblioth. 27: 389, 1985.

* Diplasiolejeunea glaziovii Tixier, Bryophyt. Biblioth. 27: 397, 1985. 294

*** Diplasiolejeunea ingekarolae Schäf.-Verw., Herzogia 19: 239, 2006.

** Diplasiolejeunea jonesii Tixier, Acta Bot. Hung. 30 (1/2): 21, 1984.

*** Diplasiolejeunea leiocarpa Jovet-Ast, Rev. Bryol. Lichénol. 16 (1/2): 34, 1947 [1948].

*** Diplasiolejeunea malleiformis (A.Evans) Tixier, Bryophyt. Biblioth. 27: 351, 1985. Bas.: Diplasiolejeunea pellucida var. malleiformis A.Evans, Bull. Torrey Bot. Club 39 (5): 215, 1912.

** Diplasiolejeunea palustrium Tixier, Acta Bot. Hung. 30 (1/2): 19, 1984.

*** Diplasiolejeunea pellucida (C.F.W.Meissn. ex Spreng.) Schiffn., Hepat. (Engl.-Prantl): 121, 1893. Bas.: Jungermannia pellucida C.F.W.Meissn. ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827.

** Diplasiolejeunea phyllarthronii Tixier, Acta Bot. Hung. 30 (1/2): 19, 1984.

*** Diplasiolejeunea pocsii Reyes Montoya, Acta Bot. Acad. Sci. Hung. 28 (1/2): 173, 1982 [1983].

** subg. Physolejeunea R.M.Schust., Bull. Torrey Bot. Club 97 (6): 348, 1970.

*** Diplasiolejeunea andringitrae Schäf.-Verw., Cryptog. Bryol. 27 (4): 447, 2006.

*** Diplasiolejeunea cornuta Steph., Sp. Hepat. (Stephani) 5: 918, 1916.

*** Diplasiolejeunea jovet-astiae Grolle, Feddes Repert. 73 (2): 84, 1966.

*** Diplasiolejeunea kraussiana (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 919, 1916. Bas.: Lejeunea kraussiana Lindenb., Syn. Hepat. 3: 393, 1845.

*** Diplasiolejeunea ornata Pócs et Schäf.-Verw., Cryptog. Bryol. 27 (4): 440, 2006.

*** Diplasiolejeunea patelligera Herzog, Svensk Bot. Tidskr. 42 (3): 240, 1948.

*** Diplasiolejeunea plicatiloba (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 45: 175, 1979. Bas.: Jungermannia plicatiloba Hook.f. et Taylor, London J. Bot. 4: 92, 1845.

*** Diplasiolejeunea pusilla Grolle, Feddes Repert. 86 (1/2): 79, 1975.

*** Diplasiolejeunea ranomafanae Pócs, Cryptog. Bryol. 27 (4): 444, 2006.

*** Diplasiolejeunea villaumei Steph., Sp. Hepat. (Stephani) 5: 921, 1916.

*** Diplasiolejeunea zakiae Tixier, Lindbergia 4 (1/2): 123, 1977.

Incertae sedis

*** Diplasiolejeunea albifolia (Taylor) E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 393, 1954. Bas.: Lejeunea albifolia Taylor, London J. Bot. 5: 399, 1846.

*** Diplasiolejeunea aulae E.W.Jones, J. Bryol. 7 (4): 552, 1973 [1974].

** Diplasiolejeunea auriculata Tixier, Rev. Bryol. Lichénol. 45 (2): 210, 1979.

*** Diplasiolejeunea buckii Grolle, Beitr. Phytotax. 15: 105, 1992.

** Diplasiolejeunea columbica Tixier, Cryptog. Bryol. Lichénol. 4 (3): 233, 1983.

* Diplasiolejeunea comorensis Tixier, Rev. Bryol. Lichénol. 45 (2): 213, 1979.

** Diplasiolejeunea cyanguguensis Tixier, Trop. Bryol. 11: 67, 1995.

*** Diplasiolejeunea deslooveri Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 217, 1977.

** Diplasiolejeunea ensifera Tixier, Rev. Bryol. Lichénol. 45 (2): 215, 1979.

* Diplasiolejeunea evansii Tixier, Bryophyt. Biblioth. 27: 360, 1985. 295

** Diplasiolejeunea gradsteinii Tixier, Trop. Bryol. 11: 67, 1995.

*** Diplasiolejeunea grandirostrata Schäf.-Verw., Cryptog. Bryol. 25 (1): 7, 2004.

*** Diplasiolejeunea grolleana Reyes Montoya, Acta Bot. Acad. Sci. Hung. 28 (1/2): 175, 1982 [1983].

** Diplasiolejeunea hamata Tixier, Rev. Bryol. Lichénol. 45 (2): 217, 1979.

** Diplasiolejeunea insignis Tixier, Lindbergia 4 (1/2): 120, 1977.

* Diplasiolejeunea integerrima Tixier, Rev. Bryol. Lichénol. 45 (2): 219, 1979. 296

*** Diplasiolejeunea lanceolata Grolle, Beitr. Phytotax. 15: 107, 1992.

*** Diplasiolejeunea latipuensis Tixier, Cryptog. Bryol. Lichénol. 16 (3): 230, 1995. Based on: Diplasiolejeunea latipuensis Tixier, Candollea 46 (2): 294, 1991, nom. inval.

** Diplasiolejeunea lemuriana Tixier, Lindbergia 4 (1/2): 120, 1977.

** Diplasiolejeunea longilobula Herzog, Trans. Brit. Bryol. Soc. 1 (4): 325, 1950.

** Diplasiolejeunea magnistipula Tixier, Ann. Fac. Sci. Yaoundé 20: 7, 1975.

*** Diplasiolejeunea mayaykuensis Schäf.-Verw. et Heinrichs, Polish Bot. J. 58 (1): 143, 2013.

*** Diplasiolejeunea onraedtii Grolle, Feddes Repert. 89 (5/6): 301, 1978.

** Diplasiolejeunea ramicola Tixier, Rev. Bryol. Lichénol. 45 (2): 222, 1979.

** Diplasiolejeunea riclefgrollei Schäf.-Verw., Cryptog. Bryol. 26 (1): 37, 2005.

** Diplasiolejeunea subcornuta Tixier, Lindbergia 4 (1/2): 122, 1977.

*** Diplasiolejeunea symoensii Vanden Berghen, Bull. Soc. Roy. Bot. Belgique 92: 126, 1960.

** Haplolejeunea Grolle, J. Hattori Bot. Lab. 39: 205, 1975.

*** Haplolejeunea cucullata (Steph.) Grolle, J. Hattori Bot. Lab. 45: 176, 1979. Bas.: Cheilolejeunea cucullata Steph., Sp. Hepat. (Stephani) 5: 644, 1914.

** Haplolejeunea sticta Grolle, J. Hattori Bot. Lab. 39: 205, 1975.

** Macrocolura R.M.Schust., J. Hattori Bot. Lab. 75: 233, 1994.

*** Macrocolura sagittistipula (Spruce) R.M.Schust., J. Hattori Bot. Lab. 75: 233, 1994. Bas.: Lejeunea sagittistipula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 304, 1884.

*** Myriocoleopsis Schiffn., Hedwigia 81 (5/6): 234, 1944.

*** Myriocoleopsis fluviatilis (Steph.) M.E.Reiner et Gradst., J. Bryol. 19 (3): 639, 1997. Bas.: Cololejeunea fluviatilis Steph., Hedwigia 34 (5): 248, 1895.

*** Myriocoleopsis gymnocolea (Spruce) M.E.Reiner et Gradst., J. Bryol. 19 (3): 640, 1997. Bas.: Lejeunea gymnocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 296, 1884.

*** Myriocoleopsis minutissima (Sm.) R.L.Zhu, Y.Yu et Pócs, Phytotaxa 183 (4): 293, 2014. Bas.: Jungermannia minutissima Sm., Engl. Bot. 23: tab. 1633, 1806.

*** Myriocoleopsis minutissima subsp. myriocarpa (Nees et Mont.) R.L.Zhu, Y.Yu et Pócs, Phytotaxa 183 (4): 294, 2014. Bas.: Lejeunea myriocarpa Nees et Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 473, 1842.

** Myriocoleopsis vuquangensis (Pócs et Ninh) Pócs, Trop. Bryol. 31: 124, 2010. Bas.: Cololejeunea vuquangensis Pócs et Ninh, Acta Bot. Hung. 47 (1/2): 156, 2005.

** Nephelolejeunea Grolle, J. Hattori Bot. Lab. 37: 252, 1973.

** Nephelolejeunea bidentata B.M.Thiers ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 65, 2015. Based on: Austrolejeunea bidentata B.M.Thiers, Bryologist 88 (4): 350, 1985 [1986], nom. inval.

** Nephelolejeunea carcharias M.A.M.Renner, Syst. Bot. 34 (4): 621, 2009.

*** Nephelolejeunea conchophylla Grolle, J. Hattori Bot. Lab. 48: 164, 1980.

** Nephelolejeunea fragilis (R.M.Schust.) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 65, 2015. Bas.: Cololejeunea fragilis R.M.Schust., Phytologia 56 (7): 458, 1985.

*** Nephelolejeunea hamata Grolle, J. Hattori Bot. Lab. 48: 167, 1980.

*** Nephelolejeunea hispida R.M.Schust. ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015. Based on: Austrolejeunea hispida R.M.Schust., Phytologia 47 (4): 305, 1981, nom. inval.

*** Nephelolejeunea jarmaniana Grolle ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015. Based on: Austrolejeunea jarmaniana Grolle, Nova Hedwigia 55 (1/2): 112, 1992, nom. inval.

*** Nephelolejeunea nudipes (Hook.f. et Taylor) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015. Bas.: Jungermannia nudipes Hook.f. et Taylor, London J. Bot. 3: 568, 1844.

** Nephelolejeunea occidentalis Pócs ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015. Based on: Austrolejeunea occidentalis Pócs, J. Hattori Bot. Lab. 99: 187, 2006, nom. inval.

*** Nephelolejeunea papillosa Glenny, New Zealand J. Bot. 34 (2): 195, 1996.

*** Nephelolejeunea radulifolia (C.Massal.) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015. Bas.: Lejeunea radulifolia C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 248, 1885.

** Nephelolejeunea secunda M.A.M.Renner ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015. Based on: Austrolejeunea secunda M.A.M.Renner, Bryologist 113 (4): 782, 2010, nom. inval.

*** Nephelolejeunea talinayi (S.W.Arnell) Grolle, J. Hattori Bot. Lab. 37: 253, 1973. Bas.: Harpalejeunea talinayi S.W.Arnell, Svensk Bot. Tidskr. 50 (2): 309, 1956.

** Schusterolejeunea Grolle, J. Bryol. 11 (1): 105, 1980. Nom. nov. pro Cladocolea R.M.Schust., Beih. Nova Hedwigia 9: 155, 1963.

*** Schusterolejeunea inundata (Spruce) Grolle, J. Bryol. 11 (1): 105, 1980. Bas.: Lejeunea inundata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 278, 1884.

** Siphonolejeunea Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 744, 1942.

** Siphonolejeunea elegantissima (Steph.) Grolle, J. Hattori Bot. Lab. 41: 405, 1976. Bas.: Trachylejeunea elegantissima Steph., Hedwigia 28 (4): 262, 1889.

** Siphonolejeunea neesii (Mont.) Bischl., Nova Hedwigia 17: 338, 1969. Bas.: Lejeunea neesii Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 62, 1836.

*** Siphonolejeunea schiffneri (Schiffn.) Herzog, Svensk Bot. Tidskr. 42 (3): 230, 1948. Bas.: Pycnolejeunea schiffneri Schiffn., Consp. Hepat. Arch. Ind.: 260, 1898.

** Tuyamaella S.Hatt., J. Hattori Bot. Lab. 5: 60, 1951.

** Tuyamaella angulistipa (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 508, 1961. Bas.: Pycnolejeunea angulistipa Steph., Hedwigia 35 (3): 123, 1896.

** Tuyamaella borneensis Tixier, Rev. Bryol. Lichénol. 39 (2): 241, 1973.

** Tuyamaella hattorii Tixier, Rev. Bryol. Lichénol. 31 (3/4): 188, 1962 [1963].

** Tuyamaella jackii (Steph.) Tixier, Rev. Bryol. Lichénol. 39 (2): 235, 1973. Bas.: Pycnolejeunea jackii Steph., Sp. Hepat. (Stephani) 5: 611, 1914.

*** Tuyamaella molischii (Schiffn.) S.Hatt., J. Hattori Bot. Lab. 5: 62, 1951. Bas.: Pycnolejeunea molischii Schiffn., Ann. Bryol. 2: 97, 1929.

** Tuyamaella molischii var. brevistipa P.C.Wu et P.J.Lin, Acta Phytotax. Sin. 16 (2): 65, 1978.

** Tuyamaella molischii var. taiwanensis R.L.Zhu et M.L.So, Nova Hedwigia 70 (1/2): 190, 2000.

** Tuyamaella serratistipa S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 118, 1951.

** subtrib. Cyclolejeuneinae Gradst.

*** Bromeliophila R.M.Schust., J. Hattori Bot. Lab. 75: 226, 1994.

*** Bromeliophila helenae Gradst., Cryptog. Bryol. Lichénol. 18 (3): 218, 1997.

*** Bromeliophila natans (Steph.) R.M.Schust., J. Hattori Bot. Lab. 75: 226, 1994. Bas.: Peltolejeunea natans Steph., Hedwigia 44 (4): 228, 1905.

*** Cyclolejeunea A.Evans, Bull. Torrey Bot. Club 31 (4): 193, 1904.

*** Cyclolejeunea accedens (Gottsche) A.Evans, Bull. Torrey Bot. Club 31 (4): 201, 1904. Bas.: Lejeunea accedens Gottsche, Syn. Hepat. 3: 339, 1845.

*** Cyclolejeunea chitonia (Taylor) A.Evans, Bull. Torrey Bot. Club 31 (4): 194, 1904. Bas.: Lejeunea chitonia Taylor, Nov. Stirp. Pug. 8: 27, 1844.

*** Cyclolejeunea convexistipa (Lehm. et Lindenb.) A.Evans, Bull. Torrey Bot. Club 31 (4): 198, 1904. Bas.: Jungermannia convexistipa Lehm. et Lindenb., Nov. Stirp. Pug. 6: 43, 1834.

* Cyclolejeunea ecuadorensis Steph., Sp. Hepat. (Stephani) 5: 194, 1913.

*** Cyclolejeunea foliorum (Nees) Grolle, J. Hattori Bot. Lab. 65: 403, 1988. Bas.: Lejeunea foliorum Nees, Syn. Hepat. 3: 326, 1845.

* Cyclolejeunea integerrima (Steph.) Steph., Sp. Hepat. (Stephani) 5: 195, 1913. Bas.: Odontolejeunea integerrima Steph., Hedwigia 44 (4): 227, 1905.

*** Cyclolejeunea luteola (Spruce) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 33: 761, 1984. Bas.: Lejeunea luteola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 205, 1884.

*** Cyclolejeunea peruviana (Lehm. et Lindenb.) A.Evans, Bull. Torrey Bot. Club 31 (4): 196, 1904. Bas.: Jungermannia peruviana Lehm. et Lindenb., Nov. Stirp. Pug. 5: 18, 1833.

* Cyclolejeunea spectabilis Steph., Sp. Hepat. (Stephani) 5: 193, 1913.

*** Prionolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893. Bas.: Lejeunea subg. Prionolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 152, 1884.

*** Prionolejeunea aemula (Gottsche) A.Evans, Bull. Torrey Bot. Club 31 (4): 219, 1904. Bas.: Lejeunea aemula Gottsche, Syn. Hepat. 3: 338, 1845.

*** Prionolejeunea ampliretis Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 178, 1955.

*** Prionolejeunea arguta (Nees) Steph., Sp. Hepat. (Stephani) 5: 204, 1913. Bas.: Lejeunea arguta Nees, Syn. Hepat. 3: 338, 1845.

*** Prionolejeunea decora (Taylor) Steph., Sp. Hepat. (Stephani) 5: 207, 1913. Bas.: Lejeunea decora Taylor, London J. Bot. 5: 393, 1846.

*** Prionolejeunea denticulata (F.Weber) Schiffn., Hepat. (Engl.-Prantl): 127, 1893. Bas.: Jungermannia denticulata F.Weber, Hist. Musc. Hepat. Prodr.: 30, 1815.

*** Prionolejeunea diversitexta (Hampe et Gottsche) Steph., Sp. Hepat. (Stephani) 5: 209, 1913. Bas.: Lejeunea diversitexta Hampe et Gottsche, Linnaea 25 (3): 357, 1852 [1853].

*** Prionolejeunea exauriculata A.Evans, Bull. Torrey Bot. Club 31 (4): 223, 1904.

*** Prionolejeunea galliotii Steph., Sp. Hepat. (Stephani) 6: 388, 1923.

*** Prionolejeunea grata (Gottsche) Schiffn., Hepat. (Engl.-Prantl): 127, 1893. Bas.: Lejeunea grata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 359, 1882.

*** Prionolejeunea grollei Ilk.-Borg. et Schäf.-Verw., Cryptog. Bryol. 26 (1): 29, 2005.

*** Prionolejeunea guadalupensis (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 211, 1913. Bas.: Lejeunea guadalupensis Lindenb., Syn. Hepat. 3: 340, 1845.

*** Prionolejeunea limpida Herzog, Hedwigia 67 (6): 252, 1927.

*** Prionolejeunea magnistipula Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 176, 1955.

*** Prionolejeunea meissneri (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 212, 1913. Bas.: Lejeunea meissneri Gottsche, Syn. Hepat. 3: 340, 1845.

*** Prionolejeunea mucronata (Sande Lac.) Steph., Hedwigia 35 (3): 119, 1896. Bas.: Lejeunea mucronata Sande Lac., Syn. hepat. jav.: 106, 1856 [1857].

*** Prionolejeunea muricatoserrulata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 223, 1913. Bas.: Lejeunea muricatoserrulata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 155, 1884.

*** Prionolejeunea principensis Vanden Berghen, Rev. Bryol. Lichénol. 29 (1/2): 65, 1960.

*** Prionolejeunea recurvula (Spruce) Steph., Sp. Hepat. (Stephani) 5: 224, 1913. Bas.: Lejeunea recurvula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 155, 1884.

*** Prionolejeunea scaberula (Spruce) Steph., Sp. Hepat. (Stephani) 5: 214, 1913. Bas.: Lejeunea scaberula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 159, 1884.

*** Prionolejeunea schlimiana (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 214, 1913. Bas.: Lejeunea schlimiana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 154, 1864.

*** Prionolejeunea trachyodes (Spruce) Steph., Sp. Hepat. (Stephani) 5: 215, 1913. Bas.: Lejeunea trachyodes Spruce, J. Linn. Soc., Bot. 30 (210): 338, 1895.

Excluded from the genus

* Prionolejeunea corbisieri Pearson, Natuurw. Tijdschr. 4 (5/6): 128, 1922. 297

* Prionolejeunea maculata Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 180, 1955. 298

** subtrib. Drepanolejeuneinae Gradst.

** Capillolejeunea S.W.Arnell, Svensk Bot. Tidskr. 59 (1): 69, 1965.

** Capillolejeunea geisslerae (Pócs) R.L.Zhu, Qiong He, Y.M.Wei et Pócs, Phytotaxa 175 (3): 167, 2014. Bas.: Drepanolejeunea geisslerae Pócs, Candollea 56 (1): 70, 2001.

** Capillolejeunea mascarena S.W.Arnell, Svensk Bot. Tidskr. 59 (1): 69, 1965.

*** Drepanolejeunea (Spruce) Steph., Hedwigia 30 (5): 209, 1891. Bas.: Lejeunea subg. Drepanolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 186, 1884.

** subg. Acantholejeunea R.M.Schust., Beih. Nova Hedwigia 9: 115, 1963.

** Drepanolejeunea dentistipula Steph., Sp. Hepat. (Stephani) 5: 343, 1913.

** Drepanolejeunea spinistipula Herzog, Svensk Bot. Tidskr. 42 (3): 238, 1948.

*** subg. Drepanolejeunea

*** Drepanolejeunea aculeata Bischl., Rev. Bryol. Lichénol. 33 (1/2): 68, 1964.

** Drepanolejeunea anderssonii (Ångstr.) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 429, 1900. Bas.: Lejeunea anderssonii Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 24, 1872.

*** Drepanolejeunea andina Herzog, Svensk Bot. Tidskr. 51 (1): 196, 1957.

*** Drepanolejeunea angustifolia (Mitt.) Grolle, J. Jap. Bot. 40 (7): 206, 1965. Bas.: Lejeunea angustifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 116, 1860 [1861].

*** Drepanolejeunea ankasica E.W.Jones, Bull. Brit. Mus. (Nat. Hist.), Bot. 11 (3): 245, 1983.

*** Drepanolejeunea anoplantha (Spruce) Steph., Sp. Hepat. (Stephani) 5: 325, 1913. Bas.: Lejeunea anoplantha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 189, 1884.

*** Drepanolejeunea appalachiana R.M.Schust., J. Elisha Mitchell Sci. Soc. 83 (4): 219, 1967.

*** Drepanolejeunea araucariae Steph., Hedwigia 35 (3): 80, 1896.

** Drepanolejeunea araucariae var. chilensis Herzog, Acta Horti Gotob. 15: 161, 1943.

** Drepanolejeunea aucklandica Steph., Sp. Hepat. (Stephani) 5: 358, 1913.

*** Drepanolejeunea aurita Bischl., Rev. Bryol. Lichénol. 33 (1/2): 146, 1964.

** Drepanolejeunea bakeri Herzog, Ann. Bryol. 3: 148, 1930.

*** Drepanolejeunea bidens (Prantl) A.Evans, Bull. Torrey Bot. Club 30 (1): 29, 1903. Bas.: Lejeunea bidens Prantl, Hedwigia 29: xiv, 1890.

*** Drepanolejeunea biocellata A.Evans, Bull. Torrey Bot. Club 30 (1): 22, 1903.

** Drepanolejeunea blumei Steph., Hedwigia 35 (3): 81, 1896.

** Drepanolejeunea brunnea Mizut., J. Hattori Bot. Lab. 33: 231, 1970.

** Drepanolejeunea caledonica Steph., Sp. Hepat. (Stephani) 5: 342, 1913.

*** Drepanolejeunea campanulata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 328, 1913. Bas.: Lejeunea campanulata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 192, 1884.

** Drepanolejeunea canceroides H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 53, 1963.

** Drepanolejeunea capulata (Taylor) J.B.Jack et Steph., Hedwigia 31 (1): 13, 1892. Bas.: Lejeunea capulata Taylor, London J. Bot. 5: 394, 1846.

** Drepanolejeunea capulata var. flagellifera S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 100, 1964.

** Drepanolejeunea ciliata Mizut., J. Hattori Bot. Lab. 33: 232, 1970.

*** Drepanolejeunea crassiretis A.Evans, Bull. Torrey Bot. Club 30 (1): 25, 1903.

*** Drepanolejeunea crucianella (Taylor) A.Evans, Bull. Torrey Bot. Club 30 (1): 33, 1903. Bas.: Lejeunea crucianella Taylor, London J. Bot. 5: 393, 1846.

*** Drepanolejeunea cultrella (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 324, 1913. Bas.: Lejeunea cultrella Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 168, 1863.

** Drepanolejeunea cutervoensis (Loitl.) Grolle, J. Hattori Bot. Lab. 69: 186, 1991. Bas.: Lejeunea cutervoensis Loitl., Diagn. pl. nov.: 21, 1894.

** Drepanolejeunea decurviloba Steph., Sp. Hepat. (Stephani) 6: 397, 1923.

* Drepanolejeunea deslooveri Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 210, 1977. 299

* Drepanolejeunea dissitifolia A.Evans, Bull. Torrey Bot. Club 30 (1): 28, 1903.

** Drepanolejeunea elegans Herzog, Ann. Bryol. 9: 128, 1936 [1937].

** Drepanolejeunea erecta (Steph.) Mizut., J. Hattori Bot. Lab. 40: 442, 1976. Bas.: Leptolejeunea erecta Steph., Bull. Soc. Roy. Bot. Belgique 38 (1): 44, 1899.

** Drepanolejeunea evansii Bischl. ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 47, 2012. Based on: Drepanolejeunea evansii Bischl., Rev. Bryol. Lichénol. 33 (1/2): 75, 1964, nom. inval.

*** Drepanolejeunea fragilis Bischl. ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 47, 2012. Based on: Drepanolejeunea fragilis Bischl., Rev. Bryol. Lichénol. 33 (1/2): 123, 1964, nom. inval.

** Drepanolejeunea fulfordiae L.Söderstr., Phytotaxa 208 (1): 98, 2015. Nom. nov. pro Drepanolejeunea papillosa Fulford, Mem. New York Bot. Gard. 23: 843, 1972, nom. illeg.

*** Drepanolejeunea granatensis (Prantl) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 150, 1964. Bas.: Lejeunea granatensis Prantl, Hedwigia 31: xvi, 1892.

** Drepanolejeunea grandis Herzog, Ann. Bryol. 12: 113, 1939.

*** Drepanolejeunea grollei M.E.Reiner et Schäf.-Verw., Candollea 51 (2): 475, 1996.

** Drepanolejeunea grossidens Steph., Sp. Hepat. (Stephani) 5: 358, 1913. Based on: Drepanolejeunea grossidens Steph., Hedwigia 28 (3): 168, 1889, nom. inval.

*** Drepanolejeunea hamatifolia (Hook.) Schiffn., Hepat. (Engl.-Prantl): 126, 1893. Bas.: Jungermannia hamatifolia Hook., Brit. Jungermann.: tab. 51, 1813.

** Drepanolejeunea hamulata Steph., Sp. Hepat. (Stephani) 5: 331, 1913.

*** Drepanolejeunea helenae Pócs, Cryptog. Bryol. Lichénol. 18 (3): 198, 1997.

** Drepanolejeunea herzogii R.L.Zhu et M.L.So, Beih. Nova Hedwigia 121: 181, 2001. Nom. nov. pro Strepsilejeunea ocellata Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 57, 1950 [1951].

*** Drepanolejeunea inchoata (C.F.W.Meissn.) Steph., Primit. fl. costar.: 115, 1892 [1893]. Bas.: Jungermannia inchoata C.F.W.Meissn., Nov. Stirp. Pug. 5: 19, 1833.

** Drepanolejeunea inchoata var. palmicola Pócs, Acta Bot. Hung. 51 (3/4): 381, 2009.

** Drepanolejeunea inchoata var. roraimae (Steph. ex Zwickel) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 48, 1964. Bas.: Drepanolejeunea roraimae Steph. ex Zwickel, Ann. Bryol. 6: 121, 1933.

*** Drepanolejeunea infundibulata (Spruce) A.Evans, Bull. Torrey Bot. Club 30 (1): 35, 1903. Bas.: Lejeunea infundibulata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 191, 1884.

*** Drepanolejeunea integribracteata Bischl., Rev. Bryol. Lichénol. 33 (1/2): 142, 1964.

** Drepanolejeunea laevis (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 357, 1913. Bas.: Lejeunea laevis Mitt., Fl. vit.: 416, 1871 [1873].

* Drepanolejeunea lancifolia (Gottsche) J.B.Jack et Steph., Hedwigia 31 (1): 13, 1892. Bas.: Lejeunea lancifolia Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 155, 1864. 300

*** Drepanolejeunea lichenicola (Spruce) Steph., Sp. Hepat. (Stephani) 5: 335, 1913. Bas.: Lejeunea lichenicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 191, 1884.

** Drepanolejeunea longii Grolle et R.L.Zhu, Ann. Bot. Fenn. 36 (2): 115, 1999.

** Drepanolejeunea macrodonta (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 357, 1913. Bas.: Lejeunea macrodonta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 116, 1860 [1861].

** Drepanolejeunea mawtmiana Ajit P.Singh et V.Nath, Hepat. Khasi Jaintia Hills: E. Himal.: 251, 2007.

** Drepanolejeunea microcarpa Pearson, J. Linn. Soc., Bot. 46 (305): 36, 1922.

** Drepanolejeunea moluccensis Herzog, Ann. Bryol. 7: 88, 1934.

*** Drepanolejeunea mosenii (Steph.) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 118, 1967 [1968]. Bas.: Leptolejeunea mosenii Steph., Sp. Hepat. (Stephani) 5: 372, 1913.

* Drepanolejeunea obliqua Steph., Hedwigia 35 (3): 82, 1896. 301

** Drepanolejeunea obtriangulata T.Kodama, J. Hattori Bot. Lab. 41: 381, 1976.

** Drepanolejeunea obtusifolia T.Yamag., J. Jap. Bot. 59 (11): 332, 1984.

*** Drepanolejeunea orthophylla (Nees et Mont.) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 102, 1967 [1968]. Bas.: Lejeunea orthophylla Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 265, 1843.

*** Drepanolejeunea palmifolia (Nees) Schiffn., Hepat. (Engl.-Prantl): 126, 1893. Bas.: Jungermannia palmifolia Nees, Fl. Bras. (Martius) 1 (1): 366, 1833.

*** Drepanolejeunea pentadactyla (Mont.) Steph., Sp. Hepat. (Stephani) 5: 357, 1913. Bas.: Lejeunea pentadactyla Mont., Ann. Sci. Nat. Bot. (sér. 3) 10: 113, 1848.

** Drepanolejeunea perissodonta (Spruce) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 73, 1964. Bas.: Lejeunea inchoata var. perissodonta Spruce, J. Linn. Soc., Bot. 30 (210): 340, 1895.

*** Drepanolejeunea physifolia (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 8, 1892. Bas.: Lejeunea physifolia Gottsche, Abh. Naturwiss. Vereins Bremen 7: 357, 1882.

** Drepanolejeunea pinnatiloba Schiffn., Bot. Jahrb. Syst. 23 (5): 591, 1897.

** Drepanolejeunea pleiodictya Herzog, Ann. Bryol. 7: 89, 1934.

** Drepanolejeunea propagulifera Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 204, 1931.

** Drepanolejeunea pseudoneura (A.Evans) Grolle, J. Hattori Bot. Lab. 65: 405, 1988. Bas.: Harpalejeunea pseudoneura A.Evans, Trans. Connecticut Acad. Arts 10 (8): 427, 1900.

** Drepanolejeunea pterocalyx (Herzog) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 114, 1967 [1968]. Bas.: Leptolejeunea pterocalyx Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 187, 1955.

** Drepanolejeunea pungens Bischl., Rev. Bryol. Lichénol. 33 (1/2): 104, 1964.

** Drepanolejeunea ramentiflora Steph., Sp. Hepat. (Stephani) 5: 338, 1913.

* Drepanolejeunea ruandensis Vanden Berghen, Bull. Soc. Roy. Bot. Belgique 93: 63, 1961. 302

*** Drepanolejeunea senticosa Bischl., Rev. Bryol. Lichénol. 33 (1/2): 96, 1964.

** Drepanolejeunea sikkimensis (Udar et U.S.Awasthi) Grolle, J. Hattori Bot. Lab. 55: 503, 1984. Bas.: Leptolejeunea sikkimensis Udar et U.S.Awasthi, Misc. Bryol. Lichenol. 8 (6): 115, 1979.

** Drepanolejeunea spinosa Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 185, 1955.

** Drepanolejeunea spinosocornuta Steph., Sp. Hepat. (Stephani) 5: 351, 1913.

* Drepanolejeunea subdissitifolia Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 65, 1950.

** Drepanolejeunea submuricata R.M.Schust., Phytotaxa 208 (1): 98, 2015. Based on: Drepanolejeunea submuricata R.M.Schust., Nova Hedwigia 62 (1/2): 34, 1996, nom. inval.

** Drepanolejeunea subquadrata (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 311, 1896. Bas.: Lejeunea subquadrata Mitt., Fl. vit.: 416, 1871 [1873].

** Drepanolejeunea subvittata (Herzog) Grolle, J. Hattori Bot. Lab. 69: 186, 1991. Bas.: Harpalejeunea subvittata Herzog, Svensk Bot. Tidskr. 51 (1): 195, 1957.

* Drepanolejeunea tenax K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 15, 1928.

** Drepanolejeunea tenera K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 20, 1928. 303

* Drepanolejeunea tenera var. litoceras Herzog, Ann. Bryol. 7: 85, 1934.

*** Drepanolejeunea ternatensis (Gottsche) Schiffn., Hepat. (Engl.-Prantl): 126, 1893. Bas.: Lejeunea ternatensis Gottsche, Syn. Hepat. 3: 346, 1845.

** Drepanolejeunea teysmannii (Gottsche) Steph., Hedwigia 35 (3): 84, 1896. Bas.: Lejeunea teysmannii Gottsche, Abh. Naturwiss. Vereins Bremen 7: 360, 1882.

** Drepanolejeunea tridactyla (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 354, 1913. Bas.: Lejeunea tridactyla Gottsche, Syn. Hepat. 3: 347, 1845.

*** Drepanolejeunea trigonophylla Steph., Hedwigia 35 (3): 85, 1896.

** Drepanolejeunea tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 7, 1958.

** Drepanolejeunea tuyamae S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 116, 1951.

** Drepanolejeunea ualanensis Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 148, 1965.

** Drepanolejeunea ungulata (Steph.) Grolle, J. Hattori Bot. Lab. 69: 187, 1991. Bas.: Harpalejeunea ungulata Steph., Sp. Hepat. (Stephani) 5: 264, 1913.

** Drepanolejeunea urceolata R.M.Schust., Phytologia 39 (6): 427, 1978.

** Drepanolejeunea valiae Jovet-Ast, Rev. Bryol. Lichénol. 18 (1/2): 38, 1949.

** Drepanolejeunea vandenberghenii Buchb. et Eb.Fisch., J. Bryol. 26 (4): 273, 2004.

*** Drepanolejeunea vesiculosa (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 356, 1913. Bas.: Lejeunea vesiculosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 116, 1860 [1861].

** Drepanolejeunea yulensis Steph., Sp. Hepat. (Stephani) 5: 356, 1913.

*** subg. Kolpolejeunea Grolle, J. Hattori Bot. Lab. 40: 193, 1976.

*** Drepanolejeunea intermedia Zwickel, Ann. Bryol. 6: 119, 1933.

*** Drepanolejeunea lyrata Grolle, J. Hattori Bot. Lab. 40: 199, 1976.

*** Drepanolejeunea madagascariensis (Steph.) Grolle, Lindbergia 2 (3/4): 232, 1974. Bas.: Leptolejeunea madagascariensis Steph., Sp. Hepat. (Stephani) 5: 363, 1913.

*** Drepanolejeunea pocsii Grolle, J. Hattori Bot. Lab. 40: 209, 1976.

** Drepanolejeunea symoensii Vanden Berghen et Grolle, J. Hattori Bot. Lab. 49: 86, 1981. Based on: Leptolejeunea symoensii Vanden Berghen, Bull. Soc. Roy. Bot. Belgique 93: 58, 1961, nom. inval.

** Drepanolejeunea symoensii var. minor Tixier, Trop. Bryol. 11: 25, 1995.

*** Drepanolejeunea trematodes (Nees) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 125, 1967 [1968]. Bas.: Lejeunea trematodes Nees, Ann. Sci. Nat. Bot. (sér. 2) 5: 63, 1836.

** subg. Pristolejeunea Grolle, J. Hattori Bot. Lab. 40: 193, 1976.

** Drepanolejeunea actinogyna Inuthai, R.L.Zhu et Chantanaorr., Bryologist 117 (2): 165, 2014.

*** Drepanolejeunea fissicornua Steph., Sp. Hepat. (Stephani) 5: 344, 1913.

* Drepanolejeunea hampeana Steph., Sp. Hepat. (Stephani) 5: 345, 1913. Nom. nov. pro Drepanolejeunea hampeana Steph., Hedwigia 29 (2): 70, 1890, nom. inval.

** Drepanolejeunea laciniata Qiong He et R.L.Zhu, Cryptog. Bryol. 33 (3): 292, 2012.

*** Drepanolejeunea levicornua Steph., Sp. Hepat. (Stephani) 5: 347, 1913.

* Drepanolejeunea longicornua (Herzog) Mizut., J. Hattori Bot. Lab. 68: 368, 1990. Bas.: Drepanolejeunea levicornua var. longicornua Herzog, Ann. Bryol. 3: 142, 1930.

* Drepanolejeunea nymanii Steph., Sp. Hepat. (Stephani) 5: 348, 1913.

** Drepanolejeunea pulla (Mitt.) Grolle, J. Hattori Bot. Lab. 46: 349, 1979. Bas.: Lejeunea pulla Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 116, 1860 [1861].

* Drepanolejeunea serricalyx Herzog, Ann. Bryol. 9: 126, 1936 [1937].

*** Drepanolejeunea thwaitesiana (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 350, 1913. Bas.: Lejeunea thwaitesiana Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 117, 1860 [1861]. 304

** Drepanolejeunea thwaitesiana var. zhengii R.L.Zhu, Beih. Nova Hedwigia 121: 197, 2001.

*** Drepanolejeunea tricornua Herzog, Ann. Bryol. 9: 124, 1936 [1937].

*** subg. Rhaphidolejeunea (Herzog) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 376, 2000. Bas.: Rhaphidolejeunea Herzog, Mitth. Thüring. Bot. Vereins 50: 104, 1943.

** Drepanolejeunea bidoupensis Pócs, Cryptog. Bryol. 34 (3): 293, 2013.

*** Drepanolejeunea bischlerae (Grolle) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 391, 2000. Bas.: Rhaphidolejeunea bischlerae Grolle, J. Hattori Bot. Lab. 38: 653, 1974.

*** Drepanolejeunea commutata Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 377, 2000.

*** Drepanolejeunea cyclops (Sande Lac.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 391, 2000. Bas.: Lejeunea cyclops Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 310, 1864.

*** Drepanolejeunea fleischeri (Steph.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 379, 2000. Bas.: Leptolejeunea fleischeri Steph., Sp. Hepat. (Stephani) 5: 382, 1913.

** Drepanolejeunea foliicola Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 85, 1932.

** Drepanolejeunea longicruris (Steph.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 392, 2000. Bas.: Leptolejeunea longicruris Steph., Hedwigia 35 (3): 106, 1896.

*** Drepanolejeunea polyrhiza (Nees) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 392, 2000. Bas.: Lejeunea polyrhiza Nees, Syn. Hepat. 3: 403, 1845.

*** Drepanolejeunea siamensis (Bischl.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 393, 2000. Bas.: Rhaphidolejeunea siamensis Bischl., Rev. Bryol. Lichénol. 36 (1/2): 86, 1968 [1969].

*** Drepanolejeunea spicata (Steph.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 384, 2000. Bas.: Leptolejeunea spicata Steph., Hedwigia 35 (3): 108, 1896.

*** Drepanolejeunea tibetana (P.C.Wu et J.S.Lou) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 386, 2000. Bas.: Rhaphidolejeunea tibetana P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 102, 1978.

*** Drepanolejeunea yunnanensis (P.C.Chen) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 388, 2000. Bas.: Rhaphidolejeunea yunnanensis P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 44, 1955.

Incertae sedis

*** Drepanolejeunea dactylophora (Nees, Lindenb. et Gottsche) J.B.Jack et Steph., Hedwigia 31 (1): 12, 1892. Bas.: Lejeunea dactylophora Nees, Lindenb. et Gottsche, Observ. bot.: 473, 1843.

*** Drepanolejeunea dactylophora var. submuricata Herzog, Ann. Bryol. 4: 92, 1931.

* Drepanolejeunea devendrae Sushil K.Singh et M.Dey, Nelumbo 54: 20, 2012.

* Drepanolejeunea integerrima Herzog, Rev. Bryol. Lichénol. 20 (1/2): 146, 1951 [1952]. 305

** Vitalianthus R.M.Schust. et Giancotti, Nova Hedwigia 57 (3/4): 447, 1993.

*** Vitalianthus bischlerianus (K.C.Pôrto et Grolle) R.M.Schust. et Giancotti, Nova Hedwigia 57 (3/4): 448, 1993. Bas.: Drepanolejeunea bischleriana K.C.Pôrto et Grolle, Cryptog. Bryol. Lichénol. 8 (4): 301, 1987.

** Vitalianthus guangxianus R.L.Zhu, Qiong He et Y.M.Wei, J. Bryol. 34 (1): 32, 2012.

** subtrib. Echinolejeuneinae Gradst.

*** Anoplolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893. Bas.: Lejeunea subg. Anoplolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 129, 1884.

*** Anoplolejeunea conferta (C.F.W.Meissn. ex Spreng.) A.Evans, Bull. Torrey Bot. Club 35 (4): 175, 1908. Bas.: Jungermannia conferta C.F.W.Meissn. ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827.

*** Echinolejeunea R.M.Schust., Beih. Nova Hedwigia 9: 187, 1963.

*** Echinolejeunea papillata (Mitt.) R.M.Schust. ex Hamlin, Rec. Domin. Mus. 7: 260, 1972. Bas.: Lejeunea papillata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 158, 1854.

** Kymatolejeunea Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 32 (6): 1005, 1984.

*** Kymatolejeunea bartlettii Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 32 (6): 1005, 1984.

** subtrib. Leiolejeuneinae Schäf.-Verw. et Heinrichs

** Leiolejeunea A.Evans, Bull. Torrey Bot. Club 35 (8): 377, 1908.

*** Leiolejeunea grandiflora A.Evans, Bull. Torrey Bot. Club 35 (8): 378, 1908.

*** subtrib. Lejeuneinae Gradst.

*** Harpalejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 126, 1893. Bas.: Lejeunea subg. Harpalejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 164, 1884.

** subg. Cleefiolejeunea Grolle et M.E.Reiner, J. Bryol. 21 (1): 33, 1999.

*** Harpalejeunea grandis Grolle et M.E.Reiner, J. Bryol. 21 (1): 32, 1999.

** subg. Harpalejeunea

* Harpalejeunea acuta S.Winkl., Rev. Bryol. Lichénol. 42 (3): 812, 1976. 306

*** Harpalejeunea ancistrodes (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893. Bas.: Lejeunea ancistrodes Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 169, 1884.

** Harpalejeunea buenaventurae Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 183, 1955.

*** Harpalejeunea cinchonae (Nees) Schiffn., Hepat. (Engl.-Prantl): 127, 1893. Bas.: Lejeunea cinchonae Nees, Syn. Hepat. 3: 342, 1845.

** Harpalejeunea cinchonae var. strigulosa Herzog, Svensk Bot. Tidskr. 51 (1): 192, 1957.

** Harpalejeunea decurvicuspis (Besch. et C.Massal.) P.Syd., Just’s Bot. Jahresber. 19: 246, 1894. Bas.: Lejeunea decurvicuspis Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 639, 1886.

** Harpalejeunea emarginata Jovet-Ast, Rev. Bryol. Lichénol. 16 (1/2): 38, 1947 [1948].

** Harpalejeunea exocellata Herzog, Rev. Bryol. Lichénol. 20 (1/2): 144, 1951 [1952].

** Harpalejeunea grandistipula R.M.Schust., J. Hattori Bot. Lab. 87: 290, 1999.

*** Harpalejeunea harpaphylla (Herzog) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 164, 1964. Bas.: Drepanolejeunea harpaphylla Herzog, Svensk Bot. Tidskr. 46 (1): 93, 1952.

** Harpalejeunea herzogii Jovet-Ast, Feddes Repert. Spec. Nov. Regni Veg. 58: 19, 1955.

** Harpalejeunea longibracteata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 254, 1913. Bas.: Lejeunea longibracteata Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cciii, 1889 [1890].

** Harpalejeunea marginalis (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 271, 1913. Bas.: Jungermannia marginalis Hook.f. et Taylor, London J. Bot. 4: 91, 1845.

*** Harpalejeunea molleri (Steph.) Grolle, Taxon 38 (1): 89, 1989. Bas.: Lejeunea molleri Steph., Hedwigia 26 (1): 3, 1887.

** Harpalejeunea molleri subsp. integra (R.M.Schust.) Damsh., Ill. Fl. Nord. Liverw. Hornw.: 615, 2002. Bas.: Harpalejeunea ovata subsp. integra R.M.Schust., J. Elisha Mitchell Sci. Soc. 83 (4): 199, 1967.

*** Harpalejeunea oxyphylla (Nees et Mont.) Steph., Sp. Hepat. (Stephani) 5: 255, 1913. Bas.: Lejeunea oxyphylla Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 264, 1843.

*** Harpalejeunea parasitica (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 268, 1913. Bas.: Jungermannia parasitica Hook.f. et Taylor, London J. Bot. 3: 477, 1844.

** Harpalejeunea pinaundensis Grolle, J. Hattori Bot. Lab. 46: 44, 1979.

** Harpalejeunea reflexula A.Evans, Bull. Torrey Bot. Club 35 (8): 375, 1908.

** Harpalejeunea scabra Gradst. et Schäf.-Verw., Cryptog. Bryol. 32 (2): 102, 2011.

*** Harpalejeunea schiffneri S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 102, 1964.

* Harpalejeunea solitaria (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 246, 1913. Bas.: Lejeunea solitaria Gottsche, Abh. Naturwiss. Vereins Bremen 7: 356, 1882.

*** Harpalejeunea stricta (Lindenb. et Gottsche) Steph., Sp. Hepat. (Stephani) 5: 259, 1913. Bas.: Lejeunea stricta Lindenb. et Gottsche, Syn. Hepat. 5: 756, 1847.

** Harpalejeunea subacuta A.Evans, Bull. Torrey Bot. Club 30 (10): 547, 1903.

* Harpalejeunea tenuicuspis (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893. Bas.: Lejeunea tenuicuspis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 170, 1884. 307

*** Harpalejeunea tridens (Besch. et Spruce) Steph., Sp. Hepat. (Stephani) 5: 263, 1913. Bas.: Lejeunea tridens Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxi, 1889 [1890].

** Harpalejeunea uncinata Steph., Hedwigia 35 (3): 97, 1896.

* Harpalejeunea uncinata var. setulosa Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 183, 1955.

Incertae sedis

* Harpalejeunea grossearmata Steph., Biblioth. Bot. 87 (2): 256, 1916. 308

* Harpalejeunea renneri Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 54, 1950 [1951]. 309

* Harpalejeunea spruceana Steph., Biblioth. Bot. 87 (2): 257, 1916. 310

* Harpalejeunea vitrea Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 94, 1952. 311

** Hattoriolejeunea Mizut., J. Hattori Bot. Lab. 61: 303, 1986 [1987].

** Hattoriolejeunea akiyamae Mizut., J. Hattori Bot. Lab. 61: 303, 1986 [1987].

*** Lejeunea Lib., Ann. Gen. Sci. Phys. 6: 373, 1820 nom. conserv. 312

** Lejeunea tunquiniensis M.E.Reiner et Drehwald, Nova Hedwigia 100 (3/4): 584, 2015.

** subg. Lejeunea

*** Lejeunea abyssinica (Gola) Cufod., Phyton (Horn) 4: 75, 1952. Bas.: Eulejeunea abyssinica Gola, Ann. Bot. (Rome) 13 (1): 70, 1914.

*** Lejeunea acanthogona Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 177, 1884.

** Lejeunea acuminata (Lehm. et Lindenb.) Lehm., Nov. Stirp. Pug. 7: 22, 1838. Bas.: Jungermannia acuminata Lehm. et Lindenb., Nov. Stirp. Pug. 6: 49, 1834.

** Lejeunea acuta Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 167, 1863.

** Lejeunea acutata (Steph.) Solari, J. Hattori Bot. Lab. 54: 541, 1983. Bas.: Strepsilejeunea acutata Steph., Hedwigia 35 (3): 127, 1896.

*** Lejeunea adpressa Nees, Repert. Pharm. 76: 45, 1842. 313

*** Lejeunea aethiopica E.W.Jones, J. Bryol. 13 (3): 387, 1985.

** Lejeunea alaskana (R.M.Schust. et Steere) Inoue et Steere, J. Hattori Bot. Lab. 44: 330, 1978. Bas.: Hygrolejeunea alaskana R.M.Schust. et Steere, Bull. Torrey Bot. Club 85 (3): 190, 1958.

*** Lejeunea alata Gottsche, Syn. Hepat. 3: 406, 1845.

** Lejeunea alata var. patriciae Pócs, Candollea 56 (1): 72, 2001.

*** Lejeunea albescens (Steph.) Mizut., J. Hattori Bot. Lab. 33: 245, 1970. Bas.: Taxilejeunea albescens Steph., Hedwigia 35 (3): 132, 1896.

* Lejeunea albiflora Colenso, Trans. & Proc. New Zealand Inst. 21: 72, 1889.

** Lejeunea aloba Sande Lac., Plagiochila Sandei: 10, 1856.

** Lejeunea alobifolia H.A.Mill., Phytologia 47 (4): 323, 1981. Nom. nov. pro Lejeunea aloba Steph., Sp. Hepat. (Stephani) 5: 767, 1915, nom. illeg.

** Lejeunea amaniensis E.W.Jones, J. Bryol. 13 (3): 392, 1985.

** Lejeunea ambigua Lindenb. et Gottsche, Syn. Hepat. 5: 764, 1847.

* Lejeunea amentulifera Steph., Sp. Hepat. (Stephani) 5: 707, 1915.

** Lejeunea androgyna R.M.Schust., Phytologia 45 (5): 432, 1980.

* Lejeunea angulifolia Mitt., Philos. Trans. 168: 400, 1879. 314

*** Lejeunea angusta (Lehm. et Lindenb.) Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 469, 1842. Bas.: Jungermannia angusta Lehm. et Lindenb., Nov. Stirp. Pug. 4: 52, 1832.

** Lejeunea anisophylla Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 263, 1843.

*** Lejeunea anomala Lindenb. et Gottsche, Linnaea 24 (6): 636, 1851 [1852].

* Lejeunea antillana Steph., Hedwigia 27 (11/12): 281, 1888.

** Lejeunea aphanes Spruce, J. Bot. 19: 36, 1881.

*** Lejeunea apiculata Sande Lac., Ned. Kruidk. Arch. 3: 421, 1854 [1855].

** Lejeunea aquatica Horik., Sci. Rep. Tôhoku Imp. Univ., Ser. 4, Biol. 5 (4): 643, 1929 [1930].

** Lejeunea aquatica var. apiculata S.Hatt., Bull. Tokyo Sci. Mus. 11: 108, 1944.

** Lejeunea armitii (Steph.) Steph., Sp. Hepat. (Stephani) 5: 768, 1915. Bas.: Eulejeunea armitii Steph., Hedwigia 28 (3): 169, 1889.

* Lejeunea asperifolia Steph., Sp. Hepat. (Stephani) 5: 708, 1915.

*** Lejeunea asperrima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 160, 1884.

** Lejeunea asperula (Steph.) Mizut., J. Hattori Bot. Lab. 33: 236, 1970. Bas.: Taxilejeunea asperula Steph., Sp. Hepat. (Stephani) 5: 499, 1914.

** Lejeunea asprella Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 175, 1884.

* Lejeunea atheatostipa Spruce, J. Bot. 33: 83, 1895.

** Lejeunea barbata (Herzog) R.L.Zhu et M.J.Lai, Ann. Bot. Fenn. 48 (5): 376, 2011. Bas.: Rectolejeunea barbata Herzog, J. Hattori Bot. Lab. 14: 49, 1955.

*** Lejeunea bermudiana (A.Evans) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 1105, 1980. Bas.: Crossotolejeunea bermudiana A.Evans, Bull. Torrey Bot. Club 33 (3): 132, 1906.

** Lejeunea bidentula Herzog, Symb. Sin. 5: 51, 1930.

** Lejeunea biformis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 162 (68), 1864.

** Lejeunea blepharogona Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 178, 1884.

** Lejeunea blomquistii R.M.Schust., J. Elisha Mitchell Sci. Soc. 78 (1): 64, 1962.

*** Lejeunea boliviensis (Steph.) R.L.Zhu et M.E.Reiner, Bryologist 107 (2): 237, 2004. Bas.: Strepsilejeunea boliviensis Steph., Biblioth. Bot. 87 (2): 257, 1916.

** Lejeunea boryana Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 47, 1838.

** Lejeunea brenanii E.W.Jones, J. Bryol. 10 (4): 391, 1979.

*** Lejeunea calcicola R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 404, 1957.

** Lejeunea calcicola var. mexicana R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 408, 1957.

** Lejeunea canariensis (Steph.) Steph., Sp. Hepat. (Stephani) 5: 802, 1915. Bas.: Eulejeunea canariensis Steph., Mém. Soc. Bot. France 7: 42, 1907.

*** Lejeunea cancellata Nees et Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 472, 1842.

** Lejeunea cantabrigiensis E.W.Jones, J. Bryol. 15 (4): 669, 1989.

*** Lejeunea capensis Gottsche, Syn. Hepat. 3: 374, 1845.

* Lejeunea caroliniana Austin, Bot. Bull. (Hanover) 1 (8): 36, 1876. 315

*** Lejeunea catinulifera Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 233, 1884.

*** Lejeunea caulicalyx (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 13, 2000. Bas.: Crossotolejeunea caulicalyx Steph., Sp. Hepat. (Stephani) 5: 237, 1913.

*** Lejeunea cavifolia (Ehrh.) Lindb., Revis. crit. icon.: 43, 1871. Bas.: Jungermannia cavifolia Ehrh., Beitr. Naturk. (Ehrhart) 4: 45, 1789.

** Lejeunea caviloba (Steph.) Besch., J. Bot. (Morot) 12: 140, 1898. Bas.: Eulejeunea caviloba Steph., Hedwigia 35 (3): 86, 1896.

*** Lejeunea cerina (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 391, 1845. Bas.: Jungermannia cerina Lehm. et Lindenb., Nov. Stirp. Pug. 5: 16, 1833.

** Lejeunea chaishanensis S.H.Lin, Yushania 9: 7, 1992.

** Lejeunea cladogyna A.Evans, Amer. J. Bot. 5 (3): 134, 1918.

** Lejeunea clavata Lindenb., Syn. Hepat. 3: 379, 1845.

** Lejeunea claviformis Lindenb. ex Steph., Sp. Hepat. (Stephani) 5: 727, 1915.

** Lejeunea cochleata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 273, 1884.

** Lejeunea cocoes Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 114, 1860 [1861].

*** Lejeunea colensoana (Steph.) M.A.M.Renner, Austral. Syst. Bot. 23 (6): 455, 2010. Bas.: Taxilejeunea colensoana Steph., Hedwigia 35 (3): 132, 1896.

** Lejeunea compacta (Steph.) Steph., Sp. Hepat. (Stephani) 5: 771, 1915. Bas.: Eulejeunea compacta Steph., Bull. Herb. Boissier 5 (2): 93, 1897.

** Lejeunea concinnula Spruce et Steph., J. Bot. 25: 39, 1887.

** Lejeunea connatistipula (Steph.) Steph., Sp. Hepat. (Stephani) 5: 772, 1915. Bas.: Eulejeunea connatistipula Steph., Hedwigia 35 (3): 87, 1896.

** Lejeunea contracta Mizut., J. Hattori Bot. Lab. 33: 248, 1970.

** Lejeunea controversa Gottsche, Hepat. Eur., Leberm. 56-57: no 556, 1873.

** Lejeunea convexiloba M.L.So et R.L.Zhu, Bryologist 101 (1): 137, 1998.

** Lejeunea corcovadae (Steph.) Bischl., Nova Hedwigia 5 (1/2): 406, 1963. Bas.: Microlejeunea corcovadae Steph., Sp. Hepat. (Stephani) 5: 820, 1915.

** Lejeunea cordiflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 283, 1884.

** Lejeunea corralensis A.Evans, Ann. Bryol. 3: 86, 1930.

*** Lejeunea corynantha Spruce, J. Linn. Soc., Bot. 30 (210): 344, 1895.

** Lejeunea crassiretis Mitt., Fl. vit.: 414, 1871 [1873].

*** Lejeunea cristulata (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 21, 2000. Bas.: Crossotolejeunea cristulata Steph., Hedwigia 35 (3): 75, 1896.

*** Lejeunea cristuliflora (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 19, 2000. Bas.: Crossotolejeunea cristuliflora Steph., Sp. Hepat. (Stephani) 5: 231, 1913.

** Lejeunea cuspidistipula (Steph.) Steph. ex Watts, Proc. Linn. Soc. New South Wales (ser. 2) 27 (108): 493, 1903. Bas.: Eulejeunea cuspidistipula Steph., Hedwigia 35 (3): 88, 1896.

** Lejeunea cyanomontana R.M.Schust., Phytologia 45 (5): 432, 1980.

** Lejeunea cyanophora R.M.Schust., J. Hattori Bot. Lab. 26: 246, 1963.

** Lejeunea cyathearum E.W.Jones, J. Bryol. 8 (1): 86, 1974.

*** Lejeunea cyathophora Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 359, 1851.

** Lejeunea denticalyx (Steph.) Steph., Sp. Hepat. (Stephani) 5: 794, 1915. Bas.: Eulejeunea denticalyx Steph., Hedwigia 28 (3): 169, 1889.

** Lejeunea denticuspis (Steph.) Mizut., J. Hattori Bot. Lab. 36: 160, 1972 [1973]. Bas.: Strepsilejeunea denticuspis Steph., Hedwigia 35 (3): 129, 1896.

** Lejeunea denudata (Pearson) J.J.Engel, Bryologist 78 (3): 361, 1975. Bas.: Eulejeunea denudata Pearson, J. Linn. Soc., Bot. 46 (305): 39, 1922.

*** Lejeunea deplanata Nees, Syn. Hepat. 3: 368, 1845.

** Lejeunea deplanata var. cuspidata (Steph.) M.E.Reiner, Nova Hedwigia 91 (3/4): 529, 2010. Bas.: Pycnolejeunea cuspidata Steph., Sp. Hepat. (Stephani) 5: 605, 1914.

** Lejeunea diaphana Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 168, 1884.

** Lejeunea dimorpha T.Kodama, J. Hattori Bot. Lab. 41: 384, 1976.

** Lejeunea dipterocarpa E.W.Jones, J. Bryol. 7 (1): 44, 1972.

*** Lejeunea discreta Lindenb., Syn. Hepat. 3: 361, 1845.

* Lejeunea disjecta Spruce, J. Linn. Soc., Bot. 30 (210): 347, 1895.

** Lejeunea diversicuspis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 176, 1884.

** Lejeunea drummondii Taylor, London J. Bot. 5: 400, 1846.

** Lejeunea ecarinata (Steph.) J.M.Coult., Barnes et Arthur, Bot. Gaz. 15 (12): 349, 1890. Bas.: Eulejeunea ecarinata Steph., Bot. Gaz. 15 (11): 283, 1890.

** Lejeunea eckloniana Lindenb., Syn. Hepat. 3: 381, 1845.

** Lejeunea eifrigii Mizut., J. Hattori Bot. Lab. 33: 244, 1970. Nom. nov. pro Taxilejeunea acutiloba Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 94, 1937.

* Lejeunea elongella Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 161, 1864.

*** Lejeunea erostrata M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 25, 2000. Nom. nov. pro Crossotolejeunea parva Steph., Sp. Hepat. (Stephani) 5: 241, 1913.

*** Lejeunea exilis (Reinw., Blume et Nees) Grolle, J. Hattori Bot. Lab. 46: 353, 1979. Bas.: Jungermannia exilis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 227, 1824 [1825].

** Lejeunea exilis var. abnormis (Herzog) G.E.Lee, Polish Bot. J. 58 (1): 61, 2013. Bas.: Byssolejeunea abnormis Herzog, Hedwigia 80 (1/2): 84, 1941.

** Lejeunea fernandeziana S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 16, 1957.

** Lejeunea firma Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 112, 1860 [1861].

** Lejeunea fissistipula (Steph.) Steph., Sp. Hepat. (Stephani) 5: 775, 1915. Bas.: Eulejeunea fissistipula Steph., Hedwigia 35 (3): 88, 1896.

** Lejeunea flagellaris Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 273, 1884.

*** Lejeunea flava (Sw.) Nees, Naturgesch. Eur. Leberm. 3: 277, 1838. Bas.: Jungermannia flava Sw., Prodr. (Swartz): 144, 1788. 316

** Lejeunea flava subsp. moorei (Lindb.) R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (1): 161, 1957. Bas.: Lejeunea moorei Lindb., Acta Soc. Sci. Fenn. 10: 487, 1875.

** Lejeunea flava subsp. orientalis R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (1): 161, 1957.

** Lejeunea flava var. pellucida Lindenb. et Gottsche, Linnaea 24 (6): 634, 1851 [1852].

** Lejeunea flava subsp. tabularis (Spreng.) S.W.Arnell, Hepat. South Africa: 199, 1963. Bas.: Jungermannia tabularis Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827.

*** Lejeunea flavovirens Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 144, 1873.

** Lejeunea fleischeri (Steph.) Mizut., J. Hattori Bot. Lab. 33: 238, 1970. Bas.: Hygrolejeunea fleischeri Steph., Sp. Hepat. (Stephani) 5: 560, 1914.

** Lejeunea floridana A.Evans, Bull. Torrey Bot. Club 32 (4): 185, 1905.

*** Lejeunea fulfordiae (Jovet-Ast) R.L.Zhu, Syst. Bot. 33 (4): 617, 2008. Bas.: Amblyolejeunea fulfordiae Jovet-Ast, Rev. Bryol. Lichénol. 17 (1/4): 25, 1948 [1949].

** Lejeunea fusagasugana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 158, 1864.

** Lejeunea galeata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 172, 1884.

** Lejeunea gayana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 157, 1864.

** Lejeunea gibbiloba (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981. Bas.: Eulejeunea gibbiloba Steph., Sp. Hepat. (Stephani) 6: 418, 1923.

*** Lejeunea glaucescens Gottsche, Syn. Hepat. 3: 378, 1845.

** Lejeunea glaucescens var. acrogyna R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 400, 1957.

* Lejeunea glaucescens var. obsoleta R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 395, 1957.

** Lejeunea globosiflora (Steph.) Steph., Sp. Hepat. (Stephani) 5: 795, 1915. Bas.: Eulejeunea globosiflora Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 65, 1900.

** Lejeunea gomphocalyx Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 174, 1884.

** Lejeunea gracilicaulis Spruce, Mem. Torrey Bot. Club 1 (3): 126, 1890.

** Lejeunea gracilipes (Taylor) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 213, 1884. Bas.: Omphalanthus gracilipes Taylor, London J. Bot. 5: 385, 1846.

** Lejeunea gracilis Steph., Sp. Hepat. (Stephani) 5: 777, 1915.

** Lejeunea gradsteiniana Pócs, Acta Biol. Pl. Agr. 1: 55, 2010 [2011]. Nom. nov. pro Ceratolejeunea aberrans Steph., Sp. Hepat. (Stephani) 6: 399, 1923.

** Lejeunea gradsteinii G.E.Lee, Damanhuri et Latiff, Acta Biol. Pl. Agr. 1: 29, 2010 [2011].

* Lejeunea grossecristata (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 5 (3): 556, 1968. Bas.: Hygrolejeunea grossecristata Steph., Hedwigia 35 (3): 102, 1896.

*** Lejeunea grossiretis (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 27, 2000. Bas.: Crossotolejeunea grossiretis Steph., Hedwigia 35 (3): 75, 1896.

** Lejeunea grossistipula Steph., Sp. Hepat. (Stephani) 5: 739, 1915.

*** Lejeunea grossitexta (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 29, 2000. Bas.: Crossotolejeunea grossitexta Steph., Sp. Hepat. (Stephani) 5: 240, 1913.

* Lejeunea grossiuscula Gottsche ex Steph., Sp. Hepat. (Stephani) 5: 739, 1915.

** Lejeunea hahnii Solari, J. Hattori Bot. Lab. 54: 543, 1983. Nom. nov. pro Microlejeunea grandistipula Steph., Hedwigia 35 (3): 114, 1896.

* Lejeunea haitica Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 263, 1843.

** Lejeunea hawaikiana M.A.M.Renner et de Lange, New Zealand J. Bot. 49 (3): 431, 2011. Nom. nov. pro Stenolejeunea acuminata R.M.Schust., J. Hattori Bot. Lab. 89: 156, 2000.

** Lejeunea helenae Pearson, Forh. Vidensk.-Selsk. Kristiania 1886 (3): 6, 1886.

** Lejeunea helmsiana (Steph.) Steph., Sp. Hepat. (Stephani) 5: 796, 1915. Bas.: Eulejeunea helmsiana Steph., Hedwigia 35 (3): 89, 1896.

*** Lejeunea hepaticola (Steph.) Steph., Sp. Hepat. (Stephani) 5: 714, 1915. Bas.: Eulejeunea hepaticola Steph., Hedwigia 27 (2): 60, 1888.

** Lejeunea hibernica Bischl., H.A.Mill. et Bonner ex Grolle, Lindbergia 3 (1/2): 48, 1975 [1976]. Based on: Lejeunea hibernica Bischl., H.A.Mill. et Bonner, Nova Hedwigia 3 (4): 455, 1961 [1962], nom. inval.

** Lejeunea hodgsoniana Grolle ex R.J.Lewington, Bever. et M.A.M.Renner, PhytoKeys 29: 2, 2013.

** Lejeunea howeana Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 219, 1982. Nom. nov. pro Cheilolejeunea wattsiana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 102, 1914.

** Lejeunea hui R.L.Zhu, Beih. Nova Hedwigia 121: 134, 2001.

** Lejeunea humefacta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 275, 1884.

** Lejeunea hyalina (Steph.) L.Söderstr. et A.Hagborg, Phytotaxa 220 (2): 188, 2015. Bas.: Pycnolejeunea hyalina Steph., Sp. Hepat. (Stephani) 5: 614, 1914.

** Lejeunea ibadana A.J.Harr. et E.W.Jones, J. Bryol. 12 (1): 40, 1982.

** Lejeunea immersa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 186, 1884.

** Lejeunea increscens Spruce, Mem. Torrey Bot. Club 1 (3): 124, 1890.

* Lejeunea inflatiloba (Steph.) H.A.Mill., Bonner et Bischl., Nova Hedwigia 14 (1): 67, 1967. Bas.: Microlejeunea inflatiloba Steph., Sp. Hepat. (Stephani) 6: 422, 1923.

*** Lejeunea inflexiloba Prantl, Hedwigia 31: xvi, 1892. Based on: Crossotolejeunea inflexiloba J.B.Jack et Steph., Hedwigia 31 (1): 16, 1892, nom. inval.

*** Lejeunea intricata Prantl, Hedwigia 31: xvi, 1892. Based on: Crossotolejeunea intricata J.B.Jack et Steph., Hedwigia 31 (1): 17, 1892, nom. inval.

*** Lejeunea isophylla E.W.Jones, Trans. Brit. Bryol. Soc. 5 (3): 559, 1968.

** Lejeunea japonica Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 203, 1891.

** Lejeunea jardinii Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: ccv, 1889 [1890].

** Lejeunea julacea Steph., Sp. Hepat. (Stephani) 5: 715, 1915.

*** Lejeunea jungneri (Steph.) Steph., Cat. Afr. Pl. (Hiern) 2 (2): 318, 1901. Bas.: Eulejeunea jungneri Steph., Hedwigia 35 (3): 90, 1896.

** Lejeunea kashyapii M.Dey, D.K.Singh et D.Singh, J. Bryol. 30 (2): 126, 2008.

* Lejeunea kilimandjarensis Steph., Sp. Hepat. (Stephani) 5: 716, 1915.

** Lejeunea kinabalensis Mizut., J. Hattori Bot. Lab. 33: 246, 1970.

** Lejeunea kodamae Ikegami et Inoue, J. Jap. Bot. 36 (1): 7, 1961.

** Lejeunea konosensis Mizut., J. Hattori Bot. Lab. 71: 127, 1992.

** Lejeunea kuerschneriana Pócs, Beih. Nova Hedwigia 138: 100, 2010.

*** Lejeunea laeta (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 380, 1845. Bas.: Jungermannia laeta Lehm. et Lindenb., Nov. Stirp. Pug. 6: 45, 1834.

** Lejeunea laii R.L.Zhu, J. Bryol. 30 (2): 173, 2008. Nom. nov. pro Microlejeunea ramulosa Herzog, J. Hattori Bot. Lab. 14: 51, 1955.

*** Lejeunea lamacerina (Steph.) Schiffn., Hedwigia 41 (5): 278, 1902. Bas.: Eulejeunea lamacerina Steph., Hedwigia 35 (3): 91, 1896. 317

** Lejeunea lamacerina subsp. gemminata R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (1): 168, 1957.

** Lejeunea latilobula (Herzog) R.L.Zhu et M.L.So, J. Bryol. 24 (2): 168, 2002. Bas.: Taxilejeunea latilobula Herzog, Symb. Sin. 5: 50, 1930.

** Lejeunea leptalea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 272, 1884.

* Lejeunea leratii (Steph.) Mizut., J. Hattori Bot. Lab. 33: 243, 1970. Bas.: Hygrolejeunea leratii Steph., Sp. Hepat. (Stephani) 5: 562, 1914. 318

** Lejeunea leucosis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxviii, 1889 [1890].

* Lejeunea litoralis Steph., Sp. Hepat. (Stephani) 5: 778, 1915.

** Lejeunea lomana E.W.Jones, Bull. Brit. Mus. (Nat. Hist.), Bot. 11 (3): 257, 1983.

** Lejeunea longicollis S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 12, 1958.

** Lejeunea longilobula Pócs, Beih. Nova Hedwigia 138: 112, 2010. Nom. nov. pro Lejeunea halei subsp. africana Pócs, J. Bryol. 29 (2): 89, 2007.

** Lejeunea lowriana Steph., Sp. Hepat. (Stephani) 5: 779, 1915.

*** Lejeunea lumbricoides (Nees) Nees, Syn. Hepat. 3: 342, 1845. Bas.: Jungermannia lumbricoides Nees, Enum. Pl. Crypt. Javae: 40, 1830.

** Lejeunea lunatigastria Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 98, 1974.

** Lejeunea lyratiflora Prantl, Hedwigia 31: xvi, 1892. Based on: Hygrolejeunea lyratiflora Steph., Hedwigia 31 (4): 169, 1892, nom. inval.

** Lejeunea magohukui Mizut., Misc. Bryol. Lichenol. 7 (7): 133, 1977.

** Lejeunea mandonii (Steph.) Müll.Frib., Leberm. Eur. 2 (9): 1281, 1958. Bas.: Microlejeunea mandonii Steph., Hedwigia 35 (3): 115, 1896.

*** Lejeunea masoalae Pócs, Beih. Nova Hedwigia 138: 103, 2010.

** Lejeunea massalongoana (Schiffn. ex P.Syd.) Solari, J. Hattori Bot. Lab. 54: 542, 1983.Bas.: Harpalejeunea massalongoana Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894.

** Lejeunea megalantha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 172, 1884.

** Lejeunea mehrana M.Dey, D.K.Singh et D.Singh, J. Bryol. 30 (2): 128, 2008.

*** Lejeunea meridensis Ilk.-Borg., Nova Hedwigia 80 (1/2): 59, 2005.

** Lejeunea micholitzii Mizut., J. Hattori Bot. Lab. 33: 236, 1970. Nom. nov. pro Hygrolejeunea parvisaccata Steph., Sp. Hepat. (Stephani) 5: 567, 1914.

** Lejeunea microloba Taylor, London J. Bot. 5: 399, 1846.

*** Lejeunea mimula Hürl., Bauhinia 11 (1): 12, 1993. Nom. nov. pro Hygrolejeunea luteola Steph., Sp. Hepat. (Stephani) 5: 553, 1914.

*** Lejeunea minutiloba A.Evans, Bull. Torrey Bot. Club 44 (11): 525, 1917.

** Lejeunea minutiloba var. heterogyna R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 425, 1957.

** Lejeunea mizutanii Grolle, J. Hattori Bot. Lab. 45: 178, 1979. Nom. nov. pro Cheilolejeunea zollingeri Steph., Hedwigia 34 (5): 245, 1895.

** Lejeunea molkenboeriana Sande Lac., Ned. Kruidk. Arch. 3: 421, 1854 [1855].

*** Lejeunea monimiae (Steph.) Steph., Sp. Hepat. (Stephani) 5: 747, 1915. Bas.: Eulejeunea monimiae Steph., Hedwigia 35 (3): 91, 1896.

*** Lejeunea multidentata M.E.Reiner et Mustelier, J. Bryol. 26 (2): 103, 2004 (Reiner-Drehwald and Mustelier 2004).

* Lejeunea musae (Spreng.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 407, 1845. Bas.: Jungermannia musae Spreng. Ann. Wetterauischen Ges. Gesammte Naturk. 1: 25, 1809.

** Lejeunea muscicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 281, 1884.

* Lejeunea muscicola var. palmicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 282, 1884.

** Lejeunea neelgherriana Gottsche, Syn. Hepat. 3: 354, 1845.

** Lejeunea nemoralis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 159 (65), 1864.

** Lejeunea nepalensis Steph., Sp. Hepat. (Stephani) 5: 780, 1915.

* Lejeunea nesiotica Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxviii, 1889 [1890].

** Lejeunea neumanniana Nees, Repert. Pharm. 76: 44, 1842.

** Lejeunea nietneri (Steph.) Steph. ex Watts, Proc. Linn. Soc. New South Wales (ser. 2) 26 (104): 633, 1902. Bas.: Eulejeunea nietneri Steph., Hedwigia 35 (3): 91, 1896.

** Lejeunea notata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 361, 1882.

** Lejeunea nymannii Steph., Sp. Hepat. (Stephani) 5: 781, 1915.

** Lejeunea obfusca Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 114, 1860 [1861].

** Lejeunea obidensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 277, 1884.

* Lejeunea obscura Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 112, 1860 [1861]. 319

** Lejeunea obtusata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 354, 1882.

** Lejeunea okomuensis E.W.Jones, Trans. Brit. Bryol. Soc. 5 (4): 787, 1969.

*** Lejeunea oligoclada Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcix, 1889 [1890].

*** Lejeunea oracola M.A.M.Renner, Austral. Syst. Bot. 23 (6): 448, 2010.

** Lejeunea osculatiana De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 233, 1857.

** Lejeunea otiana S.Hatt., Bot. Mag. (Tokyo) 65 (763/764): 15, 1952.

** Lejeunea ovalifolia Steph., Sp. Hepat. (Stephani) 5: 751, 1915.

** Lejeunea pacifica Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 262, 1843.

** Lejeunea pallescens Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 360, 1851.

** Lejeunea pallida Lindenb. et Gottsche, Syn. Hepat. 5: 762, 1847.

* Lejeunea pallidissima Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 27 (2/4): 247, 1920. 320

*** Lejeunea papilionacea Prantl, Hedwigia 31: xvii, 1892. Based on: Hygrolejeunea papilionacea Steph., Hedwigia 31 (4): 169, 1892, nom. inval.

* Lejeunea paratropa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 176, 1884. 321

** Lejeunea patagonica (Steph.) Steph., Sp. Hepat. (Stephani) 5: 797, 1915. Bas.: Eulejeunea patagonica Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 66, 1900.

** Lejeunea patens Lindb., Acta Soc. Sci. Fenn. 10: 482, 1875.

*** Lejeunea patersonii (Steph.) Steph., Sp. Hepat. (Stephani) 5: 784, 1915. Bas.: Eulejeunea patersonii Steph., Hedwigia 35 (3): 92, 1896.

*** Lejeunea patriciae Schäf.-Verw., Candollea 56 (1): 64, 2001. Nom. nov. pro Lejeunea pilifera Tixier, Gard. Bull. Singapore 25 (3): 351, 1971, nom. illeg.

*** Lejeunea paucidentata (Steph.) Grolle, J. Hattori Bot. Lab. 69: 191, 1991. Bas.: Crossotolejeunea paucidentata Steph., Hedwigia 35 (3): 76, 1896.

*** Lejeunea pectinella Mizut., J. Hattori Bot. Lab. 33: 239, 1970.

** Lejeunea perigonialis Gottsche, Mexik. Leverm.: 223, 1863.

*** Lejeunea perpapillosa M.E.Reiner et K.C.Pôrto, Nova Hedwigia 85 (3/4): 542, 2007 (Reiner-Drehwald and Pôrto 2007).

* Lejeunea pertusa (Corda ex Nees et Mont.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 407, 1845. Bas.: Jungermannia pertusa Corda ex Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 63, 1836.

*** Lejeunea phyllobola Nees et Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 471, 1842.

* Lejeunea phyllobola var. turgidula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 259, 1884.

*** Lejeunea planiloba A.Evans, Proc. Wash. Acad. Sci. 8: 147, 1906.

** Lejeunea polilloensis Steph., Sp. Hepat. (Stephani) 5: 786, 1915.

** Lejeunea praetervisa Steph., Sp. Hepat. (Stephani) 5: 752, 1915.

** Lejeunea primordialis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 375, 1845. Bas.: Jungermannia primordialis Hook.f. et Taylor, London J. Bot. 4: 92, 1845.

** Lejeunea princeps (Steph.) Mizut., J. Hattori Bot. Lab. 34: 454, 1971. Bas.: Hygrolejeunea princeps Steph., Sp. Hepat. (Stephani) 5: 568, 1914.

** Lejeunea procumbens Mitt., Fl. vit.: 413, 1871 [1873].

** Lejeunea propagulifera Gradst., Phytotaxa 9: 54, 2010. Nom. nov. pro Trachylejeunea schiffneri Herzog, Svensk Bot. Tidskr. 42 (3): 239, 1948.

** Lejeunea pteridis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxvii, 1889 [1890].

*** Lejeunea ptosimophylla C.Massal., Nuovo Giorn. Bot. Ital. 13 (2): 123, 1881.

*** Lejeunea puiggariana Steph., Sp. Hepat. (Stephani) 5: 754, 1915.

*** Lejeunea pulverulenta (Steph.) M.E.Reiner, Cryptog. Bryol. 26 (1): 60, 2005. Bas.: Taxilejeunea pulverulenta Steph., Sp. Hepat. (Stephani) 5: 477, 1914.

*** Lejeunea pulvinata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 61, 1836. Nom. nov. pro Jungermannia pulvinata Lehm. et Lindenb., Nov. Stirp. Pug. 5: 15, 1833, nom. illeg.

*** Lejeunea raddiana Lindenb., Syn. Hepat. 3: 342, 1845.

** Lejeunea radicans Lindenb. et Gottsche, Syn. Hepat. 5: 766, 1847.

** Lejeunea ramosissima Steph., Bot. Jahrb. Syst. 8 (2): 88, 1886.

** Lejeunea rara Steph., Sp. Hepat. (Stephani) 5: 798, 1915. 322

* Lejeunea ravenelii Austin, Bot. Bull. (Hanover) 1 (8): 35, 1876. 323

** Lejeunea recurva M.E.Reiner, Polish Bot. J. 58 (2): 423, 2013. Nom. nov. pro Hygrolejeunea herzogii Steph., Biblioth. Bot. 87 (2): 265, 1916, nom. illeg.

*** Lejeunea reflexistipula (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 335, 1845. Bas.: Jungermannia reflexistipula Lehm. et Lindenb., Nov. Stirp. Pug. 5: 10, 1833.

** Lejeunea reflexistipula var. costaricensis (Steph.) M.E.Reiner, Nova Hedwigia 81 (3/4): 408, 2005. Bas.: Hygrolejeunea costaricensis Steph., Hedwigia 35 (3): 100, 1896.

*** Lejeunea reinerae Ilk.-Borg., Nova Hedwigia 80 (1/2): 61, 2005. Nom. nov. pro Echinocolea herzogii Mizut. et Grolle, Bot. Mag. (Tokyo) 77 (915): 333, 1964.

** Lejeunea resupinata (Steph.) Steph., Sp. Hepat. (Stephani) 5: 757, 1915. Bas.: Eulejeunea resupinata Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 22, 1897.

** Lejeunea reticulata Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 747, 1942.

*** Lejeunea rhigophila M.A.M.Renner, Austral. Syst. Bot. 23 (6): 453, 2010.

** Lejeunea rhodesiae (Sim) R.M.Schust., J. Hattori Bot. Lab. 25: 71, 1962. Bas.: Stylolejeunea rhodesiae Sim, Trans. Roy. Soc. South Africa 15 (1): 68, 1926.

*** Lejeunea rionegrensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 579, 1885. Nom. nov. pro Lejeunea implexa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 240, 1884, nom. inval.

** Lejeunea riparia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 113, 1860 [1861].

* Lejeunea rothii (Schwägr.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 407, 1845. Bas.: Jungermannia rothii Schwägr., Hist. Musc. Hepat. Prodr.: 17, 1814.

*** Lejeunea rotundifolia Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 359, 1851.

** Lejeunea sanctae-helenae M.Wigginton, J. Bryol. 29 (1): 12, 2007.

** Lejeunea schusteri Grolle, Haussknechtia 8: 60, 2001. Nom. nov. pro Rectolejeunea denudata R.M.Schust., J. Hattori Bot. Lab. 89: 143, 2000.

** Lejeunea semiscabrida Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 154 (60), 1864.

** Lejeunea semperi Steph., Sp. Hepat. (Stephani) 5: 788, 1915.

** Lejeunea seriata Lindenb. et Gottsche, Syn. Hepat. 5: 762, 1847.

** Lejeunea sessiliflora (Steph.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 37: 171, 1988. Bas.: Macrolejeunea sessiliflora Steph., Sp. Hepat. (Stephani) 5: 512, 1914.

** Lejeunea setacea (Steph.) Steph., Sp. Hepat. (Stephani) 5: 719, 1914. Bas.: Eulejeunea setacea Steph., Bull. Mus. Natl. Hist. Nat. 18 (2): 120, 1912.

** Lejeunea setiloba Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 281, 1884.

** Lejeunea sharpii (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 991, 1980. Bas.: Taxilejeunea sharpii R.M.Schust., J. Elisha Mitchell Sci. Soc. 81 (1): 41, 1965.

** Lejeunea siccata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 284, 1884.

** Lejeunea silvatica Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 159 (65), 1864.

** Lejeunea smaragdina Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxii, 1889 [1890].

*** Lejeunea soae R.L.Zhu, Y.M.Wei, L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 81 (1): 1, 2013. Nom. nov. pro Trachylejeunea chinensis Herzog, Symb. Sin. 5: 49, 1930.

** Lejeunea solanicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 280, 1884.

*** Lejeunea sordida (Nees) Nees, Naturgesch. Eur. Leberm. 3: 278, 1838. Bas.: Jungermannia sordida Nees, Enum. Pl. Crypt. Javae: 41, 1830.

*** Lejeunea spiniloba Lindenb. et Gottsche, Syn. Hepat. 5: 770, 1847.

** Lejeunea spinuliflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 177, 1884.

** Lejeunea sporadica Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxx, 1889 [1890].

* Lejeunea squarrosa (Steph.) Steph., Sp. Hepat. (Stephani) 5: 719, 1915. Bas.: Eulejeunea squarrosa Steph., Bot. Jahrb. Syst. 20 (3): 317, 1895.

** Lejeunea squarrosula (Herzog) Solari, J. Hattori Bot. Lab. 54: 543, 1983. Bas.: Strepsilejeunea squarrosula Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 741, 1942.

* Lejeunea stephaniana Mizut., J. Hattori Bot. Lab. 27: 143, 1964. Nom. nov. pro Strepsilejeunea heterophylla Steph., Sp. Hepat. (Stephani) 6: 395, 1923. 324

** Lejeunea stevensiana (Steph.) Mizut., J. Hattori Bot. Lab. 34: 452, 1971. Bas.: Taxilejeunea stevensiana Steph., Hedwigia 35 (3): 136, 1896.

** Lejeunea subacuta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 113, 1860 [1861].

* Lejeunea subaquatica Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 397, 1955.

* Lejeunea subbifida Steph. ex Duss, Enum. musc. Antilles franç., Hép.: 10, 1903. Nom. nov. pro Odontolejeunea subbifida Steph., Symb. Antill. (Urban) 3 (2): 277, 1902, nom. illeg.

** Lejeunea subigiensis (Steph.) Steph., Hedwigia 35 (3): 94, 1896. Bas.: Eulejeunea subigiensis Steph., Hedwigia 35 (3): 94, 1896.

** Lejeunea subrufula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 289, 1884.

** Lejeunea subsessilis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 282, 1884.

*** Lejeunea subspathulata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 173, 1884.

** Lejeunea succulenta Herzog, Rev. Bryol. Lichénol. 20 (1/2): 171, 1951 [1952].

** Lejeunea suffruticola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 279, 1884.

** Lejeunea syoshii Inoue, Bull. Natl. Sci. Mus. Tokyo, B 3 (4): 143, 1977.

*** Lejeunea talamancensis M.E.Reiner et Schäf.-Verw., Nova Hedwigia 87 (3/4): 414, 2008.

*** Lejeunea tamasii M.E.Reiner, N.Salazar et C.Chung, Polish Bot. J. 58 (2): 420, 2013.

** Lejeunea tamaspocsii G.E.Lee, Polish Bot. J. 58 (1): 66, 2013.

*** Lejeunea tapajosensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 223, 1884.

** Lejeunea tarapotensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 282, 1884.

** Lejeunea tenella Taylor, London J. Bot. 5: 398, 1846.

** Lejeunea thallophora (Eifrig) Gradst., Phytotaxa 9: 54, 2010. Bas.: Taxilejeunea thallophora Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 99, 1937.

*** Lejeunea tonduzana (Steph.) M.E.Reiner, Polish Bot. J. 58 (2): 421, 2013. Bas.: Hygrolejeunea tonduzana Steph., Hedwigia 35 (3): 105, 1896.

** Lejeunea trachygona Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 175, 1884.

*** Lejeunea trinitensis Lindenb., Syn. Hepat. 3: 381, 1845.

** Lejeunea trukensis H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 56, 1963.

** Lejeunea tuberculosa Steph., Sp. Hepat. (Stephani) 5: 790, 1915.

** Lejeunea tumida Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 157, 1854.

** Lejeunea uleana (Steph.) Steph., Sp. Hepat. (Stephani) 5: 765, 1915. Bas.: Eulejeunea uleana Steph., Hedwigia 35 (3): 94, 1896.

*** Lejeunea umbilicata (Nees) Nees, Observ. bot.: 472, 1843. Bas.: Jungermannia umbilicata Nees, Enum. Pl. Crypt. Javae: 42, 1830.

** Lejeunea utriculata (Steph.) Mizut., J. Hattori Bot. Lab. 33: 242, 1970. Bas.: Pycnolejeunea utriculata Steph., Hedwigia 35 (3): 126, 1896.

** Lejeunea vesicata Mitt., Fl. vit.: 415, 1871 [1873].

*** Lejeunea villaumei (Steph.) Grolle, J. Bryol. 9 (4): 536, 1977 [1978]. Bas.: Otigoniolejeunea villaumei Steph., Sp. Hepat. (Stephani) 5: 516, 1914.

** Lejeunea vojtkoi Pócs, Beih. Nova Hedwigia 138: 107, 2010.

** Lejeunea vulgariformis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 355, 1882.

** Lejeunea wattsiana (Steph.) H.A.Mill., Bonner et Bischl., Nova Hedwigia 14 (1): 66, 1967. Bas.: Microlejeunea wattsiana Steph., Sp. Hepat. (Stephani) 5: 834, 1916.

* Lejeunea wichurae (Steph.) Steph., Sp. Hepat. (Stephani) 5: 792, 1915. Bas.: Eulejeunea wichurae Steph., Hedwigia 35 (3): 94, 1896.

** Lejeunea wightii Lindenb., Syn. Hepat. 3: 379, 1845.

*** Lejeunea xiphophylla (Herzog) M.E.Reiner, Nova Hedwigia 87 (3/4): 409, 2008. Bas.: Taxilejeunea xiphophylla Herzog, Svensk Bot. Tidskr. 46 (1): 99, 1952.

** subg. Nanolejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 1092, 1980.

** Lejeunea curviloba Steph., Sp. Hepat. (Stephani) 5: 774, 1915.

*** Lejeunea laetevirens Nees et Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 469, 1842.

** Lejeunea pallidevirens S.Hatt., J. Hattori Bot. Lab. 12: 80, 1954. Nom. nov. pro Microlejeunea rotundistipula var. pallida S.Hatt., J. Hattori Bot. Lab. 5: 53, 1951.

** Lejeunea parva (S.Hatt.) Mizut., Misc. Bryol. Lichenol. 5 (10/12): 178, 1971. Bas.: Microlejeunea rotundistipula f. parva S.Hatt., Bull. Tokyo Sci. Mus. 11: 123, 1944.

*** Lejeunea prionoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 238, 1884.

*** Lejeunea ramulosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 274, 1884.

*** subg. Neopotamolejeunea (M.E.Reiner) Gradst. et M.E.Reiner, Syst. Bot. 32 (3): 487, 2007. Bas.: Neopotamolejeunea M.E.Reiner, Nova Hedwigia 71 (3/4): 449, 2000.

** Lejeunea juruana Gradst. et M.E.Reiner, Syst. Bot. 32 (3): 488, 2007. Nom. nov. pro Potamolejeunea uleana Steph., Sp. Hepat. (Stephani) 5: 641, 1914.

*** Lejeunea polyantha Mont., Ann. Sci. Nat. Bot. (sér. 4) 5: 350, 1856.

** Lejeunea tenera (Sw.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 406, 1845. Bas.: Jungermannia tenera Sw., Prodr. (Swartz): 143, 1788.

*** Lejeunea topoensis Gradst. et M.E.Reiner, Syst. Bot. 32 (3): 488, 2007.

** subg. Papillolejeunea (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998. Bas.: Papillolejeunea Pócs, Trop. Bryol. 13: 2, 1997.

** Lejeunea balazsii (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998. Bas.: Papillolejeunea balazsii Pócs, Trop. Bryol. 13: 3, 1997.

** Lejeunea candida (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998. Bas.: Papillolejeunea candida Pócs, Trop. Bryol. 13: 8, 1997.

** Lejeunea falcata (Pócs et J.Eggers) Pócs, Phytotaxa 208 (1): 99, 2015. Bas.: Papillolejeunea falcata Pócs et J.Eggers, Bryobrothera 5: 163, 1999.

** Lejeunea koponenii (Pócs et J.Eggers) Pócs, Phytotaxa 208 (1): 99, 2015. Bas.: Papillolejeunea koponenii Pócs et J.Eggers, Bryobrothera 5: 159, 1999.

** Lejeunea papuana (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998. Bas.: Papillolejeunea papuana Pócs, Trop. Bryol. 13: 14, 1997.

** Lejeunea touwii (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998. Bas.: Papillolejeunea touwii Pócs, Trop. Bryol. 13: 11, 1997.

Incertae sedis

* Lejeunea apiahyna (Steph.) Sushil K.Singh, Phytotaxa 96 (1): 63, 2013. Bas.: Otigoniolejeunea apiahyna Steph., Sp. Hepat. (Stephani) 5: 514, 1914.

* Lejeunea aptycta Gottsche, Syn. Hepat. 3: 369, 1845.

** Lejeunea asthenica Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 222, 1884.

* Lejeunea bethanica Gottsche, Syn. Hepat. 3: 381, 1845 (Gottsche et al. 1945b)

** Lejeunea bombonasensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 222, 1884.

** Lejeunea bornmuelleri (Steph.) M.E.Reiner, Nova Hedwigia 95 (3/4): 471, 2012. Bas.: Rectolejeunea bornmuelleri Steph., Sp. Hepat. (Stephani) 5: 682, 1914.

** Lejeunea caracensis Lindenb., Syn. Hepat. 3: 355, 1845.

** Lejeunea caripensis Lindenb. et Gottsche, Syn. Hepat. 5: 758, 1847.

** Lejeunea chamissonis Lindenb., Syn. Hepat. 3: 378, 1845.

** Lejeunea chimborazensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 215, 1884.

*** Lejeunea combuensis O.S.Moura, Ilk.-Borg. et M.E.Reiner, Nova Hedwigia 95 (1/2): 198, 2012.

** Lejeunea concava Lindenb. et Gottsche, Syn. Hepat. 5: 759, 1847.

*** Lejeunea conformis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 260, 1843.

** Lejeunea cordistipula Lindenb. et Gottsche, Syn. Hepat. 5: 758, 1847.

** Lejeunea cyrtotis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 229, 1884.

*** Lejeunea debilis (Lehm. et Lindenb.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 60, 1836. Bas.: Jungermannia debilis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 51, 1832.

** Lejeunea devendrae (Sushil K.Singh) P.K.Verma et K.K.Rawat, J. Bryol. 36 (2): 161, 2014. Bas.: Rectolejeunea devendrae Sushil K.Singh, Indian J. Forest. 34 (3): 341, 2011.

** Lejeunea dictyocalyx Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 218, 1884.

** Lejeunea dipterota (Eifrig) G.E.Lee, Polish Bot. J. 58 (1): 61, 2013. Bas.: Taxilejeunea dipterota Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 96, 1937.

*** Lejeunea drehwaldii Heinrichs et Schäf.-Verw., Phytotaxa 69: 14, 2012. Nom. nov. pro Sphaerolejeunea umbilicata Herzog, Ann. Bryol. 11: 88, 1938.

** Lejeunea duncaniae (Sim) M.E.Reiner, Phytotaxa 208 (1): 98, 2015. Bas.: Stylolejeunea duncaniae Sim, Trans. Roy. Soc. South Africa 15 (1): 68, 1926.

** Lejeunea edentata L.Söderstr., Phytotaxa 208 (1): 98, 2015. Nom. nov. pro Cyclolejeunea marginata R.M.Schust., Phytologia 39 (6): 430, 1978.

* Lejeunea emarginuliflora Gottsche ex Steph., Sp. Hepat. (Stephani) 5: 734, 1915. 325

* Lejeunea epibrya Taylor, London J. Bot. 7: 199, 1848.

** Lejeunea estrellamontana M.A.M.Renner et Pócs, Phytotaxa 81 (1): 9, 2013. Nom. nov. pro Stenolejeunea fissistipula R.M.Schust., J. Hattori Bot. Lab. 89: 167, 2000.

* Lejeunea fawcettiae D.J.Carr, Proc. Roy. Soc. Victoria 117 (2): 325, 2005. Based on: Lejeunea fawcettiae D.J.Carr, Proc. Roy. Soc. Victoria 116 (2): 229, 2004, nom. inval.

** Lejeunea flaccida Lindenb. et Gottsche, Syn. Hepat. 5: 758, 1847.

** Lejeunea flavida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 113, 1860 [1861].

** Lejeunea florida Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 221, 1884.

** Lejeunea fulva Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 237, 1884.

** Lejeunea gottscheana Lindenb., Syn. Hepat. 3: 382, 1845.

** Lejeunea graminicolor Spruce, J. Linn. Soc., Bot. 30 (210): 343, 1895.

*** Lejeunea grolleana (Bernecker) R.L.Zhu et W.Ye, J. Syst. Evol. 51 (4): 472, 2013. Bas.: Oryzolejeunea grolleana Bernecker, Haussknechtia, Beih. 9: 37, 1999.

*** Lejeunea herminieri (Steph.) R.L.Zhu, Phytotaxa 208 (1): 99, 2015. Bas.: Archilejeunea herminieri Steph., Sp. Hepat. (Stephani) 4: 714, 1911.

** Lejeunea heterocheila Taylor, London J. Bot. 5: 394, 1846.

* Lejeunea hygrophila Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 157, 1864.

** Lejeunea laevicalyx Gottsche, Mexik. Leverm.: 221, 1863.

* Lejeunea laxa (Nees) Lindenb., Syn. Hepat. 3: 378, 1845. Bas.: Jungermannia thymifolia δ laxa Nees, Enum. Pl. Crypt. Javae: 43, 1830.

** Lejeunea leiantha Spruce, J. Linn. Soc., Bot. 30 (210): 345, 1895.

** Lejeunea leptoscypha Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: ccv, 1889 [1890].

*** Lejeunea lusoria (Lindenb. et Gottsche) Steph., Hedwigia 29 (3): 141, 1890. Bas.: Omphalanthus lusorius Lindenb. et Gottsche, Syn. Hepat. 5: 747, 1847.

** Lejeunea luzonensis (Steph.) R.L.Zhu et M.J.Lai, Ann. Bot. Fenn. 48 (5): 376, 2011. Bas.: Taxilejeunea luzonensis Steph., Hedwigia 35 (3): 134, 1896.

** Lejeunea macrorhyncha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 220, 1884.

** Lejeunea malangensis (Herzog) R.L.Zhu et Y.M.Wei, J. Bryol. 34 (4): 319, 2012. Bas.: Trachylejeunea malangensis Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 203, 1931.

** Lejeunea marasmodes Spruce, Mem. Torrey Bot. Club 1 (3): 125, 1890.

** Lejeunea morobensis (Grolle) M.A.M.Renner et Pócs, Phytotaxa 81 (1): 9, 2013. Bas.: Stenolejeunea morobensis Grolle, J. Hattori Bot. Lab. 29: 76, 1966.

** Lejeunea novoguineensis Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 238, 1893.

*** Lejeunea obtusangula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 221, 1884.

* Lejeunea oerstediana Lindenb. et Hampe, Linnaea 24 (6): 641, 1851 [1852]. 326

** Lejeunea paraensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 224, 1884.

** Lejeunea parviloba Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 87, 1876 [1877].

* Lejeunea pfleidereri Sushil K.Singh, Phytotaxa 96 (1): 63, 2013. Nom. nov. pro Otigoniolejeunea indica Steph., Sp. Hepat. (Stephani) 6: 408, 1923.

* Lejeunea prominula Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 157, 1864.

*** Lejeunea pterigonia (Lehm. et Lindenb.) Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 337, 1840. Bas.: Jungermannia pterigonia Lehm. et Lindenb., Nov. Stirp. Pug. 6: 44, 1834.

** Lejeunea quinqueumbonata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 230, 1884.

** Lejeunea quinqueumbonata var. rotundata (Herzog) Sushil K.Singh, Phytotaxa 96 (1): 64, 2013. Bas.: Otigoniolejeunea quinqueumbonata var. rotundata Herzog, Rev. Bryol. Lichénol. 20 (1/2): 154, 1951 [1952].

** Lejeunea remotifolia Hampe ex Steph., Sp. Hepat. (Stephani) 5: 756, 1915.

*** Lejeunea ruthii (A.Evans) R.M.Schust., J. Hattori Bot. Lab. 25: 23, 1962. Bas.: Microlejeunea ruthii A.Evans, Mem. Torrey Bot. Club 8 (2): 161, 1902.

** Lejeunea ruthii var. alata R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 1063, 1980.

** Lejeunea sikorae (Steph.) Steph., Bull. Soc. Roy. Bot. Belgique, Mém. 32: 120, 1893 [1894]. Bas.: Taxilejeunea sikorae Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 34, 1893 [1894].

** Lejeunea srivastavae P.K.Verma et K.K.Rawat, Taiwania 58 (1): 8, 2013.

** Lejeunea stenodentata M.A.M.Renner et Pócs, Phytotaxa 81 (1): 8, 2013. Nom. nov. pro Drepanolejeunea dentata Steph., Hedwigia 35 (3): 82, 1896.

*** Lejeunea subelobata Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1039, 1888.

** Lejeunea subolivacea Mizut., J. Hattori Bot. Lab. 28: 121, 1965. Nom. nov. pro Eulejeunea olivacea Steph., Hedwigia 29 (2): 85, 1890.

** Lejeunea subplana (Steph.) C.J.Bastos, J. Bryol. 36 (3): 249, 2014. Bas.: Trachylejeunea subplana Steph., Sp. Hepat. (Stephani) 5: 310, 1913.

*** Lejeunea sulphurea (Lehm. et Lindenb.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 217, 1884. Bas.: Jungermannia sulphurea Lehm. et Lindenb., Nov. Stirp. Pug. 5: 14, 1833.

** Lejeunea terricola Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxci, 1889 [1890].

** Lejeunea urbanii (Steph.) Steph., Sp. Hepat. (Stephani) 5: 766, 1915. Bas.: Eulejeunea urbanii Steph., Hedwigia 27 (11/12): 301, 1888.

** Lejeunea venezuelana (R.M.Schust.) R.L.Zhu et W.Ye, J. Syst. Evol. 51 (4): 473, 2013. Bas.: Cyrtolejeunea venezuelana R.M.Schust., Phytologia 39 (6): 426, 1978.

** Lejeunea viridis R.M.Schust. ex L.Söderstr. et A.Hagborg, Phytotaxa 208 (1): 99, 2015. Based on: Prionocolea viridissima R.M.Schust., J. Hattori Bot. Lab. 75: 215, 1994, nom. inval.

* Lejeunea zacuapana (Steph.) Prantl, Hedwigia 29: xviii, 1890. Bas.: Eulejeunea zacuapana Steph., Hedwigia 29 (2): 87, 1890.

Excluded from the genus

* Lejeunea elegans Gottsche, Syn. Hepat. 3: 364, 1845. 327

* Lejeunea hieronymii Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cciii, 1889 [1890]. 328

* Lejeunea proboscidea Gottsche, Mexik. Leverm.: 225, 1863. 329

* Lejeunea scabriflora Loitl., Diagn. pl. nov.: 22, 1894. 330

* Lejeunea trochantha Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcii, 1889 [1890]. 331

** Microlejeunea (Spruce) Steph., Hedwigia 27 (2): 61, 1888. Bas.: Lejeunea subg. Microlejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 286, 1884.

*** Microlejeunea acutifolia Steph., Hedwigia 35 (3): 113, 1896.

** Microlejeunea africana Steph., Hedwigia 27 (2): 61, 1888.

** Microlejeunea aligera (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 827, 1915. Bas.: Lejeunea aligera Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 113, 1860 [1861].

** Microlejeunea ankasica E.W.Jones, J. Bryol. 10 (4): 394, 1979.

** Microlejeunea aphanella (Spruce) Steph., Sp. Hepat. (Stephani) 5: 816, 1915. Bas.: Lejeunea aphanella Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 290, 1884.

** Microlejeunea atsuana Steph., Hedwigia 35 (3): 113, 1896.

** Microlejeunea bischlerae (B.M.Thiers) B.M.Thiers, Phytotaxa 65: 59, 2012. Bas.: Lejeunea bischlerae B.M.Thiers, Cryptog. Bryol. Lichénol. 18 (3): 223, 1997.

*** Microlejeunea bullata (Taylor) Steph., Hedwigia 29 (2): 90, 1890. Bas.: Lejeunea bullata Taylor, London J. Bot. 5: 398, 1846.

* Microlejeunea byssoides (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892. Bas.: Lejeunea byssoides Gottsche, Abh. Naturwiss. Vereins Bremen 7: 358, 1882.

** Microlejeunea capillaris (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 819, 1915. Bas.: Lejeunea capillaris Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 163, 1864.

** Microlejeunea cochlearifolia Steph., Hedwigia 27 (3/4): 113, 1888. 332

** Microlejeunea colombiana Bischl., Nova Hedwigia 5 (1/2): 373, 1963.

** Microlejeunea constricta (Grolle) Grolle, J. Bryol. 21 (1): 41, 1999. Bas.: Harpalejeunea constricta Grolle, Nova Hedwigia 16: 150, 1968.

** Microlejeunea crenulifolia (Gottsche) Steph., Hedwigia 35 (3): 114, 1896. Bas.: Lejeunea crenulifolia Gottsche, Mexik. Leverm.: 227, 1863.

*** Microlejeunea cystifera Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 68, 1950.

** Microlejeunea dispar Jovet-Ast, Rev. Bryol. Lichénol. 27 (3/4): 191, 1959.

*** Microlejeunea epiphylla Bischl., Nova Hedwigia 5 (1/2): 378, 1963.

*** Microlejeunea filicuspis (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 52, 2013. Bas.: Drepanolejeunea filicuspis Steph., Sp. Hepat. (Stephani) 5: 344, 1913.

*** Microlejeunea fischeri (Tixier) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 52, 2013. Bas.: Harpalejeunea fischeri Tixier, Trop. Bryol. 11: 29, 1995.

** Microlejeunea fissistipula Steph., Sp. Hepat. (Stephani) 5: 810, 1915.

*** Microlejeunea globosa (Spruce) Steph., Sp. Hepat. (Stephani) 5: 821, 1915. Bas.: Lejeunea globosa Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxciii, 1889 [1890].

** Microlejeunea herzogiana Steph., Biblioth. Bot. 87 (2): 266, 1916.

** Microlejeunea indica (Udar et U.S.Awasthi) Y.M.Wei et R.L.Zhu, Phytotaxa 97 (2): 63, 2013. Bas.: Lejeunea indica Udar et U.S.Awasthi, Cryptog. Bryol. Lichénol. 2 (3): 345, 1981.

** Microlejeunea inflata Steph., Sp. Hepat. (Stephani) 5: 811, 1915.

** Microlejeunea kinabaluensis (Mizut.) Grolle, J. Bryol. 21 (1): 42, 1999. Bas.: Harpalejeunea kinabaluensis Mizut., J. Hattori Bot. Lab. 37: 200, 1973.

*** Microlejeunea latitans (Hook.f. et Taylor) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 52, 2013. Bas.: Jungermannia latitans Hook.f. et Taylor, London J. Bot. 3: 399, 1844.

** Microlejeunea lunulatiloba Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 27, 1931. 333

* Microlejeunea magnilobula Gola, Ann. Bot. (Rome) 6 (2): 274, 1907.

** Microlejeunea mammillosa (Mizut.) Grolle, J. Bryol. 21 (1): 42, 1999. Bas.: Harpalejeunea mammillosa Mizut., J. Hattori Bot. Lab. 37: 192, 1973.

** Microlejeunea minutissima (Mizut.) Grolle, J. Bryol. 21 (1): 42, 1999. Bas.: Harpalejeunea minutissima Mizut., J. Hattori Bot. Lab. 37: 202, 1973.

* Microlejeunea minutistipula Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907–08), Bot. 2: 131, 1911. 334

*** Microlejeunea moniliata (Mizut.) R.L.Zhu et Y.M.Wei, Cryptog. Bryol. 34 (3): 308, 2013. Bas.: Lejeunea moniliata Mizut., J. Hattori Bot. Lab. 46: 357, 1979.

* Microlejeunea nepalensis Steph., Sp. Hepat. (Stephani) 5: 832, 1915.

** Microlejeunea nyandaruensis Pócs, Polish Bot. J. 47 (1): 14, 2002.

** Microlejeunea oblongistipula (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892. Bas.: Lejeunea oblongistipula Gottsche, Abh. Naturwiss. Vereins Bremen 7: 357, 1882.

** Microlejeunea ocellata (Herzog) Grolle, Haussknechtia 8: 60, 2001. Bas.: Rectolejeunea ocellata Herzog, Trans. & Proc. Roy. Soc. New Zealand 77 (2): 255, 1949.

* Microlejeunea ovistipula Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 131, 1911. 335

** Microlejeunea papulosa (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892. Bas.: Lejeunea papulosa Gottsche, Abh. Naturwiss. Vereins Bremen 7: 358, 1882.

** Microlejeunea perpusilla (Spruce) Steph., Hedwigia 35 (3): 113, 1896. Bas.: Lejeunea perpusilla Spruce, J. Bot. 19: 36, 1881.

** Microlejeunea punctiformis (Taylor) Steph., Hedwigia 29 (2): 90, 1890. Bas.: Lejeunea punctiformis Taylor, London J. Bot. 5: 398, 1846. 336

** Microlejeunea pyriformis (Lindenb. et Gottsche) Steph., Sp. Hepat. (Stephani) 5: 824, 1915. Bas.: Lejeunea pyriformis Lindenb. et Gottsche, Syn. Hepat. 5: 767, 1847.

** Microlejeunea spinosa (Mizut.) Grolle, J. Bryol. 21 (1): 43, 1999. Bas.: Harpalejeunea spinosa Mizut., J. Hattori Bot. Lab. 37: 196, 1973.

*** Microlejeunea squarrosa (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 51, 2013. Bas.: Strepsilejeunea squarrosa Steph., Hedwigia 35 (3): 130, 1896.

** Microlejeunea strasbergii Bardat et Ah-Peng, Bryologist 114 (4): 669, 2011.

*** Microlejeunea subulistipa Steph., Hedwigia 35 (3): 115, 1896.

* Microlejeunea szechuanensis P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 46, 1955. 337

** Microlejeunea udarii P.K.Verma et S.C.Srivast., J. Bombay Nat. Hist. Soc. 108 (2): 122, 2011 [2012].

*** Microlejeunea ulicina (Taylor) Steph., Hedwigia 29 (2): 88, 1890. Bas.: Jungermannia ulicina Taylor, Trans. Bot. Soc. Edinburgh 1 (1/4): 115, 1844.

* Microlejeunea usambarensis Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 730, 1913.

** Microlejeunea valenciana Steph., Sp. Hepat. (Stephani) 5: 815, 1915.

* Microlejeunea victoriensis D.J.Carr, Proc. Roy. Soc. Victoria 117 (2): 322, 2005.

*** Microlejeunea wallichiana (Lehm.) R.L.Zhu et Y.M.Wei, Cryptog. Bryol. 34 (3): 308, 2013. Bas.: Jungermannia wallichiana Lehm., Nov. Stirp. Pug. 3: 5, 1831.

* Taxilejeunea (Spruce) Steph., Hedwigia 28 (4): 262, 1889 nom. rejic. Bas.: Lejeunea subg. Taxilejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 212, 1884.

** Taxilejeunea acutifolia (Steph.) Steph., Sp. Hepat. (Stephani) 5: 481, 1914. Bas.: Dicranolejeunea acutifolia Steph., Hedwigia 35 (3): 76, 1896.

** Taxilejeunea antillana Steph., Sp. Hepat. (Stephani) 5: 482, 1914.

** Taxilejeunea apiculata (Gottsche) J.B.Jack et Steph., Hedwigia 31 (1): 13, 1892. Bas.: Omphalanthus apiculatus Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 149, 1864.

** Taxilejeunea argentina Steph., Sp. Hepat. (Stephani) 5: 482, 1914.

** Taxilejeunea arsenii Steph., Sp. Hepat. (Stephani) 6: 400, 1923.

** Taxilejeunea auriculata Steph., Sp. Hepat. (Stephani) 5: 459, 1914.

** Taxilejeunea berteroana Steph., Sp. Hepat. (Stephani) 5: 483, 1914.

** Taxilejeunea beyrichiana Steph., Sp. Hepat. (Stephani) 5: 460, 1914.

** Taxilejeunea biapiculata Steph., Sp. Hepat. (Stephani) 5: 460, 1914.

* Taxilejeunea boliviana Steph., Biblioth. Bot. 87 (2): 260, 1916.

** Taxilejeunea brasiliensis Steph., Hedwigia 35 (3): 132, 1896.

** Taxilejeunea coilantha Herzog, Rev. Bryol. Lichénol. 20 (1/2): 159, 1951 [1952].

* Taxilejeunea compressiuscula Steph., Sp. Hepat. (Stephani) 5: 501, 1914.

* Taxilejeunea convoluta Herzog, Biblioth. Bot. 88: 31, 1920 [1921].

* Taxilejeunea cuervi (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 486, 1914. Bas.: Omphalanthus cuervi Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 147, 1864.

* Taxilejeunea cuneistipula Steph., Sp. Hepat. (Stephani) 6: 401, 1923.

* Taxilejeunea cuspidata Steph., Biblioth. Bot. 87 (2): 260, 1916.

** Taxilejeunea decurrens Steph., Sp. Hepat. (Stephani) 5: 487, 1914.

** Taxilejeunea deflexa Steph., Sp. Hepat. (Stephani) 5: 502, 1914.

** Taxilejeunea densiflora A.Evans, Bull. Torrey Bot. Club 48 (4): 121, 1921.

** Taxilejeunea diaphana (Lehm.) Steph., Sp. Hepat. (Stephani) 5: 463, 1914. Bas.: Omphalanthus diaphanus Lehm., Nov. Stirp. Pug. 10: 12, 1857.

* Taxilejeunea dissitifolia Steph., Symb. Antill. 2: 472, 1901.

** Taxilejeunea eggersiana Schiffn., Bot. Jahrb. Syst. 23 (5): 579, 1897. Based on: Taxilejeunea eggersiana Steph., Hedwigia 27 (11/12): 285, 1888, nom. inval.

* Taxilejeunea elobulata Sim, Trans. Roy. Soc. South Africa 15 (1): 66, 1926.

** Taxilejeunea fissistipula Steph., Sp. Hepat. (Stephani) 5: 488, 1914.

** Taxilejeunea foliicola Steph., Sp. Hepat. (Stephani) 5: 466, 1914.

** Taxilejeunea furcicornuta Grolle, J. Bryol. 8 (1): 93, 1974.

** Taxilejeunea fuscorufa Steph., Hedwigia 35 (3): 133, 1896.

** Taxilejeunea galapagensis Onr., Misc. Bryol. Lichenol. 9 (6): 117, 1982.

* Taxilejeunea ghatensis P.K.Verma et S.C.Srivast., Proc. Natl. Acad. Sci. India, B 77 (2): 211, 2007.

* Taxilejeunea giulianettii Steph., Sp. Hepat. (Stephani) 5: 502, 1914.

** Taxilejeunea gomphocalyx Herzog, Beih. Bot. Centralbl. 61B (3): 580, 1942.

* Taxilejeunea grandifolia Steph., Biblioth. Bot. 87 (2): 261, 1916.

* Taxilejeunea grandistipula Steph., Sp. Hepat. (Stephani) 5: 504, 1914.

* Taxilejeunea hamatifolia Steph., Biblioth. Bot. 87 (2): 261, 1916.

** Taxilejeunea himalayensis Herzog, Ann. Bryol. 12: 86, 1939.

* Taxilejeunea immersa Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 96, 1937.

** Taxilejeunea irregularis Steph., Sp. Hepat. (Stephani) 5: 490, 1914.

** Taxilejeunea jamaicensis A.Evans, Bull. Torrey Bot. Club 48 (4): 117, 1921.

** Taxilejeunea jeringii Steph., Hedwigia 35 (3): 134, 1896.

** Taxilejeunea killipii Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 190, 1955.

* Taxilejeunea laevis (Gottsche) Steph., Hedwigia 44 (4): 229, 1905. Bas.: Omphalanthus laevis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 148, 1864.

* Taxilejeunea langiana Pearson, Ann. Bryol. 4: 102, 1931.

** Taxilejeunea lindbergiana Steph., Sp. Hepat. (Stephani) 5: 491, 1914.

** Taxilejeunea linguifolia Steph., Sp. Hepat. (Stephani) 5: 471, 1914.

* Taxilejeunea maxima Steph., Sp. Hepat. (Stephani) 5: 473, 1914.

** Taxilejeunea mexicana Steph., Sp. Hepat. (Stephani) 6: 402, 1923.

* Taxilejeunea microstipula Steph., Sp. Hepat. (Stephani) 5: 493, 1914.

* Taxilejeunea mucronata Steph., Sp. Hepat. (Stephani) 5: 473, 1914.

** Taxilejeunea multiflora Steph., Hedwigia 35 (3): 135, 1896.

* Taxilejeunea muscicola Steph., Biblioth. Bot. 87 (2): 263, 1916.

* Taxilejeunea nilgiriensis P.K.Verma et S.C.Srivast., Proc. Natl. Acad. Sci. India, B 77 (2): 207, 2007.

** Taxilejeunea nymannii Steph., Sp. Hepat. (Stephani) 5: 507, 1914.

** Taxilejeunea obtusifolia Steph., Sp. Hepat. (Stephani) 5: 474, 1914.

** Taxilejeunea papuliflora Steph., Sp. Hepat. (Stephani) 5: 494, 1914.

** Taxilejeunea parvibracteata Steph., Hedwigia 35 (3): 136, 1896.

** Taxilejeunea parvistipula Steph., Sp. Hepat. (Stephani) 6: 403, 1923.

* Taxilejeunea paucidens Steph., Biblioth. Bot. 87 (2): 263, 1916.

* Taxilejeunea pendula Steph., Biblioth. Bot. 87 (2): 263, 1916.

* Taxilejeunea peruviana Steph., Sp. Hepat. (Stephani) 5: 475, 1914.

** Taxilejeunea planilobula Herzog, Rev. Bryol. Lichénol. 20 (1/2): 160, 1951 [1952].

** Taxilejeunea pulchriflora Pearson, Ark. Bot. 19 (5): 15, 1924.

* Taxilejeunea pusilla Steph., Biblioth. Bot. 87 (2): 264, 1916.

** Taxilejeunea renistipula (Lindenb.) Steph., Hedwigia 29 (3): 142, 1890. Bas.: Omphalanthus renistipulus Lindenb., Syn. Hepat. 3: 308, 1845.

* Taxilejeunea rufescens Steph., Biblioth. Bot. 87 (2): 264, 1916.

*** Taxilejeunea serpillifolioides (Raddi) D.P.Costa, J. Bryol. 31 (4): 230, 2009. Bas.: Jungermannia serpillifolioides Raddi, Critt. Brasil.: 17, 1822.

** Taxilejeunea setchellii Pearson, Amer. Samoa: 146, 1924.

** Taxilejeunea speciosa Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 188, 1955.

* Taxilejeunea splendida Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 88, 1937.

** Taxilejeunea stephanii Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 90, 1937. Nom. nov. pro Hygrolejeunea nymanii Steph., Sp. Hepat. (Stephani) 5: 564, 1914.

** Taxilejeunea steyermarkii H.Rob., Phytologia 34 (1): 67, 1976.

** Taxilejeunea surinamensis (Lindenb. et Gottsche) Steph., Hedwigia 29 (3): 142, 1890. Bas.: Omphalanthus surinamensis Lindenb. et Gottsche, Linnaea 24 (6): 628, 1851 [1852].

* Taxilejeunea suringarii Steph., Sp. Hepat. (Stephani) 5: 479, 1914.

** Taxilejeunea tenerrima Steph., Sp. Hepat. (Stephani) 6: 406, 1923.

** Taxilejeunea tenuiplica Steph., Sp. Hepat. (Stephani) 5: 480, 1914.

* Taxilejeunea tjibodensis (Steph.) Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 91, 1937. Bas.: Hygrolejeunea tjibodensis Steph., Sp. Hepat. (Stephani) 5: 571, 1914.

** Taxilejeunea tonduzana Steph., Sp. Hepat. (Stephani) 5: 498, 1914.

* Taxilejeunea uleana Steph., Hedwigia 35 (3): 136, 1896.

* Taxilejeunea umbonata Steph., Sp. Hepat. (Stephani) 5: 481, 1914.

* Taxilejeunea urbanii Steph., Symb. Antill. (Urban) 3 (2): 278, 1902.

** Taxilejeunea vallis-gratiae Steph., Hedwigia 35 (3): 137, 1896.

** subtrib. Lepidolejeuneinae Gradst.

*** Lepidolejeunea R.M.Schust., Beih. Nova Hedwigia 9: 139, 1963. 338

** subg. Kingiolejeunea (H.Rob.) R.M.Schust., Phytologia 45 (5): 425, 1980. Bas.: Kingiolejeunea H.Rob., Bryologist 70 (1): 53, 1967.

** Lepidolejeunea auriculata Schäf.-Verw. et Heinrichs, Taxon 64 (2): 224, 2015.

*** Lepidolejeunea cordifissa (Taylor) M.E.Reiner, Nova Hedwigia 83 (3/4): 478, 2006. Bas.: Lejeunea cordifissa Taylor, London J. Bot. 5: 395, 1846.

*** Lepidolejeunea grossepapulosa (Steph.) Piippo, Acta Bot. Fenn. 132: 49, 1986. Bas.: Prionolejeunea grossepapulosa Steph., Sp. Hepat. (Stephani) 5: 220, 1913.

*** Lepidolejeunea involuta (Gottsche) Grolle, J. Hattori Bot. Lab. 55: 504, 1984. Bas.: Lejeunea involuta Gottsche, Syn. Hepat. 3: 350, 1845.

** subg. Lepidolejeunea

*** Lepidolejeunea bidentula (Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980. Bas.: Pycnolejeunea bidentula Steph., Bot. Centralbl. 60 (4): 107, 1894.

*** Lepidolejeunea bidentula var. novae-caledoniae Piippo, Acta Bot. Fenn. 132: 26, 1986.

*** Lepidolejeunea falcata (Herzog) R.M.Schust., Beih. Nova Hedwigia 9: 139, 1963. Bas.: Hygrolejeunea falcata Herzog, Ark. Bot. (n.ser.) 3 (3): 57, 1953.

*** Lepidolejeunea graeffei (J.B.Jack et Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980. Bas.: Archilejeunea graeffei J.B.Jack et Steph., Bot. Centralbl. 60 (4): 104, 1894.

*** Lepidolejeunea integristipula (J.B.Jack et Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980. Bas.: Pycnolejeunea integristipula J.B.Jack et Steph., Bot. Centralbl. 60 (4): 107, 1894.

** subg. Perilejeunea (Kachroo et R.M.Schust.) R.M.Schust., Phytologia 45 (5): 424, 1980. Bas.: Pycnolejeunea subg. Perilejeunea Kachroo et R.M.Schust., J. Linn. Soc., Bot. 56 (368): 493, 1961.

** Lepidolejeunea cuspidata (Gottsche) Heinrichs et Schäf.-Verw., Taxon 64 (2): 224, 2015. Bas.: Lejeunea cuspidata Gottsche, Syn. Hepat. 3: 351, 1845.

*** Lepidolejeunea delessertii (Nees et Mont.) Grolle, J. Hattori Bot. Lab. 55: 505, 1984. Bas.: Lejeunea delessertii Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 260, 1843.

*** Lepidolejeunea eluta (Nees) R.M.Schust., Beih. Nova Hedwigia 9: 139, 1963. Bas.: Jungermannia eluta Nees, Fl. Bras. (Martius) 1 (1): 362, 1833.

*** Lepidolejeunea serrulata (Steph.) Grolle, J. Hattori Bot. Lab. 55: 505, 1984. Bas.: Trachylejeunea serrulata Steph., Sp. Hepat. (Stephani) 5: 300, 1913.

*** Lepidolejeunea sullivantii (Gottsche) M.E.Reiner, Nova Hedwigia 83 (3/4): 479, 2006. Bas.: Lejeunea sullivantii Gottsche, Mexik. Leverm.: 196, 1863.

Incertae sedis

*** Lepidolejeunea borneensis (Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980. Bas.: Hygrolejeunea borneensis Steph., Sp. Hepat. (Stephani) 5: 557, 1914.

*** Lepidolejeunea longilobula (Amakawa) Xiao L.He, Acta Bot. Fenn. 163: 59, 1999. Bas.: Pycnolejeunea longilobula Amakawa, J. Jap. Bot. 40 (10): 307, 1965.

*** Otolejeunea Grolle et Tixier, Nova Hedwigia 32: 609, 1980.

*** subg. Allorgella (Tixier) Grolle, Haussknechtia 2: 53, 1985. Bas.: Allorgella Tixier, Nova Hedwigia 32: 612, 1980.

*** Otolejeunea australiensis B.M.Thiers, Brittonia 44 (2): 162, 1992.

*** Otolejeunea hoana (Tixier) Grolle, Haussknechtia 2: 54, 1985 [1986]. Bas.: Allorgella hoana Tixier, Nova Hedwigia 32: 615, 1980.

*** Otolejeunea rabenorii Tixier, Nova Hedwigia 46 (3/4): 376, 1988.

*** Otolejeunea schmidii (Tixier) Grolle, Haussknechtia 2: 54, 1985 [1986]. Bas.: Allorgella schmidii Tixier, Nova Hedwigia 32: 613, 1980.

*** Otolejeunea schnellii (Tixier) R.L.Zhu et M.L.So, Ann. Bot. Fenn. 34 (4): 287, 1997. Bas.: Allorgella schnellii Tixier, Cryptog. Bryol. Lichenol. 16 (3): 230, 1995.

*** Otolejeunea semperiana (Steph.) Grolle, Haussknechtia 2: 53, 1985 [1986]. Bas.: Prionolejeunea semperiana Steph., Sp. Hepat. (Stephani) 5: 227, 1913.

** Otolejeunea subana Pocs, Acta Acad. Ped. Agr., Sect. Biol. 25: 50, 2004 (Pocs 2004).

*** Otolejeunea zantenii Grolle, Haussknechtia 2: 54, 1985 [1986].

*** subg. Otolejeunea

*** Otolejeunea moniliata Grolle, Nova Hedwigia 32: 609, 1980. World checklist of hornworts and liverworts 353

*** subg. Phoxolejeunea Grolle, Haussknechtia 2: 49, 1985.

** Otolejeunea philippinensis R.L.Zhu et M.L.So, Syst. Bot. 23 (2): 231, 1998.

*** Otolejeunea streimannii Grolle, Haussknechtia 2: 49, 1985 [1986].

*** Rectolejeunea A.Evans, Bull. Torrey Bot. Club 33 (1): 8, 1906.

* Rectolejeunea colombiana R.M.Schust., J. Hattori Bot. Lab. 89: 146, 2000.

*** Rectolejeunea emarginuliflora (Schiffn.) A.Evans, Bull. Torrey Bot. Club 33 (1): 14, 1906. Bas.: Cheilolejeunea emarginuliflora Schiffn., Bot. Jahrb. Syst. 23 (5): 585, 1897.

*** Rectolejeunea flagelliformis A.Evans, Bull. Torrey Bot. Club 33 (1): 9, 1906.

* Rectolejeunea flagelliformis subsp. hamata R.M.Schust., J. Hattori Bot. Lab. 89: 137, 2000.

*** Rectolejeunea queenslandica (B.M.Thiers) Xiao L.He, Ann. Bot. Fenn. 34 (2): 129, 1997. Bas.: Lepidolejeunea queenslandica B.M.Thiers, Mem. New York Bot. Gard. 45: 556, 1987.

** Rectolejeunea truncatilobula C.J.Bastos, J. Bryol. 34 (2): 144, 2012.

*** Rectolejeunea versifolia (Schiffn.) L.Söderstr. et A.Hagborg, Phytotaxa 220 (2): 188, 2015. Bas.: Cheilolejeunea versifolia Schiffn., Bot. Jahrb. Syst. 23 (5): 597, 1897.

Excluded from the genus

* Rectolejeunea lindenbergii Steph., Sp. Hepat. (Stephani) 5: 689, 1914. 339

* Rectolejeunea lindigiana Steph., Sp. Hepat. (Stephani) 5: 690, 1914. 340

* Rectolejeunea pachyderma R.M.Schust., J. Hattori Bot. Lab. 89: 148, 2000. 341

** subtrib. Leptolejeuneinae Heinrichs et Schäf.-Verw.

*** Leptolejeunea (Spruce) Steph., Hedwigia 30 (6): 270, 1891. Bas.: Lejeunea subg. Leptolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 193, 1884.

*** Leptolejeunea amphiophthalma Zwickel, Ann. Bryol. 6: 117, 1933.

** Leptolejeunea apiculata (Horik.) S.Hatt., J. Hattori Bot. Lab. 5: 46, 1951. Bas.: Drepanolejeunea apiculata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 266, 1934.

** Leptolejeunea arunachalensis Sudipa Das et D.K.Singh, J. Jap. Bot. 83 (6): 343, 2008.

*** Leptolejeunea astroidea (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 363, 1913. Bas.: Lejeunea astroidea Mitt., Trans. Linn. Soc. London 23 (1): 58, 1860.

** Leptolejeunea australis Steph., Sp. Hepat. (Stephani) 5: 389, 1913. Based on: Leptolejeunea australis Steph., Hedwigia 28 (3): 173, 1889, nom. inval.

** Leptolejeunea balansae Steph., Hedwigia 35 (3): 105, 1896.

* Leptolejeunea borneensis Herzog, Flora 135: 394, 1942. 342

** Leptolejeunea brasiliensis Bischl., Nova Hedwigia 17: 301, 1969.

* Leptolejeunea convexistipa Bischl., Nova Hedwigia 17: 325, 1969.

* Leptolejeunea curvatifolia Steph., Sp. Hepat. (Stephani) 6: 398, 1923. 343

** Leptolejeunea denticulata Steph., Sp. Hepat. (Stephani) 5: 389, 1913. Based on: Leptolejeunea denticulata Steph., Hedwigia 28 (3): 174, 1889, nom. inval.

** Leptolejeunea dentistipula Steph., Sp. Hepat. (Stephani) 5: 379, 1913.

** Leptolejeunea diversilobulata Bischl., Nova Hedwigia 17: 313, 1969.

** Leptolejeunea dolabriformis Pearson, J. Linn. Soc., Bot. 46 (305): 37, 1922.

*** Leptolejeunea elliptica (Lehm. et Lindenb.) Besch., Rev. Bryol. 19 (1): 14, 1892. Bas.: Jungermannia elliptica Lehm. et Lindenb., Nov. Stirp. Pug. 5: 13, 1833.

* Leptolejeunea emarginata (Horik.) S.Hatt., J. Hattori Bot. Lab. 5: 46, 1951. Bas.: Drepanolejeunea emarginata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 267, 1934. 344

*** Leptolejeunea epiphylla (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 380, 1913. Bas.: Lejeunea epiphylla Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 118, 1860 [1861].

*** Leptolejeunea exocellata (Spruce) A.Evans, Bull. Torrey Bot. Club 29 (8): 498, 1902. Bas.: Lejeunea exocellata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 195, 1884.

** Leptolejeunea foliicola Steph., Hedwigia 35 (3): 106, 1896.

** Leptolejeunea integristipula Steph., Sp. Hepat. (Stephani) 6: 398, 1923.

* Leptolejeunea jamaicensis R.M.Schust., J. Elisha Mitchell Sci. Soc. 83 (4): 229, 1967.

** Leptolejeunea lancifolia Steph., Sp. Hepat. (Stephani) 5: 382, 1913. Nom. nov. pro Lejeunea lancifolia Mitt., Fl. vit.: 415, 1871 [1873], nom. illeg.

** Leptolejeunea latifolia Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 58, 1950 [1951].

** Leptolejeunea lepinii Steph., Sp. Hepat. (Stephani) 5: 383, 1913.

** Leptolejeunea ligulata Herzog, Flora 135: 429, 1942.

*** Leptolejeunea maculata (Mitt.) Schiffn., Consp. Hepat. Arch. Ind.: 275, 1898. Bas.: Lejeunea maculata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 118, 1860 [1861].

* Leptolejeunea massartiana Schiffn. ex Herzog, Flora 135: 421, 1942. 345

** Leptolejeunea micronesica Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 149, 1965.

** Leptolejeunea minima Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 60, 1950 [1951].

** Leptolejeunea mirikana M.Dey et D.K.Singh, Taiwania 55 (4): 355, 2010.

*** Leptolejeunea moniliata Steph., Sp. Hepat. (Stephani) 5: 371, 1913.

*** Leptolejeunea obfuscata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 373, 1913. Bas.: Lejeunea obfuscata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 579, 1885.

* Leptolejeunea punctata Herzog, Flora 135: 432, 1942.

*** Leptolejeunea radicosa (Nees ex Mont.) Grolle, J. Hattori Bot. Lab. 45: 178, 1979. Bas.: Lejeunea radicosa Nees ex Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 475, 1842.

* Leptolejeunea renneri Herzog, Flora 135: 422, 1942.

* Leptolejeunea revoluta P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 44, 1955. 346

** Leptolejeunea rosulans Steph., Sp. Hepat. (Stephani) 5: 390, 1913. Based on: Leptolejeunea rosulans Steph., Hedwigia 28 (3): 174, 1889, nom. inval.

** Leptolejeunea serratifolia Schiffn., Bot. Jahrb. Syst. 23 (5): 594, 1897.

** Leptolejeunea serrulata Herzog, Flora 135: 426, 1942.

** Leptolejeunea spinistipula (Mizut.) Xiao L.He, Ann. Bot. Fenn. 34 (2): 127, 1997. Bas.: Pycnolejeunea spinistipula Mizut., J. Hattori Bot. Lab. 33: 255, 1970.

** Leptolejeunea subdentata Schiffn. ex Herzog, Flora 135: 403, 1942.

*** Leptolejeunea subrotundifolia Herzog, Flora 135: 424, 1942.

*** Leptolejeunea tridentata Bischl., Nova Hedwigia 17: 335, 1969.

* Leptolejeunea trigonostipa (Spruce) Steph., Sp. Hepat. (Stephani) 5: 376, 1913. Bas.: Lejeunea trigonostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 197, 1884.

** Leptolejeunea tripuncta (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 388, 1913. Bas.: Lejeunea tripuncta Mitt., Fl. vit.: 415, 1871 [1873].

** Leptolejeunea truncatifolia Steph., Sp. Hepat. (Stephani) 5: 388, 1913.

* Leptolejeunea udarii M.Dey et D.K.Singh, Taiwania 55 (4): 359, 2010. 347

*** Leptolejeunea vitrea (Nees) Schiffn., Hepat. (Engl.-Prantl): 126, 1893. Bas.: Jungermannia vitrea Nees, Enum. Pl. Crypt. Javae: 56, 1830.

** subtrib. Pycnolejeuneinae Heinrichs et Schäf.Verw.

** Pycnolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 124, 1893. Bas.: Lejeunea subg. Pycnolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 246, 1884.

* Pycnolejeunea anotomensis Tixier, Bull. Soc. Hist. Nat. Afrique N. 63: 10, 1972. 348

** Pycnolejeunea borneensis Steph., Sp. Hepat. (Stephani) 5: 632, 1914.

* Pycnolejeunea cavistipula (Steph.) Mizut., J. Hattori Bot. Lab. 36: 161, 1972 [1973]. Bas.: Strepsilejeunea cavistipula Steph., Hedwigia 35 (3): 128, 1896. 349

* Pycnolejeunea connivens Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894. Based on: Pycnolejeunea connivens Gottsche ex Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 32, 1890, nom. inval.

*** Pycnolejeunea contigua (Nees) Grolle, J. Hattori Bot. Lab. 45: 179, 1979. Bas.: Jungermannia contigua Nees, Fl. Bras. (Martius) 1 (1): 360, 1833.

* Pycnolejeunea convexifolia (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 636, 1914. Bas.: Lejeunea convexifolia Mitt., Fl. vit.: 416, 1871 [1873]. 350

*** Pycnolejeunea decurviloba Steph., Hedwigia 35 (3): 125, 1896.

*** Pycnolejeunea densistipula (Lehm. et Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 602, 1914. Bas.: Lejeunea densistipula Lehm. et Lindenb., Nov. Stirp. Pug. 7: 20, 1838.

** Pycnolejeunea gradsteinii Ilk.-Borg., Bol. Inst. Bot. (São Paulo) 21 (1): 1, 2011.

** Pycnolejeunea grandiocellata Steph., Bot. Tidsskr. 24 (3): 279, 1902.

* Pycnolejeunea grossiloba Steph., Sp. Hepat. (Stephani) 5: 629, 1914. 351

*** Pycnolejeunea macroloba (Nees et Mont.) Schiffn., Hepat. (Engl.-Prantl): 124, 1893. Bas.: Lejeunea macroloba Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 260, 1843.

*** Pycnolejeunea monophthalma (R.M.Schust.) Xiao L.He, Acta Bot. Fenn. 163: 52, 1999. Bas.: Trachylejeunea monophthalma R.M.Schust., Bull. Torrey Bot. Club 97 (6): 345, 1970 [1971].

** Pycnolejeunea novae-caledoniae (Steph.) Horik., Acta Phytotax. Geobot. 13: 214, 1943. Bas.: Archilejeunea novae-caledoniae Steph., Sp. Hepat. (Stephani) 4: 729, 1911.

* Pycnolejeunea palmicola Steph., Sp. Hepat. (Stephani) 6: 413, 1923. 352

*** Pycnolejeunea papillosa Xiao L.He, Acta Bot. Fenn. 163: 55, 1999.

** Pycnolejeunea porrectilobula C.J.Bastos et O.Yano, Nova Hedwigia 74 (3/4): 440, 2002.

** Pycnolejeunea retusa R.M.Schust., J. Hattori Bot. Lab. 100: 402, 2006.

* Pycnolejeunea schlimiana Steph., Sp. Hepat. (Stephani) 5: 615, 1914. 353

*** Pycnolejeunea schwaneckei (Steph.) Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894. Bas.: Lejeunea schwaneckei Steph., Hedwigia 27 (11/12): 290, 1888.

** Pycnolejeunea sphaeroides (Sande Lac.) J.B.Jack et Steph., Bot. Centralbl. 60 (4): 107, 1894. Bas.: Lejeunea sphaeroides Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 309, 1864.

** subtrib. Xylolejeuneinae Heinrichs et Schäf.Verw.

*** Xylolejeunea Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 27, 2001.

*** Xylolejeunea aquarius (Spruce) Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 29, 2001. Bas.: Lejeunea aquarius Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 185, 1884.

*** Xylolejeunea crenata (Nees et Mont.) Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 36, 2001. Bas.: Lejeunea crenata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 48, 1838.

*** Xylolejeunea grolleana (Pócs) Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 32, 2001. Bas.: Trachylejeunea grolleana Pócs, Haussknechtia, Beih. 9: 285, 1999.

*** Xylolejeunea muricella Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 34, 2001.

** trib. Symbiezidieae Gradst.

*** Symbiezidium Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 402, 1877.

*** Symbiezidium barbiflorum (Lindenb. et Gottsche) A.Evans, Bull. Torrey Bot. Club 34 (11): 540, 1907 [1908]. Bas.: Lejeunea barbiflora Lindenb. et Gottsche, Linnaea 24 (6): 630, 1851 [1852].

*** Symbiezidium dentatum Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 175, 1955.

*** Symbiezidium madagascariense Steph., Sp. Hepat. (Stephani) 5: 99, 1912.

*** Symbiezidium transversale (Sw.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 403, 1877. Bas.: Jungermannia transversalis Sw., Prodr. (Swartz): 144, 1788.

*** Symbiezidium transversale var. hookerianum (Gottsche, Lindenb. et Nees) Gradst. et J.Beek, Beih. Nova Hedwigia 80: 237, 1985. Bas.: Lejeunea transversalis β hookeriana Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 311, 1845.

*** Ptychanthoideae Mizut.

*** Acrolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 nom. conserv. Bas.: Lejeunea subg. Acrolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 115, 1884.

*** Acrolejeunea sandvicensis (Gottsche) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 312, 1896. Bas.: Phragmicoma sandvicensis Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 344, 1857.

** sect. Acrolejeunea

*** Acrolejeunea emergens (Mitt.) Steph., Pflanzenw. Ost-Afrikas C: 65, 1895. Bas.: Phragmicoma emergens Mitt., Philos. Trans. 168: 397, 1879.

*** Acrolejeunea emergens var. confertissima (Steph.) Gradst., Bryophyt. Biblioth. 4: 76, 1975. Bas.: Acrolejeunea confertissima Steph., Hedwigia 31 (4): 165, 1892.

*** Acrolejeunea heterophylla (A.Evans) Grolle et Gradst., J. Hattori Bot. Lab. 38: 332, 1974. Bas.: Ptychocoleus heterophyllus A.Evans, Amer. J. Bot. 5 (3): 144, 1918.

*** Acrolejeunea torulosa (Lehm. et Lindenb.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Jungermannia torulosa Lehm. et Lindenb., Nov. Stirp. Pug. 6: 41, 1834.

** sect. Minores (Verd.) L.Söderstr. et A.Hagborg, Bryophyte Diversity Evol. 36 (1): 41, 2014. Bas.: Ptychocoleus sect. Minores Verd., Ann. Bryol., Suppl. 4: 132, 1934.

*** Acrolejeunea arcuata (Nees) Grolle et Gradst., J. Hattori Bot. Lab. 38: 332, 1974. Bas.: Jungermannia arcuata Nees, Enum. Pl. Crypt. Javae: 38, 1830.

** Acrolejeunea arcuata subsp. gradsteinii M.A.M.Renner, Phytotaxa 83 (1): 42, 2013.

*** Acrolejeunea fertilis (Reinw., Blume et Nees) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Jungermannia fertilis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 211, 1824 [1825].

*** Acrolejeunea parvula (Mizut.) Gradst., Bryophyt. Biblioth. 4: 115, 1975. Bas.: Ptychocoleus parvulus Mizut., Dansk Bot. Ark. 27 (1): 97, 1969.

*** Acrolejeunea pycnoclada (Taylor) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Ptychanthus pycnocladus Taylor, London J. Bot. 5: 385, 1846.

*** Acrolejeunea pycnoclada subsp. latistipula Gradst., Bryophyt. Biblioth. 4: 113, 1975.

*** Acrolejeunea tjibodensis (Verd.) Grolle et Gradst., J. Hattori Bot. Lab. 38: 333, 1974. Bas.: Ptychocoleus tjibodensis Verd., Recueil Trav. Bot. Neerl. 30: 227, 1933.

** sect. Pusillae Gradst., Bryophyt. Biblioth. 4: 59, 1975.

*** Acrolejeunea pusilla (Steph.) Grolle et Gradst., J. Hattori Bot. Lab. 38: 332, 1974. Bas.: Archilejeunea pusilla Steph., Sp. Hepat. (Stephani) 4: 731, 1911.

*** Acrolejeunea sikkimensis (Mizut.) Gradst., Bryophyt. Biblioth. 4: 83, 1975. Bas.: Ptychocoleus sikkimensis Mizut., Fl. E. Himalaya: 532, 1966.

** sect. Recurvatae Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 39, 2014.

*** Acrolejeunea recurvata Gradst., Bryophyt. Biblioth. 4: 79, 1975.

** sect. Regulares (Verd.) Gradst., Bryophyt. Biblioth. 4: 63, 1975. Bas.: Ptychocoleus sect. Regulares Verd., Ann. Bryol., Suppl. 4: 143, 1934.

*** Acrolejeunea allisonii Gradst., Bryophyt. Biblioth. 4: 103, 1975.

*** Acrolejeunea aulacophora (Mont.) Steph., Bot. Jahrb. Syst. 20 (3): 317, 1895. Bas.: Phragmicoma aulacophora Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 259, 1843.

*** Acrolejeunea mollis (Hook.f. et Taylor) Schiffn., Hedwigia 33 (4): 178, 1894. Bas.: Ptychanthus mollis Hook.f. et Taylor, London J. Bot. 5: 384, 1846.

*** Acrolejeunea securifolia (Nees) Steph., Hedwigia 34 (2): 59, 1895. Bas.: Jungermannia securifolia Nees, Prodr. Fl. Norfolk.: 5, 1833.

*** Acrolejeunea securifolia subsp. caledonica (Steph.) Gradst., Bryophyt. Biblioth. 4: 100, 1975. Bas.: Ptychocoleus caledonicus Steph., Sp. Hepat. (Stephani) 5: 39, 1912.

*** Acrolejeunea securifolia subsp. hartmannii (Steph.) Gradst., Bryophyt. Biblioth. 4: 99, 1975. Bas.: Ptychocoleus hartmannii Steph., Sp. Hepat. (Stephani) 5: 44, 1912.

*** Acrolejeunea securifolia subsp. pallida (Ångstr.) Gradst., Bryophyt. Biblioth. 4: 101, 1975. Bas.: Phragmicoma pallida Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 132, 1873.

** sect. Trocholejeunea (Schiffn.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 38, 2014. Bas.: Trocholejeunea Schiffn., Ann. Bryol. 5: 160, 1932.

*** Acrolejeunea crassicaulis (Steph.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 38, 2014. Bas.: Hygrolejeunea crassicaulis Steph., Sp. Hepat. (Stephani) 5: 550, 1914.

*** Acrolejeunea infuscata (Mitt.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 38, 2014. Bas.: Lejeunea infuscata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 111, 1860 [1861].

** Acrolejeunea meghalayensis (Ajit P.Singh et V.Nath) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 39, 2014. Bas.: Trocholejeunea meghalayensis Ajit P.Singh et V.Nath, J. Jap. Bot. 83 (1): 2, 2008.

** Acrolejeunea sinensis (Jian Wang bis, R.L.Zhu et Gradst.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 39, 2014. Bas.: Trocholejeunea sinensis Jian Wang bis, R.L.Zhu et Gradst., Phytotaxa 174 (5): 296, 2014.

Excluded from the genus

* Acrolejeunea abnormis (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 4, 1892. Bas.: Phragmicoma abnormis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 352, 1882. 354

* Acrolejeunea comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 33, 1922. 355

* Acrolejeunea inflexa (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 4, 1892. Bas.: Phragmicoma inflexa Gottsche, Abh. Naturwiss. Vereins Bremen 7: 351, 1882. 356

*** Archilejeunea (Spruce) Steph., Hedwigia 27 (3/4): 113, 1888. Bas.: Lejeunea subg. Archilejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 88, 1884. 357

*** Archilejeunea abbreviata (Mitt.) Vanden Berghen, Rev. Bryol. Lichénol. 20 (1/2): 117, 1951. Bas.: Lejeunea abbreviata Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 167, 1863.

** Archilejeunea africana Steph., Sp. Hepat. (Stephani) 4: 705, 1911.

** Archilejeunea alata Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 33, 1893 [1894].

** Archilejeunea amakawana Inoue, J. Jap. Bot. 41 (1): 16, 1966. Nom. nov. pro Archilejeunea falcata Amakawa, J. Jap. Bot. 39 (5): 137, 1964, nom. illeg.

*** Archilejeunea auberiana (Mont.) Steph., Hedwigia 29 (3): 134, 1890. Bas.: Lejeunea auberiana Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 483, 1842.

*** Archilejeunea autoica Vanden Berghen, Rev. Bryol. Lichénol. 20 (1/2): 119, 1951 (Vanden Bergen 1951b).

*** Archilejeunea badia (Spruce) Steph., Sp. Hepat. (Stephani) 4: 711, 1911. Bas.: Lejeunea badia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 92, 1884.

* Archilejeunea bilabiata (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 723, 1911. Bas.: Phragmicoma bilabiata Mitt., Fl. vit.: 412, 1871 [1873]. 358

*** Archilejeunea bischleriana Gradst., Fl. Neotrop. Monogr. 62: 62, 1994.

** Archilejeunea bongardii Steph., Hedwigia 29 (1): 20, 1890.

** Archilejeunea brachyantha J.B.Jack et Steph., Bot. Centralbl. 60 (4): 104, 1894.

** Archilejeunea brevilobula Steph., Sp. Hepat. (Stephani) 4: 706, 1911.

*** Archilejeunea crispistipula (Spruce) Steph., Sp. Hepat. (Stephani) 4: 712, 1911. Bas.: Lejeunea crispistipula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 93, 1884.

* Archilejeunea eberhardtii Steph., Sp. Hepat. (Stephani) 4: 725, 1911.

** Archilejeunea elobulata Steph., Sp. Hepat. (Stephani) 4: 707, 1911.

** Archilejeunea gradsteinii X.Q.Shi et R.L.Zhu, Nova Hedwigia 100 (3/4): 592, 2015 (Shi and Zhu 2015).

** Archilejeunea incrassata Steph., Rev. Bryol. 35 (2): 30, 1908.

** Archilejeunea jonesii Vanden Berghen, Rev. Bryol. Lichénol. 20 (1/2): 116, 1951.

*** Archilejeunea juliformis (Nees) Gradst., Bryophyt. Biblioth. 4: 127, 1975. Bas.: Jungermannia juliformis Nees, Fl. Bras. (Martius) 1 (1): 351, 1833.

*** Archilejeunea kiushiana (Horik.) Verd., Ann. Bryol., Suppl. 4: 46, 1934. Bas.: Lopholejeunea kiushiana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 129, 1932.

*** Archilejeunea ludoviciana (De Not.) P.Geissler et Gradst., J. Hattori Bot. Lab. 75: 202, 1994. Bas.: Phragmicoma ludoviciana De Not., Nov. Stirp. Pug. 10: 11, 1857.

*** Archilejeunea ludoviciana subsp. porelloides (Spruce) Gradst., Fl. Neotrop. Monogr. 62: 58, 1994. Bas.: Lejeunea porelloides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 90, 1884.

** Archilejeunea nebeliana Gradst. et Schäf.-Verw., Cryptog. Bryol. 33 (2): 108, 2012.

*** Archilejeunea olivacea (Hook.f. et Taylor) Steph., Hedwigia 29 (3): 134, 1890. Bas.: Jungermannia olivacea Hook.f. et Taylor, London J. Bot. 3: 568, 1844.

*** Archilejeunea parviflora (Nees) Steph., Hedwigia 29 (3): 134, 1890. Bas.: Jungermannia parviflora Nees, Fl. Bras. (Martius) 1 (1): 353, 1833.

** Archilejeunea planifolia (Horik.) Mizut., J. Hattori Bot. Lab. 73: 175, 1993. Bas.: Leucolejeunea planifolia Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 199, 1933. 359

*** Archilejeunea planiuscula (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 731, 1911. Bas.: Lejeunea planiuscula Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 111, 1860 [1861].

Excluded from the genus

* Archilejeunea negrensis Steph., Sp. Hepat. (Stephani) 4: 716, 1911. 360

* Archilejeunea ovata Herzog, Rev. Bryol. Lichénol. 20 (1/2): 130, 1951 [1952]. 361

*** Bryopteris (Nees) Lindenb., Syn. Hepat. 2: 284, 1845. Bas.: Frullania subg. Bryopteris Nees, Naturgesch. Eur. Leberm. 3: 211, 1838.

*** Bryopteris diffusa (Sw.) Nees, Syn. Hepat. 2: 286, 1845. Bas.: Jungermannia diffusa Sw., Prodr. (Swartz): 144, 1788.

*** Bryopteris filicina (Sw.) Nees, Syn. Hepat. 2: 284, 1845. Bas.: Jungermannia filicina Sw., Prodr. (Swartz): 145, 1788.

* Bryopteris fissiloba Steph., Sp. Hepat. (Stephani) 6: 568, 1924. 362

*** Bryopteris gaudichaudii Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 340, 1857.

*** Caudalejeunea Schiffn., Hepat. (Engl.-Prantl): 129, 1893.

** subg. Acaudalejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 779, 1980.

*** Caudalejeunea grolleana Gradst., Acta Bot. Neerl. 23 (3): 334, 1974.

** subg. Caudalejeunea

** Caudalejeunea africana (Steph.) Schiffn., Hepat. (Engl.-Prantl): 129, 1893. Bas.: Thysananthus africanus Steph., Bot. Jahrb. Syst. 8 (2): 93, 1886.

*** Caudalejeunea hanningtonii (Mitt.) Schiffn., Hepat. (Engl.-Prantl): 129, 1893. Bas.: Lejeunea hanningtonii Mitt., J. Linn. Soc., Bot. 22 (146): 324, 1886.

** Caudalejeunea katangensis Vanden Berghen, Explor. Hydrobiol. Lac Bangweolo Luapula: 94, 1972.

*** Caudalejeunea lehmanniana (Gottsche) A.Evans, Bull. Torrey Bot. Club 34 (11): 554, 1907 [1908]. Bas.: Lejeunea lehmanniana Gottsche, Syn. Hepat. 3: 325, 1845.

** Caudalejeunea lewallei Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 42 (4): 434, 1972.

** subg. Vermilejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 778, 1980.

*** Caudalejeunea cristiloba (Steph.) Gradst., Acta Bot. Neerl. 23 (3): 340, 1974. Bas.: Acrolejeunea cristiloba Steph., Hedwigia 34 (2): 56, 1895.

*** Caudalejeunea cristiloba subsp. samoana (Steph.) Gradst., Acta Bot. Neerl. 23 (3): 343, 1974. Bas.: Caudalejeunea samoana Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 296, 1907.

** Caudalejeunea dusenii Steph., Sp. Hepat. (Stephani) 5: 11, 1912.

*** Caudalejeunea reniloba (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 16, 1912. Bas.: Phragmicoma reniloba Gottsche, Syn. Hepat. 2: 301, 1845.

** Caudalejeunea yangambiensis (Vanden Berghen) E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 192, 1957. Bas.: Ptychocoleus yangambiensis Vanden Berghen, Bull. Soc. Roy. Bot. Belgique 84: 61, 1951.

Incertae sedis

** Caudalejeunea acutifolia Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 478, 1947.

* Caudalejeunea mauritiana Horik., Acta Phytotax. Geobot. 13: 214, 1943. Nom. nov. pro Dicranolejeunea africana Steph., Sp. Hepat. (Stephani) 5: 158, 1912.

** Caudalejeunea streimannii Gyarmati, Trop. Bryol. 22: 129, 2002.

** Caudalejeunea tridentata R.L.Zhu, Y.M.Wei et Qiong He, Bryologist 114 (3): 469, 2011.

** Cephalantholejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 798, 1980.

*** Cephalantholejeunea temnanthoides (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 807, 1980. Bas.: Potamolejeunea temnanthoides R.M.Schust., Beih. Nova Hedwigia 9: 123, 1963.

** Cephalolejeunea Mizut., J. Hattori Bot. Lab. 46: 359, 1979.

*** Cephalolejeunea parvilobula Mizut., J. Hattori Bot. Lab. 46: 359, 1979.

*** Frullanoides Raddi, Critt. Brasil.: 13, 1822.

*** Frullanoides bahamensis (A.Evans) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 81, 1985. Bas.: Brachiolejeunea bahamensis A.Evans, Bull. Torrey Bot. Club 35 (8): 383, 1908.

*** Frullanoides corticalis (Lehm. et Lindenb.) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 84, 1985. Bas.: Jungermannia corticalis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 50, 1832.

*** Frullanoides densifolia Raddi, Critt. Brasil.: 14, 1822.

*** Frullanoides densifolia subsp. grandidentata (L.Clark) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 95, 1985. Bas.: Brachiolejeunea grandidentata L.Clark, Proc. Calif. Acad. Sci. (ser. 4) 27 (18): 595, 1953.

*** Frullanoides laciniatiflora (Loitl.) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 100, 1985. Bas.: Lejeunea laciniatiflora Loitl., Diagn. pl. nov.: 19, 1894.

*** Frullanoides liebmaniana (Lindenb. et Gottsche) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 102, 1985. Bas.: Phragmicoma liebmaniana Lindenb. et Gottsche, Syn. Hepat. 5: 744, 1847.

*** Frullanoides mexicana van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 106, 1985.

*** Frullanoides tristis (Steph.) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 110, 1985. Bas.: Brachiolejeunea tristis Steph., Sp. Hepat. (Stephani) 5: 112, 1912.

*** Fulfordianthus Gradst., Bryologist 95 (1): 44, 1992.

*** Fulfordianthus evansii (Fulford) Gradst., Bryologist 95 (1): 46, 1992. Bas.: Thysananthus evansii Fulford, Bull. Torrey Bot. Club 68 (1): 34, 1941.

*** Fulfordianthus pterobryoides (Spruce) Gradst., Bryologist 95 (1): 44, 1992. Bas.: Lejeunea pterobryoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 109, 1884.

*** Lopholejeunea (Spruce) Steph., Bot. Gaz. 15 (11): 285, 1890 nom. rejic. Bas.: Lejeunea subg. Lopholejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 119, 1884.

** subg. Lopholejeunea

** Lopholejeunea borbonica Steph., Hedwigia 35 (3): 109, 1896.

** Lopholejeunea jonesii Vanden Berghen, Bull. Jard. Bot. État Bruxelles 20 (2): 178, 1950.

** Lopholejeunea laciniata E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 194, 1957.

** Lopholejeunea minima Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 437, 1984.

* Lopholejeunea obtusilacera Herzog, Bull. Jard. Bot. État Bruxelles 20 (2): 172, 1950.

* Lopholejeunea paramultilacera Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 435, 1984.

* Lopholejeunea quinquecarinata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 408, 1984.

** Lopholejeunea renistipula (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 94, 1912. Bas.: Phragmicoma renistipula Mitt., Fl. vit.: 413, 1871 [1873].

** Lopholejeunea revoluta E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 207, 1957.

** sect. Eulophae Verd., Ann. Bryol., Suppl. 4: 87, 1934.

*** Lopholejeunea applanata (Reinw., Blume et Nees) Schiffn., Hepat. (Engl.-Prantl): 129, 1893. Bas.: Jungermannia applanata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 210, 1824 [1825].

*** Lopholejeunea borneensis (Steph.) Verd., Ann. Bryol., Suppl. 4: 83, 1934. Bas.: Mastigolejeunea borneensis Steph., Sp. Hepat. (Stephani) 4: 777, 1912.

*** Lopholejeunea erugata B.M.Thiers, Brittonia 36 (2): 174, 1984. Nom. nov. pro Ptychocoleus inermis Steph., Sp. Hepat. (Stephani) 5: 27, 1912.

*** Lopholejeunea eulopha (Taylor) Schiffn., Hepat. (Engl.-Prantl): 129, 1893. Bas.: Lejeunea eulopha Taylor, London J. Bot. 5: 391, 1846.

*** Lopholejeunea evansiana Verd., Nova Guinea 18: 4, 1934.

*** Lopholejeunea grollei R.L.Zhu et Gradst., Monogr. Syst. Bot. Missouri Bot. Gard. 74: 36, 2005.

*** Lopholejeunea herzogiana Verd., Recueil Trav. Bot. Neerl. 30: 217, 1933.

*** Lopholejeunea hispidissima Steph., Sp. Hepat. (Stephani) 5: 80, 1912.

*** Lopholejeunea loheri Steph., Sp. Hepat. (Stephani) 5: 77, 1912.

*** Lopholejeunea minuta R.L.Zhu et Gradst., Nova Hedwigia 78 (3/4): 436, 2004.

*** Lopholejeunea nigricans (Lindenb.) Schiffn., Consp. Hepat. Arch. Ind.: 293, 1898. Bas.: Lejeunea nigricans Lindenb., Syn. Hepat. 3: 316, 1845.

*** Lopholejeunea plicatiscypha (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 96, 1912. Bas.: Phragmicoma plicatiscypha Hook.f. et Taylor, London J. Bot. 5: 386, 1846.

* Lopholejeunea proxima Steph., Sp. Hepat. (Stephani) 5: 89, 1912. 363

*** Lopholejeunea streimannii B.M.Thiers et Gradst., Mem. New York Bot. Gard. 52: 37, 1989.

** Lopholejeunea vojtkoana Gyarmati, Nova Hedwigia 87 (3/4): 480, 2008.

** sect. Lopholejeunea

*** Lopholejeunea ceylanica Steph., Sp. Hepat. (Stephani) 5: 86, 1912.

*** Lopholejeunea horticola Schiffn., Ann. Bryol. 6: 133, 1933.

*** Lopholejeunea latilobula Verd., Nova Guinea 18: 4, 1934.

*** Lopholejeunea magna Mizut., J. Hattori Bot. Lab. 32: 131, 1969.

*** Lopholejeunea recurvata Mizut., J. Hattori Bot. Lab. 46: 369, 1979.

*** Lopholejeunea soae R.L.Zhu et Gradst., Monogr. Syst. Bot. Missouri Bot. Gard. 74: 69, 2005.

*** Lopholejeunea subfusca (Nees) Schiffn., Bot. Jahrb. Syst. 23 (5): 593, 1897. Bas.: Jungermannia subfusca Nees, Enum. Pl. Crypt. Javae: 36, 1830.

** Lopholejeunea subfusca var. elongata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 445, 1984.

*** Lopholejeunea wiltensii Steph., Hedwigia 35 (3): 112, 1896.

* Lopholejeunea yapensis Steph., Sp. Hepat. (Stephani) 5: 81, 1912. 364

*** Lopholejeunea zollingeri (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 296, 1898. Bas.: Lejeunea zollingeri Steph., Hedwigia 29 (1): 14, 1890.

** subg. Pholianthus B.M.Thiers et Gradst., Mem. New York Bot. Gard. 52: 25, 1989.

*** Lopholejeunea colensoi Steph., Sp. Hepat. (Stephani) 5: 97, 1912.

*** Lopholejeunea pocsii Gyarmati, Cryptog. Bryol. 26 (4): 404, 2005.

** subg. Pteryganthus B.M.Thiers, Brittonia 35 (1): 85, 1983.

** Lopholejeunea grandicrista Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 34, 1893 [1894].

*** Lopholejeunea leioptera Gyarmati, Candollea 56 (1): 80, 2001.

** Lopholejeunea onraedtii Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 452, 1984.

** Lopholejeunea sphaerophora (Lehm. et Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 68, 1912. Bas.: Jungermannia sphaerophora Lehm. et Lindenb., Nov. Stirp. Pug. 5: 9, 1833.

* Lopholejeunea tixieriana Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 454, 1984.

** Lopholejeunea utriculata Steph., Sp. Hepat. (Stephani) 5: 69, 1912.

Incertae sedis

** Lopholejeunea lepidoscypha Kiaer et Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 5, 1892.

* Lopholejeunea multilacera Steph., Bot. Gaz. 15 (11): 285, 1890.

** Lopholejeunea udarii M.Dey et D.K.Singh, Nelumbo 53: 197, 2011.

Excluded from the genus

* Lopholejeunea aberrantia Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 256, 1934. 365

* Lopholejeunea vietnamica Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 99, 1974. 366

*** Marchesinia Gray, Nat. Arr. Brit. Pl. 1: 689, 1821 nom. conserv.

*** subg. Marchesinia

*** Marchesinia mackaii (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 689, 1821. Bas.: Jungermannia mackaii Hook., Brit. Jungermann.: tab. 53, 1813.

*** subg. Marchesiniopsis R.M.Schust., J. Hattori Bot. Lab. 72: 358, 1992.

*** Marchesinia bongardiana (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 405, 1877. Bas.: Lejeunea bongardiana Lehm. et Lindenb., Nov. Stirp. Pug. 7: 18, 1838.

*** Marchesinia brachiata (Sw.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Jungermannia brachiata Sw., Prodr. (Swartz): 144, 1788.

*** Marchesinia deslooveri Vanden Berghen, Rev. Bryol. Lichénol. 42 (4): 926, 1976.

*** Marchesinia excavata (Mitt.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Lejeunea excavata Mitt., Trans. Linn. Soc. London 23 (1): 58, 1860.

*** Marchesinia languida (Nees et Mont.) Steph., Sp. Hepat. (Stephani) 5: 149, 1912. Bas.: Lejeunea languida Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 59, 1836.

*** Marchesinia nobilis (Gottsche) X.Q.Shi, R.L.Zhu et Gradst., Phytotaxa 195 (3): 249, 2015 (b). Bas.: Lejeunea nobilis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 353, 1882.

*** Marchesinia robusta (Mitt.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893. Bas.: Lejeunea robusta Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 359, 1851.

** Mastigolejeunea (Spruce) Steph., Hedwigia 30 (5): 206, 1891. Bas.: Lejeunea subg. Mastigolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 100, 1884.

*** Mastigolejeunea auriculata (Wilson et Hook.) Steph., Bot. Gaz. 17 (6): 171, 1892. Bas.: Jungermannia auriculata Wilson et Hook., Musci Amer., S. States: no. 170, 1841 (; non vidi).

** Mastigolejeunea auriculata var. rhodesica (Vanden Berghen) Sukkharak et Gradst., Nova Hedwigia 99 (3/4): 297, 2014. Bas.: Brachiolejeunea rhodesica Vanden Berghen, Bull. Jard. Bot. État Bruxelles 21 (1/2): 94, 1951.

*** Mastigolejeunea calcarata (Steph.) Verd., Blumea 1 (1): 230, 1934. Bas.: Archilejeunea calcarata Steph., Sp. Hepat. (Stephani) 4: 724, 1911.

*** Mastigolejeunea florea (Mitt.) Paris, Rev. Bryol. 33 (3): 42, 1906. Bas.: Phragmicoma florea Mitt., J. Linn. Soc., Bot. 22 (146): 323, 1886.

*** Mastigolejeunea frauenfeldii (Reichardt) Verd., Blumea 1 (1): 230, 1934. Bas.: Thysananthus frauenfeldii Reichardt, Verh. K.K. Zool.-Bot. Ges. Wien 16: 958, 1866.

** Mastigolejeunea gradsteinii Sukkharak, J. Bryol. 36 (1): 56, 2014.

*** Mastigolejeunea humilis (Gottsche) Schiffn., Hepat. (Engl.-Prantl): 129, 1893. Bas.: Phragmicoma humilis Gottsche, Syn. Hepat. 2: 299, 1845. 367

*** Mastigolejeunea indica Steph., Sp. Hepat. (Stephani) 4: 776, 1912.

*** Mastigolejeunea innovans (Spruce) Steph., Sp. Hepat. (Stephani) 4: 765, 1912. Bas.: Lejeunea innovans Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 103, 1884.

*** Mastigolejeunea ligulata (Lehm. et Lindenb.) Schiffn., Hepat. (Engl.-Prantl): 129, 1893. Bas.: Jungermannia ligulata Lehm. et Lindenb., Nov. Stirp. Pug. 6: 39, 1834.

*** Mastigolejeunea nigra Steph., Hedwigia 30 (5): 206, 1891.

*** Mastigolejeunea plicatiflora (Spruce) Steph., Sp. Hepat. (Stephani) 4: 766, 1912. Bas.: Lejeunea plicatiflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 104, 1884.

*** Mastigolejeunea recondita (Steph.) Mizut., J. Hattori Bot. Lab. 32: 134, 1969. Bas.: Ptycholejeunea recondita Steph., Hedwigia 35 (3): 122, 1896.

*** Mastigolejeunea repleta (Taylor) A.Evans, Mem. Torrey Bot. Club 8 (2): 131, 1902. Bas.: Lejeunea repleta Taylor, London J. Bot. 5: 392, 1846.

*** Mastigolejeunea truncata Mizut., J. Hattori Bot. Lab. 61: 292, 1986 [1987].

** Mastigolejeunea turgida Steph., Hedwigia 31 (4): 170, 1892.

*** Mastigolejeunea virens (Ångstr.) Steph., Sp. Hepat. (Stephani) 4: 776, 1912, nom. conserv. Bas.: Thysananthus virens Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 131, 1873.

** Phaeolejeunea Mizut., J. Hattori Bot. Lab. 31: 130, 1968.

*** Phaeolejeunea amicorum (Hürl.) Pócs, Fieldiana, Bot. (n.ser.) 47: 140, 2008. Bas.: Phaeolejeunea etesseana subsp. amicorum Hürl., Bauhinia 9 (4): 263, 1991.

** Phaeolejeunea etesseana (Steph.) Mizut., J. Hattori Bot. Lab. 31: 133, 1968. Bas.: Brachiolejeunea etesseana Steph., Sp. Hepat. (Stephani) 5: 133, 1912.

** Phaeolejeunea inermis (Steph.) Mizut., J. Hattori Bot. Lab. 31: 134, 1968. Bas.: Lopholejeunea inermis Steph., Sp. Hepat. (Stephani) 5: 92, 1912.

*** Phaeolejeunea latistipula (Schiffn. ex P.Syd.) Mizut., J. Hattori Bot. Lab. 31: 131, 1968. Bas.: Hygrolejeunea latistipula Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894.

*** Ptychanthus Nees, Naturgesch. Eur. Leberm. 3: 211, 1838.

** Ptychanthus africanus Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 131, 1911.

* Ptychanthus stephensonianus (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 754, 1912. Bas.: Lejeunea stephensoniana Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 155, 1854. 368

*** Ptychanthus striatus (Lehm. et Lindenb.) Nees, Naturgesch. Eur. Leberm. 3: 212, 1838. Bas.: Jungermannia striata Lehm. et Lindenb., Nov. Stirp. Pug. 4: 16, 1832.

** Ptychanthus striatus var. intermedius (Gottsche) Verd., Ann. Bryol., Suppl. 4: 122, 1934. Bas.: Ptychanthus intermedius Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 576, 1853.

*** Schiffneriolejeunea Verd., Ann. Bryol. 6: 89, 1933.

** sect. Pappeanae R.M.Schust. ex Gradst. et Vanden Berghen, Beih. Nova Hedwigia 80: 174, 1985. Based on: Phragmilejeunea R.M.Schust., J. Hattori Bot. Lab. 11: 27, 1954.

*** Schiffneriolejeunea fragilis Gradst. et E.W.Jones, J. Bryol. 12 (1): 45, 1982.

*** Schiffneriolejeunea madagascariensis (Steph.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974. Bas.: Ptychocoleus madagascariensis Steph., Sp. Hepat. (Stephani) 5: 27, 1912.

*** Schiffneriolejeunea pappeana var. bidentata Gradst. et Vanden Berghen, Beih. Nova Hedwigia 80: 182, 1985.

*** Schiffneriolejeunea pappeana var. integra Gradst. et Vanden Berghen, Beih. Nova Hedwigia 80: 182, 1985.

*** Schiffneriolejeunea parviloba (Steph.) Gradst., J. Hattori Bot. Lab. 38: 335, 1974. Bas.: Acrolejeunea parviloba Steph., Bot. Gaz. 15 (11): 286, 1890.

*** Schiffneriolejeunea pappeana (Nees) Gradst., J. Hattori Bot. Lab. 38: 335, 1974. Bas.: Phragmicoma pappeana Nees, Syn. Hepat. 2: 296, 1845.

** sect. Schiffneriolejeunea, Occas. Pap. Farlow Herb. Cryptog. Bot. 16: 72, 1981.

*** Schiffneriolejeunea altimontana Vanden Berghen, Rev. Bryol. Lichénol. 42 (4): 923, 1976.

*** Schiffneriolejeunea amazonica Gradst., Beih. Nova Hedwigia 80: 25, 1985.

*** Schiffneriolejeunea cumingiana (Mont.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974. Bas.: Phragmicoma cumingiana Mont., London J. Bot. 4: 7, 1845.

*** Schiffneriolejeunea ferruginea (Steph.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974. Bas.: Acrolejeunea ferruginea Steph., Hedwigia 34 (2): 57, 1895.

*** Schiffneriolejeunea occulta (Steph.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974. Bas.: Ptychocoleus occultus Steph., Sp. Hepat. (Stephani) 5: 25, 1912.

*** Schiffneriolejeunea omphalanthoides Verd., Ann. Bryol. 6: 91, 1933.

*** Schiffneriolejeunea tumida (Nees) Gradst., J. Hattori Bot. Lab. 38: 335, 1974. Bas.: Ptychanthus tumidus Nees, Naturgesch. Eur. Leberm. 3: 213, 1838.

*** Schiffneriolejeunea tumida var. haskarliana (Gottsche) Gradst. et Terken, Occas. Pap. Farlow Herb. Cryptog. Bot. 16: 77, 1981. Bas.: Phragmicoma haskarliana Gottsche, Syn. Hepat. 2: 299, 1845.

*** Schiffneriolejeunea nymannii (Steph.) Gradst. et Terken, Occas. Pap. Farlow Herb. Cryptog. Bot. 16: 79, 1981. Bas.: Archilejeunea nymannii Steph., Sp. Hepat. (Stephani) 4: 730, 1911.

*** Schiffneriolejeunea polycarpa (Nees) Gradst., J. Hattori Bot. Lab. 38: 335, 1974. Bas.: Jungermannia polycarpa Nees, Fl. Bras. (Martius) 1 (1): 350, 1833.

*** Schiffneriolejeunea pulopenangensis (Gottsche) Gradst., J. Hattori Bot. Lab. 38: 335, 1974. Bas.: Phragmicoma pulopenangensis Gottsche, Syn. Hepat. 2: 299, 1845.

** Spruceanthus Verd., Ann. Bryol., Suppl. 4: 151, 1934.

*** Spruceanthus macrostipulus (Steph.) Gradst., Trop. Bryol. 4: 13, 1991. Bas.: Mastigolejeunea macrostipula Steph., Sp. Hepat. (Stephani) 4: 767, 1912.

** Spruceanthus mamillilobulus (Herzog) Verd., Hepat. Select. Crit. 9: no. 447, 1936 (; non vidi). Bas.: Ptychanthus mamillilobulus Herzog, Symb. Sin. 5: 44, 1930.

*** Spruceanthus pluriplicatus (Steph.) Gradst., Schriftenreihe Mensch, Kultur Umwelt z. Bergl. W Neug. 7: 14, 1981. Bas.: Brachiolejeunea pluriplicata Steph., Sp. Hepat. (Stephani) 5: 135, 1912.

*** Spruceanthus polymorphus (Sande Lac.) Verd., Ann. Bryol., Suppl. 4: 155, 1934. Bas.: Phragmicoma polymorpha Sande Lac., Ned. Kruidk. Arch. 3: 420, 1854 [1855].

*** Spruceanthus semirepandus (Nees) Verd., Ann. Bryol., Suppl. 4: 153, 1934. Bas.: Jungermannia semirepanda Nees, Enum. Pl. Crypt. Javae: 39, 1830.

*** Spruceanthus sulcatus (Nees) Gradst., Beih. Nova Hedwigia 80: 26, 1985. Bas.: Jungermannia sulcata Nees, Enum. Pl. Crypt. Javae: 36, 1830.

*** Spruceanthus theobromae (Spruce) Gradst., Beih. Nova Hedwigia 80: 26, 1985. Bas.: Lejeunea theobromae Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 99, 1884.

*** Spruceanthus thozetianus (Gottsche et F.Muell.) B.M.Thiers et Gradst., Mem. New York Bot. Gard. 52: 62, 1989. Bas.: Phragmicoma thozetiana Gottsche et F.Muell., Fragm. (Mueller): 63, 1880.

*** Thysananthus Lindenb., Nov. Stirp. Pug. 8: 24, 1844.

** sect. Thysananthus

** subsect. Anguiformes Sukkharak, Phytotaxa 193 (1): 37, 2015.

*** Thysananthus anguiformis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 289, 1845. Bas.: Jungermannia anguiformis Hook.f. et Taylor, London J. Bot. 3: 567, 1844.

*** Thysananthus pancheri (Steph.) Hürl., Bauhinia 9 (2): 167, 1989. Bas.: Mastigolejeunea pancheri Steph., Sp. Hepat. (Stephani) 4: 771, 1912.

** subsect. Thysananthus

*** Thysananthus aculeatus Herzog, Ann. Bryol. 4: 89, 1931.

*** Thysananthus amazonicus (Spruce) Schiffn., Hepat. (Engl.-Prantl): 130, 1893. Bas.: Lejeunea amazonica Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 106, 1884.

*** Thysananthus appendiculatus Steph., Sp. Hepat. (Stephani) 4: 794, 1912.

** Thysananthus ciliaris (Sande Lac.) Sukkharak, Nova Hedwigia 99 (3/4): 339, 2014. Bas.: Phragmicoma ciliaris Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 307, 1864.

*** Thysananthus comosus Lindenb., Nov. Stirp. Pug. 8: 25, 1844.

*** Thysananthus convolutus Lindenb., Syn. Hepat. 2: 288, 1845.

** Thysananthus convolutus var. laceratus (Steph.) Sukkharak, Phytotaxa 193 (1): 30, 2015. Bas.: Thysananthus laceratus Steph., Sp. Hepat. (Stephani) 4: 796, 1912.

*** Thysananthus discretus Sukkharak et Gradst., Cryptog. Bryol. 31 (2): 113, 2010.

*** Thysananthus gottschei (J.B.Jack et Steph.) Steph., Sp. Hepat. (Stephani) 4: 787, 1912. Bas.: Thysanolejeunea gottschei J.B.Jack et Steph., Hedwigia 31 (1): 20, 1892.

** Thysananthus gottschei var. continuus Sukkharak, Phytotaxa 193 (1): 33, 2015.

*** Thysananthus spathulistipus (Reinw., Blume et Nees) Lindenb., Syn. Hepat. 2: 287, 1845. Bas.: Jungermannia spathulistipa Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 212, 1824 [1825].

** sect. Vittatae Verd., Ann. Bryol., Suppl. 4: 182, 1934.

** subsect. Sandeanthus (B.M.Thiers et Gradst.) Sukkharak, Phytotaxa 193 (1): 43, 2015. Bas.: Thysananthus subg. Sandeanthus B.M.Thiers et Gradst., Mem. New York Bot. Gard. 52: 66, 1989.

*** Thysananthus mollis Steph., Sp. Hepat. (Stephani) 4: 798, 1912.

*** Thysananthus montanus Gradst., Xiao L.He et Piippo, Acta Bot. Fenn. 174: 77, 2002.

*** Thysananthus retusus (Reinw., Blume et Nees) B.M.Thiers et Gradst., Mem. New York Bot. Gard. 52: 67, 1989. Bas.: Jungermannia retusa Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 214, 1824 [1825].

** Thysananthus retusus subsp. sellingii (Herzog) Sukkharak, Phytotaxa 193 (1): 47, 2015. Bas.: Mastigolejeunea sellingii Herzog, Ark. Bot. (n.ser.) 3 (3): 60, 1953.

** subsect. Vittatae (Verd.) Sukkharak, Phytotaxa 193 (1): 40, 2015. Bas.: Thysananthus sect. Vittatae Verd., Ann. Bryol., Suppl. 4: 182, 1934.

*** Thysananthus fruticosus (Lindenb. et Gottsche) Schiffn., Hepat. (Engl.-Prantl): 130, 1893. Bas.: Bryopteris fruticosa Lindenb. et Gottsche, Syn. Hepat. 5: 737, 1847.

** Tuzibeanthus S.Hatt., Biosphaera 1 (1): 7, 1947.

*** Tuzibeanthus chinensis (Steph.) Mizut., J. Hattori Bot. Lab. 24: 151, 1961. Bas.: Ptychanthus chinensis Steph., Sp. Hepat. (Stephani) 4: 744, 1911.

** Verdoornianthus Gradst., Bryologist 80 (4): 607, 1977 [1978].

*** Verdoornianthus griffinii Gradst., Bryologist 80 (4): 609, 1977 [1978].

*** Verdoornianthus marsupiifolius (Spruce) Gradst., Bryologist 80 (4): 609, 1977 [1978]. Bas.: Lejeunea marsupiifolia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 118, 1884.

*** Goebeliellaceae Verd.

by M. von Konrat and M.A.M. Renner

*** Goebeliella Steph., Hedwigia 51 (1): 61, 1911.

*** Goebeliella cornigera (Mitt.) Steph., Hedwigia 51 (1): 62, 1911. Bas.: Frullania cornigera Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 163, 1855.

*** Lepidolaenaceae Nakai

by M. von Konrat

The generic composition of Lepidolaenaceae follows, except that the genus Jubulopsis was reduced to a synonym of Lepidolaena by.

*** Gackstroemia Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877.

** subg. Gackstroemia

** Gackstroemia ljungneri (Herzog) Grolle, J. Hattori Bot. Lab. 30: 17, 1967. Bas.: Lepidolaena ljungneri Herzog, Ark. Bot. 29A (21): 6, 1940.

*** Gackstroemia magellanica (Lam.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877. Bas.: Jungermannia magellanica Lam., Encycl. (Lamarck) 3 (1): 284, 1789.

*** Gackstroemia novae-zelandiae R.M.Schust. et J.J.Engel, Phytotaxa 118 (1): 10, 2013.

*** Gackstroemia weindorferi (Herzog) Grolle, J. Hattori Bot. Lab. 30: 20, 1967. Bas.: Lepidolaena weindorferi Herzog, Ann. Bryol. 6: 103, 1933.

** subg. Hariotiella (Besch. et C.Massal. ex Schiffn.) Grolle, J. Hattori Bot. Lab. 30: 12, 1967. Bas.: Lepidolaena subg. Hariotiella Besch. et C.Massal. ex Schiffn., Hepat. (Engl.-Prantl): 110, 1893.

*** Gackstroemia hariotiana (Besch. et C.Massal.) Grolle, J. Hattori Bot. Lab. 30: 12, 1967. Bas.: Polyotus hariotianus Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 639, 1886.

*** Gackstroemia patagonica (Steph.) Grolle, J. Hattori Bot. Lab. 30: 14, 1967. Bas.: Lepidolaena patagonica Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 76, 1911.

** Gackstroemia schwabei (Herzog) Grolle, J. Hattori Bot. Lab. 30: 16, 1967. Bas.: Lepidolaena schwabei Herzog, Rev. Bryol. Lichénol. 29 (3/4): 191, 1960 [1961].

Incertae sedis

*** Gackstroemia alpina R.M.Schust., J. Hattori Bot. Lab. 36: 349, 1972 [1973].

*** Lepidogyna R.M.Schust., Phytologia 45 (5): 419, 1980.

*** Lepidogyna hodgsoniae (Grolle) R.M.Schust., Phytologia 45 (5): 419, 1980. Bas.: Lepidolaena hodgsoniae Grolle, J. Hattori Bot. Lab. 30: 29, 1967.

*** Lepidogyna menziesii (Hook.) R.M.Schust., Phytologia 45 (5): 419, 1980. Bas.: Jungermannia menziesii Hook., Musci Exot. 2: tab. 118, 1820.

*** Lepidolaena Dumort., Recueil Observ. Jungerm.: 13, 1835. 369

*** Lepidolaena berggrenii E.A.Hodgs., Trans. Roy. Soc. New Zealand 87 (3/4): 205, 1959.

** Lepidolaena brachyclada (Lehm.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877. Bas.: Frullania brachyclada Lehm., Nov. Stirp. Pug. 8: 21, 1844.

*** Lepidolaena clavigera (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877. Bas.: Jungermannia clavigera Hook., Musci Exot. 1: tab. 70, 1818.

*** Lepidolaena novae-zelandiae (E.A.Hodgs. et S.W.Arnell) von Konrat, L.Söderstr. et A.Hagborg, Phytotaxa 65: 51, 2012. Bas.: Jubula novae-zelandiae E.A.Hodgs. et S.W.Arnell, Trans. Roy. Soc. New Zealand, Bot. 3 (4): 90, 1965.

*** Lepidolaena palpebrifolia (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877. Bas.: Jungermannia palpebrifolia Hook., Musci Exot. 1: tab. 71, 1818.

*** Lepidolaena reticulata (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877. Bas.: Jungermannia reticulata Hook.f. et Taylor, London J. Bot. 3: 395, 1844.

*** Lepidolaena taylorii (Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877. Bas.: Polyotus taylorii Gottsche, Syn. Hepat. 2: 246, 1845.

*** Porellaceae Cavers nom. conserv.

*** Ascidiota C.Massal., Nuovo Giorn. Bot. Ital. (n.ser.) 5 (2): 256, 1898.

*** Ascidiota blepharophylla C.Massal., Nuovo Giorn. Bot. Ital. (n.ser.) 5 (2): 257, 1898.

** Ascidiota blepharophylla subsp. alaskana Steere et R.M.Schust., Bull. Torrey Bot. Club 87 (3): 213, 1960.

*** Porella L., Sp. Pl. 1: 1106, 1753.

** Porella abyssinica (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877. Bas.: Madotheca abyssinica Nees, Syn. Hepat. 2: 281, 1845.

** Porella abyssinica var. hoehnelii (Steph.) Pócs, Fragm. Florist. Geobot. 39 (1): 229, 1994. Bas.: Porella hoehnelii Steph., Hedwigia 30 (6): 266, 1891.

** Porella acutifolia (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877. Bas.: Madotheca acutifolia Lehm. et Lindenb., Nov. Stirp. Pug. 7: 8, 1838.

** Porella acutifolia var. hattoriana (Pócs) S.Hatt., Misc. Bryol. Lichenol. 8 (4): 79, 1979. Bas.: Porella plumosa var. hattoriana Pócs, J. Hattori Bot. Lab. 31: 82, 1968.

* Porella acutifolia var. linguifolia (Steph.) M.L.So, Syst. Bot. 27 (1): 5, 2002. Bas.: Madotheca linguifolia Steph., Sp. Hepat. (Stephani) 4: 291, 1910. 370

** Porella acutifolia subsp. tosana (Steph.) S.Hatt., J. Hattori Bot. Lab. 44: 100, 1978. Bas.: Madotheca tosana Steph., Bull. Herb. Boissier 5 (2): 97, 1897.

* Porella andica (Gottsche) Hässel, Beih. Nova Hedwigia 134: 452, 2009. Bas.: Madotheca andica Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 339, 1857. 371

*** Porella arboris-vitae (With.) Grolle, Trans. Brit. Bryol. Soc. 5 (4): 770, 1969. Bas.: Jungermannia arboris-vitae With., Bot. arr. veg. Gr. Brit. 2: 697, 1776.

** Porella arboris-vitae subsp. nitidula (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 40: 123, 1976. Bas.: Madotheca nitidula C.Massal., Bull. Soc. Bot. Ital. 1906: 141, 1906.

** Porella baueri (Schiffn.) C.E.O.Jensen, Danmarks mosser: 240, 1915. Bas.: Madotheca baueri Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 346, 1900.

** Porella bolanderi (Austin) Pearson, List. Canad. Hepat.: 7, 1890. Bas.: Madotheca bolanderi Austin, Bull. Torrey Bot. Club 3 (3): 14, 1872.

** Porella borellii (Gola) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 15, 1962. Bas.: Madotheca borellii Gola, Atti Reale Accad. Sci. Torino, Cl. Sci. Fis. Mat. Nat. 49: 760, 1914.

*** Porella brachiata (Taylor) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 334, 1885. Bas.: Madotheca brachiata Taylor, London J. Bot. 6: 341, 1847.

*** Porella brasiliensis (Raddi) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 246, 1893. Bas.: Schulthesia brasiliensis Raddi, Critt. Brasil.: 10, 1822.

* Porella brasiliensis var. ciliata (Gottsche, Lindenb. et Nees) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 246, 1893. Bas.: Madotheca brasiliensis β ciliata Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 271, 1845.

* Porella brasiliensis var. laevior (Gottsche, Lindenb. et Nees) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 246, 1893. Bas.: Madotheca brasiliensis α laevior Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 271, 1845.

** Porella caespitans (Steph.) S.Hatt., J. Hattori Bot. Lab. 33: 50, 1970. Bas.: Madotheca caespitans Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 218, 1894. 372

** Porella caespitans var. cordifolia (Steph.) S.Hatt. ex T.Katag. et T.Yamag., Bryol. Res. 10 (5): 133, 2011. Bas.: Madotheca cordifolia Steph., Sp. Hepat. (Stephani) 4: 315, 1910.

** Porella caespitans subsp. latior (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 40: 127, 1976. Bas.: Porella acutifolia subsp. latior S.Hatt., J. Hattori Bot. Lab. 32: 325, 1969.

** Porella caespitans var. nipponica S.Hatt., J. Hattori Bot. Lab. 33: 57, 1970.

** Porella caespitans var. reflexigastria (Pócs) S.Hatt., J. Hattori Bot. Lab. 40: 127, 1976. Bas.: Porella reflexigastria Pócs, J. Hattori Bot. Lab. 31: 71, 1968.

** Porella campylophylla (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877. Bas.: Jungermannia campylophylla Lehm. et Lindenb., Nov. Stirp. Pug. 6: 40, 1834.

** Porella campylophylla subsp. lancistipula (Steph.) S.Hatt., J. Hattori Bot. Lab. 44: 102, 1978. Bas.: Madotheca lancistipula Steph., Sp. Hepat. (Stephani) 6: 524, 1924.

** Porella campylophylla var. ligulifera (Taylor) S.Hatt., J. Hattori Bot. Lab. 32: 333, 1969. Bas.: Madotheca ligulifera Taylor, Nov. Stirp. Pug. 8: 10, 1844.

** Porella campylophylla var. tixieri (Pócs) S.Hatt., J. Hattori Bot. Lab. 40: 128, 1976. Bas.: Porella plumosa var. tixieri Pócs, J. Hattori Bot. Lab. 31: 82, 1968.

** Porella canariensis (F.Weber) Underw., Rep. (Annual) Missouri Bot. Gard. 8: 186, 1897. Bas.: Jungermannia platyphylla var. canariensis F.Weber, Hist. Musc. Hepat. Prodr.: 16, 1815.

* Porella capehorniensis Swails, Nova Hedwigia 19: 244, 1970 (Swails 1970). 373

** Porella capensis (Gottsche) Mitt., J. Linn. Soc., Bot. 22 (146): 323, 1886. Bas.: Madotheca capensis Gottsche, Syn. Hepat. 2: 270, 1845.

* Porella caucasica Steph., Bot. Centralbl. 50 (3): 71, 1892.

** Porella chenii S.Hatt., J. Hattori Bot. Lab. 30: 129, 1967.

*** Porella chilensis (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877. Bas.: Jungermannia chilensis Lehm. et Lindenb., Nov. Stirp. Pug. 6: 36, 1834.

* Porella chilensis var. antucensis (Gottsche) Hässel, Beih. Nova Hedwigia 134: 452, 2009. Bas.: Madotheca chilensis β antucensis Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 340, 1857.

** Porella chilensis var. fernandeziensis (Herzog) Swails, Nova Hedwigia 19: 236, 1970 (Swails 1970). Bas.: Madotheca chilensis f. fernandeziensis Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 736, 1942.

** Porella chilensis var. microloba (Herzog) Swails, Nova Hedwigia 19: 236, 1970 (Swails 1970). Bas.: Madotheca chilensis f. microloba Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 736, 1942.

** Porella chinensis (Steph.) S.Hatt., J. Hattori Bot. Lab. 30: 131, 1967. Bas.: Madotheca chinensis Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 218, 1894.

** Porella chinensis var. crispata Udar et Shaheen, Misc. Bryol. Lichenol. 9 (4): 74, 1982.

** Porella chinensis var. decurrens (Steph.) S.Hatt., J. Hattori Bot. Lab. 44: 102, 1978. Bas.: Madotheca decurrens Steph., Sp. Hepat. (Stephani) 4: 289, 1910.

** Porella chinensis var. hattorii Udar et Shaheen, Misc. Bryol. Lichenol. 9 (7): 146, 1983.

** Porella chinensis var. irregularis (Steph.) S.Hatt., J. Hattori Bot. Lab. 39: 270, 1975. Bas.: Madotheca irregularis Steph., Sp. Hepat. (Stephani) 4: 304, 1910.

** Porella circinnata Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 355, 1874.

*** Porella cordaeana (Huebener) Moore, Proc. Roy. Irish Acad. (ser. 2) 2: 618, 1877. Bas.: Jungermannia cordaeana Huebener, Hepaticol. germ.: 291, 1834.

** Porella cranfordii Steph., Hedwigia 28 (4): 270, 1889.

*** Porella crispata (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877. Bas.: Jungermannia crispata Hook., Pl. crypt. (Hooker): tab. 4b, 1816.

** Porella cucullistipula Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 38, 1893 [1894].

** Porella densifolia (Steph.) S.Hatt., J. Jap. Bot. 20: 109, 1944. Bas.: Madotheca densifolia Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 219, 1894. 374

** Porella densifolia subsp. andamana S.Hatt., J. Hattori Bot. Lab. 32: 346, 1969.

** Porella densifolia subsp. appendiculata (Steph.) S.Hatt., J. Hattori Bot. Lab. 32: 343, 1969. Bas.: Madotheca appendiculata Steph., Sp. Hepat. (Stephani) 4: 301, 1910.

** Porella densifolia var. oviloba (Steph.) N.Kitag., Acta Phytotax. Geobot. 19 (2/3): 64, 1962. Bas.: Madotheca oviloba Steph., Sp. Hepat. (Stephani) 4: 312, 1910.

** Porella densifolia var. paraphyllina (P.C.Chen) Pócs, J. Hattori Bot. Lab. 31: 84, 1968. Bas.: Madotheca paraphyllina P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 42, 1955.

** Porella densifolia var. pilosa S.Hatt. et K.C.Chang, Bull. Bot. Res., Harbin 8 (2): 44, 1988.

** Porella densifolia var. robusta (Steph.) S.Hatt., J. Hattori Bot. Lab. 32: 343, 1969. Bas.: Madotheca robusta Steph., Sp. Hepat. (Stephani) 4: 313, 1910.

*** Porella elegantula (Mont.) E.A.Hodgs., Svensk Bot. Tidskr. 42 (3): 277, 1948. Bas.: Madotheca elegantula Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 255, 1843.

** Porella faurieri (Steph.) S.Hatt., J. Jap. Bot. 20: 109, 1944. Bas.: Madotheca faurieri Steph., Sp. Hepat. (Stephani) 4: 315, 1910.

** Porella fengii P.C.Chen et S.Hatt., J. Hattori Bot. Lab. 30: 133, 1967.

** Porella geheebii (Steph.) S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 114, 1951. Bas.: Madotheca geheebii Steph., Sp. Hepat. (Stephani) 4: 290, 1910.

** Porella gracillima Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 202, 1891.

** Porella grandifolia (Steph.) S.Hatt., J. Hattori Bot. Lab. 30: 136, 1967. Bas.: Madotheca grandifolia Steph., Sp. Hepat. (Stephani) 4: 289, 1910.

** Porella grandiloba Lindb., Acta Soc. Sci. Fenn. 10: 234, 1872 [1873].

** Porella grollei S.Hatt., J. Hattori Bot. Lab. 34: 411, 1971.

** Porella handelii S.Hatt., J. Hattori Bot. Lab. 33: 65, 1970.

** Porella hattorii Udar et Shaheen, Lindbergia 9 (1): 70, 1983.

** Porella hoeana S.Hatt., Misc. Bryol. Lichenol. 7 (5): 86, 1976.

* Porella imbricata Lour., Fl. Cochinch. 2: 683, 1790.

** Porella inaequalis (Gottsche) Perss., Arch. Soc. Zool. Bot. Fenn. “Vanamo”, suppl. 9: 225, 1955. Bas.: Madotheca inaequalis Gottsche, Sp. Hepat. (Stephani) 4: 251, 1910.

** Porella japonica (Sande Lac.) Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 202, 1891. Bas.: Madotheca japonica Sande Lac., Syn. hepat. jav.: 105, 1856 [1857].

** Porella japonica subsp. appalachiana R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 682, 1980.

** Porella japonica var. densespinosa S.Hatt. et M.X.Zhang, J. Jap. Bot. 60 (11): 324, 1985.

** Porella javanica (Gottsche) Inoue, J. Hattori Bot. Lab. 30: 60, 1967. Bas.: Madotheca javanica Gottsche, Sp. Hepat. (Stephani) 4: 290, 1910.

** Porella latifolia J.S.Lou et Q.Li, Acta Phytotax. Sin. 25 (6): 482, 1987.

*** Porella leiboldii (Lehm.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877. Bas.: Madotheca leiboldii Lehm., Nov. Stirp. Pug. 8: 11, 1844.

** Porella longifolia (Steph.) S.Hatt., J. Hattori Bot. Lab. 32: 351, 1969. Bas.: Madotheca longifolia Steph., Sp. Hepat. (Stephani) 4: 305, 1910.

** Porella madagascariensis (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877. Bas.: Lejeunea madagascariensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 57, 1836.

* Porella maxima (Steph.) M.L.So, Syst. Bot. 27 (1): 11, 2002. Bas.: Madotheca maxima Steph., Sp. Hepat. (Stephani) 4: 291, 1910. 375

*** Porella mexicana (Hampe ex Gottsche, Lindenb. et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877. Bas.: Madotheca mexicana Hampe ex Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 273, 1845.

** Porella montantii (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 460, 1963. Bas.: Madotheca montantii Steph., Sp. Hepat. (Stephani) 4: 259, 1910.

*** Porella navicularis (Lehm. et Lindenb.) Pfeiff., Fl. Niederhessen 2: 234, 1855. Bas.: Jungermannia navicularis Lehm. et Lindenb., Nov. Stirp. Pug. 6: 38, 1834.

** Porella nitens (Steph.) S.Hatt., Fl. E. Himalaya: 525, 1966. Bas.: Madotheca nitens Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 220, 1894.

** Porella oblongifolia S.Hatt., J. Jap. Bot. 19 (7): 200, 1943. 376

*** Porella obtusata (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877. Bas.: Madotheca obtusata Taylor, London J. Bot. 5: 380, 1846.

** Porella obtusata var. macroloba (Steph.) S.Hatt. et M.X.Zhang, J. Jap. Bot. 60 (11): 325, 1985. Bas.: Madotheca macroloba Steph., Sp. Hepat. (Stephani) 4: 292, 1910.

** Porella obtusiloba S.Hatt., J. Hattori Bot. Lab. 33: 69, 1970.

** Porella perrottetiana (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877. Bas.: Madotheca perrottetiana Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 15, 1842.

** Porella perrottetiana var. angustifolia Pócs, J. Hattori Bot. Lab. 31: 75, 1968.

** Porella perrottetiana var. ciliatodentata (P.C.Chen et P.C.Wu) S.Hatt., J. Hattori Bot. Lab. 30: 144, 1967. Bas.: Porella ciliatodentata P.C.Chen et P.C.Wu, Obs. fl. Hwangs.: 8, 1965.

** Porella perrottetiana var. triciliata (Steph.) Pócs, J. Hattori Bot. Lab. 31: 75, 1968. Bas.: Madotheca triciliata Steph., Sp. Hepat. (Stephani) 4: 308, 1910.

*** Porella pinnata L., Sp. Pl. 1: 1106, 1753.

** Porella planifolia J.S.Lou, Coll. Pap. Quing-Zang Huang-Den 1: 277, 1983.

*** Porella platyphylla (L.) Pfeiff., Fl. Niederhessen 2: 234, 1855. Bas.: Jungermannia platyphylla L., Sp. Pl. 1: 1134, 1753.

*** Porella platyphylloidea (Schwein.) Lindb., Morgonbladet (Helsinki) 1876 (287, 10 Dec.): 1, 1876. Bas.: Jungermannia platyphylloidea Schwein., Spec. Fl. Amer. Crypt.: 9, 1821.

** Porella plicata J.S.Lou, Acta Phytotax. Sin. 18 (1): 119, 1980.

** Porella plumosa (Mitt.) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 17, 1962. Bas.: Madotheca plumosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 108, 1860 [1861].

** Porella prolixa (Gottsche) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 460, 1963. Bas.: Madotheca prolixa Gottsche, Sp. Hepat. (Stephani) 4: 260, 1910.

** Porella pulcherrima Herzog et S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12 (1): 34, 1986.

*** Porella reflexa (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877. Bas.: Jungermannia reflexa Lehm. et Lindenb., Nov. Stirp. Pug. 5: 5, 1833.

** Porella revoluta (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877. Bas.: Jungermannia revoluta Lehm. et Lindenb., Nov. Stirp. Pug. 4: 18, 1832.

** Porella revoluta var. propinqua (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 30: 148, 1967. Bas.: Madotheca propinqua C.Massal., Hepat. Shen-si: 27, 1897.

** Porella roellii Steph., Bot. Centralbl. 45: 203, 1891.

*** Porella saccata M.L.So, New Zealand J. Bot. 43 (1): 302, 2005.

** Porella sichuanensis S.Hatt. et K.C.Chang, Bull. Bot. Res., Harbin 8 (2): 43, 1988.

** Porella spinulosa (Steph.) S.Hatt., J. Hattori Bot. Lab. 33: 74, 1970. Bas.: Madotheca spinulosa Steph., Sp. Hepat. (Stephani) 6: 529, 1924.

*** Porella squamulifera (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877. Bas.: Madotheca squamulifera Taylor, London J. Bot. 5: 378, 1846.

** Porella stephaniana (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 5: 81, 1951. Bas.: Madotheca stephaniana C.Massal., Hepat. Shen-si: 23, 1897. 377

** Porella subdentata (Mitt.) Steph., Hedwigia 30 (5): 203, 1891. Bas.: Madotheca subdentata Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 167, 1863.

** Porella subdentata var. camerunensis E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 456, 1963.

** Porella subobtusa (Steph.) S.Hatt., J. Jap. Bot. 20: 111, 1944. Bas.: Madotheca subobtusa Steph., Sp. Hepat. (Stephani) 4: 311, 1910.

** Porella subparaphyllina J.S.Lou, Acta Phytotax. Sin. 25 (6): 483, 1987.

*** Porella subsquarrosa (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877. Bas.: Lejeunea subsquarrosa Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 57, 1836.

*** Porella swailsii Grolle, J. Bryol. 10 (3): 270, 1979. Nom. nov. pro Madotheca apiculata Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 197, 1955.

*** Porella swartziana (F.Weber) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877. Bas.: Jungermannia swartziana F.Weber, Hist. Musc. Hepat. Prodr.: 18, 1815.

** Porella triquetra (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 454, 1963. Bas.: Madotheca triquetra Steph., Bot. Jahrb. Syst. 20 (3): 321, 1895.

** Porella truncata J.S.Lou, Acta Phytotax. Sin. 18 (1): 119, 1980.

** Porella ulophylla (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 92, 1944. Bas.: Madotheca ulophylla Steph., Bull. Herb. Boissier 5 (2): 97, 1897.

** Porella undatorevoluta J.S.Lou, Acta Phytotax. Sin. 25 (6): 485, 1987.

** Porella urceolata S.Hatt., J. Hattori Bot. Lab. 33: 66, 1970.

** Porella urogea (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 32: 349, 1969. Bas.: Madotheca urogea C.Massal., Hepat. Shen-si: 28, 1897.

** Porella vallis-gratiae (Gottsche) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 450, 1963. Bas.: Madotheca vallis-gratiae Gottsche, Sp. Hepat. (Stephani) 4: 261, 1910.

** Porella variabilis (Kashyap et R.S.Chopra) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 17, 1962. Bas.: Madotheca variabilis Kashyap et R.S.Chopra, Liverworts W. Himal. 2: 33, 1932.

** Porella vernicosa Lindb., Acta Soc. Sci. Fenn. 10: 223, 1872 [1873].

** Porella viridissima (Mitt.) Grolle, J. Hattori Bot. Lab. 36: 83, 1972 [1973]. Bas.: Madotheca viridissima Mitt., Fl. vit.: 411, 1871 [1873].

** Porella wataugensis (Sull.) Underw. ex M.Howe, Bull. Torrey Bot. Club 24 (11): 519, 1897. Bas.: Madotheca wataugensis Sull., Musc. Hepat. U.S.: 700, 1856.

*** Radulaceae Müll.Frib.

by M.A.M. Renner

The treatment of Radulaceae follows. Taxonomic and nomenclatural notes can also be found in, ).

*** Radula Dumort., Commentat. Bot. (Dumortier): 112, 1822 nom. conserv.

*** subg. Amentuloradula Devos, M.A.M.Renner, Gradst., A.J.Shaw et Vanderp., Taxon 60 (6): 1630, 2011.

*** Radula amentulosa Mitt., Bonplandia 9 (24): 367, 1861.

*** Radula aneurismalis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 262, 1845. Bas.: Jungermannia aneurismalis Hook.f. et Taylor, London J. Bot. 4: 86, 1845.

** Radula ceylanica K.Yamada, J. Jap. Bot. 50 (12): 373, 1975.

*** Radula fissifolia Steph., Sp. Hepat. (Stephani) 6: 507, 1924.

*** Radula formosa (C.F.W.Meissn. ex Spreng.) Nees, Syn. Hepat. 2: 258, 1845. Bas.: Jungermannia formosa C.F.W.Meissn. ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827.

*** Radula helix (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 260, 1845. Bas.: Jungermannia helix Hook.f. et Taylor, London J. Bot. 3: 475, 1844.

*** Radula hicksiae K.Yamada, Cryptog. Bryol. Lichénol. 5 (1/2): 191, 1984.

*** Radula iwatsukii K.Yamada, J. Hattori Bot. Lab. 45: 275, 1979.

** Radula morobeana K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 358, 1989.

** Radula multiamentula E.A.Hodgs., Rec. Domin. Mus. 4 (11): 122, 1962.

*** Radula ornata E.A.Br. et Pócs, Telopea 9 (3): 436, 2001.

*** Radula physoloba Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 255, 1843.

*** Radula pseudoscripta M.A.M.Renner, New Zealand J. Bot. 44 (3): 340, 2006.

** Radula queenslandica K.Yamada, J. Hattori Bot. Lab. 62: 192, 1987.

*** Radula scariosa Mitt., Bonplandia 9 (24): 367, 1861.

*** Radula splendida M.A.M.Renner et Devos, Nova Hedwigia 90 (1/2): 113, 2010.

* Radula squarrosa K.Yamada, J. Jap. Bot. 65 (1): 1, 1990. 378

*** Radula thiersiae K.Yamada, J. Hattori Bot. Lab. 62: 198, 1987.

*** Radula uvifera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 258, 1845. Bas.: Jungermannia uvifera Hook.f. et Taylor, London J. Bot. 3: 292 [392], 1844.

* Radula vagans Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 85, 1911.

** Radula verrucosa K.Yamada, J. Hattori Bot. Lab. 45: 277, 1979.

*** subg. Cladoradula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 315, 1885.

** Radula auriculata Steph., Bull. Herb. Boissier 5 (2): 105, 1897.

* Radula bipinnata Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 166, 1863. 379

*** Radula boryana (F.Weber) Nees ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 13, 1842. Bas.: Jungermannia boryana F.Weber, Hist. Musc. Hepat. Prodr.: 58, 1815.

*** Radula campanigera Mont., London J. Bot. 3: 634, 1844.

*** Radula campanigera subsp. obiensis (S.Hatt.) K.Yamada, J. Hattori Bot. Lab. 45: 309, 1979. Bas.: Radula obiensis S.Hatt., Bull. Tokyo Sci. Mus. 11: 83, 1944.

** Radula chinensis Steph., Nuovo Giorn. Bot. Ital. (n.ser.) 13 (4): 355, 1906.

*** Radula gottscheana Taylor, London J. Bot. 5: 374, 1846.

*** Radula perrottetii Gottsche, Hedwigia 23 (10): 154, 1884.

** Radula tenax Lindb., Acta Soc. Sci. Fenn. 10: 492, 1875.

*** subg. Dactyloradula Devos, M.A.M.Renner, Gradst., A.J.Shaw et Vanderp., Taxon 60 (6): 1630, 2011.

*** Radula brunnea Steph., Sp. Hepat. (Stephani) 4: 232, 1910.

*** subg. Metaradula R.M.Schust., Phytologia 56 (2): 69, 1984.

** Radula acuminata Steph., Sp. Hepat. (Stephani) 4: 230, 1910. 380

** Radula aguirrei R.M.Schust., Phytotaxa 202 (1): 70, 2015. Based on: Radula aguirrei R.M.Schust., J. Hattori Bot. Lab. 70: 56, 1991, nom. inval.

** Radula anisotoma M.A.M.Renner, PhytoKeys 27: 30, 2013.

*** Radula assamica Steph., Hedwigia 23 (10): 151, 1884.

*** Radula australiana K.Yamada, J. Hattori Bot. Lab. 51: 323, 1982.

*** Radula buccinifera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 261, 1845. Bas.: Jungermannia buccinifera Hook.f. et Taylor, London J. Bot. 3: 580, 1844.

*** Radula demissa M.A.M.Renner, PhytoKeys 27: 53, 2013.

** Radula evansii Castle, Ann. Bryol. 11: 37, 1938.

** Radula flaccida Lindenb. et Gottsche, Syn. Hepat. 5: 726, 1847.

** Radula flaccida var. brachycalyx Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 321, 1885.

*** Radula forficata M.A.M.Renner, Austral. Syst. Bot. 26 (4): 307, 2013.

** Radula grevilleana Taylor, Ann. Mag. Nat. Hist. 20 (135): 380, 1847.

*** Radula imposita M.A.M.Renner, PhytoKeys 27: 65, 2013.

** Radula jovetiana K.Yamada, Cryptog. Bryol. Lichénol. 5 (1/2): 193, 1984.

** Radula kilgourii M.A.M.Renner, Austral. Syst. Bot. 26 (4): 313, 2013.

** Radula loriana Castle, J. Hattori Bot. Lab. 21: 6, 1959.

*** Radula mammosa Spruce, Mem. Torrey Bot. Club 1 (3): 127, 1890.

*** Radula mittenii Steph., Hedwigia 23 (10): 148, 1884.

*** Radula myriopoda M.A.M.Renner, Austral. Syst. Bot. 26 (4): 323, 2013.

*** Radula notabilis M.A.M.Renner, PhytoKeys 27: 77, 2013.

** Radula nymannii Steph., Sp. Hepat. (Stephani) 4: 229, 1910.

** Radula protensa Lindenb., Bot. Zeitung (Berlin) 6 (25): 462, 1848.

** Radula protensa var. erectilobula Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 247, 1893.

** Radula pseudoflaccida E.W.Jones, J. Bryol. 9 (4): 501, 1977 [1978].

** Radula psychosis M.A.M.Renner, Austral. Syst. Bot. 26 (4): 328, 2013.

*** Radula ratkowskiana K.Yamada, J. Jap. Bot. 59 (3): 94, 1984.

*** Radula robinsonii Steph., Sp. Hepat. (Stephani) 4: 214, 1910.

** Radula stenocalyx Mont., Ann. Sci. Nat. Bot. (sér. 4) 3 (5): 315, 1855.

*** Radula strangulata Hook.f. et Taylor, London J. Bot. 5: 377, 1846.

* Radula tjibodensis K.I.Goebel, Ann. Jard. Bot. Buitenzorg 7 (1): 53, 1888. 381

*** Radula ventricosa Steph., Sp. Hepat. (Stephani) 4: 187, 1910.

*** Radula yanoella R.M.Schust., Phytologia 56 (2): 72, 1984.

*** subg. Odontoradula K.Yamada, J. Hattori Bot. Lab. 45: 209, 1979.

** Radula acuta Mitt., Fl. vit.: 410, 1871 [1873].

** Radula allisonii Castle, Rev. Bryol. Lichénol. 31 (3/4): 148, 1962 [1963].

** Radula amoena Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 192, 1931.

** Radula anceps Sande Lac., Ned. Kruidk. Arch. 3: 419, 1854 [1855].

* Radula crenulata Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 21, 1890. 382

*** Radula cuspidata Steph., Sp. Hepat. (Stephani) 4: 156, 1910.

*** Radula decora Gottsche, Hedwigia 23 (10): 145, 1884.

** Radula emarginata K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 352, 1989.

** Radula kojana Steph., Bull. Herb. Boissier 5 (2): 105, 1897.

*** Radula lacerata Steph., Rev. Bryol. 35 (2): 33, 1908.

*** Radula novae-hollandiae Hampe, Nov. Stirp. Pug. 7: 24, 1838.

*** Radula ocellata K.Yamada, J. Hattori Bot. Lab. 45: 209, 1979.

*** Radula plicata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 154, 1854.

** Radula pugioniformis M.A.M.Renner, PhytoKeys 27: 84, 2013.

*** Radula pulchella Mitt., Hedwigia 23 (10): 149, 1884.

*** Radula retroflexa Taylor, London J. Bot. 5: 378, 1846.

*** Radula tasmanica Steph., Sp. Hepat. (Stephani) 4: 212, 1910.

*** Radula weymouthiana Steph., Sp. Hepat. (Stephani) 4: 190, 1910.

*** subg. Radula

*** Radula acutiloba Steph., Hedwigia 28 (4): 271, 1889. 383

** Radula appressa Mitt., Philos. Trans. 168: 397, 1879.

*** Radula aquilegia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 260, 1845. Bas.: Jungermannia aquilegia Hook.f. et Taylor, London J. Bot. 3: 291 [391], 1844.

** Radula australis Austin, Bot. Bull. (Hanover) 1 (7): 32, 1876.

** Radula borneensis Steph., Sp. Hepat. (Stephani) 4: 209, 1910.

** Radula caduca K.Yamada, J. Hattori Bot. Lab. 45: 225, 1979.

*** Radula carringtonii J.B.Jack, Flora 64 (25): 385, 1881.

*** Radula complanata (L.) Dumort., Syll. Jungerm. Europ.: 38, 1831. Bas.: Jungermannia complanata L., Sp. Pl. 1: 1133, 1753.

** Radula constricta Steph., Sp. Hepat. (Stephani) 6: 506, 1924.

** Radula evelynae K.Yamada, J. Jap. Bot. 50 (4): 115, 1975.

** Radula fendleri Gottsche, Hedwigia 23 (10): 146, 1884. 384

*** Radula grandis Steph., J. Linn. Soc., Bot. 29 (201): 271, 1892.

** Radula japonica Gottsche, Hedwigia 23 (10): 152, 1884.

** Radula javanica Gottsche, Syn. Hepat. 2: 257, 1845.

*** Radula jonesii Bouman, Dirkse et K.Yamada, J. Bryol. 15 (1): 161, 1988.

*** Radula lindenbergiana Gottsche ex C.Hartm., Handb. Skand. fl. (ed.9): 98, 1864.

** Radula madagascariensis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 349, 1882.

** Radula marojezica E.W.Jones, J. Bryol. 17 (2): 307, 1992.

* Radula multiflora Gottsche ex Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 20, 1890. 385

** Radula novoguineensis K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 360, 1989.

** Radula obconica Sull., Manual (Gray): 688, 1848.

** Radula obtusiloba Steph., Bull. Herb. Boissier 5 (2): 105, 1897.

** Radula obtusiloba subsp. polyclada (A.Evans) S.Hatt., J. Hattori Bot. Lab. 29: 275, 1966. Bas.: Radula polyclada A.Evans, Bull. Torrey Bot. Club 41 (12): 607, 1914 [1915].

* Radula oceania Castle, Rev. Bryol. Lichénol. 33 (3/4): 390, 1965. 386

** Radula oreopsis M.A.M.Renner, Telopea 17: 123, 2014.

** Radula portoricensis Steph., Hedwigia 27 (11/12): 298, 1888.

*** Radula prolifera Arnell, Ark. Bot. 13 (2): 12, 1913.

*** Radula quadrata Gottsche, Syn. Hepat. 2: 255, 1845.

*** Radula reflexa Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 255, 1843.

** Radula sharpii K.Yamada, J. Jap. Bot. 60 (9): 260, 1985.

** Radula sumatrana Steph., Sp. Hepat. (Stephani) 4: 204, 1910.

** Radula tokiensis Steph., Hedwigia 23 (10): 150, 1884.

*** Radula van-zantenii K.Yamada, J. Hattori Bot. Lab. 45: 260, 1979.

** Radula varilobula Castle, J. Hattori Bot. Lab. 21: 19, 1959.

*** Radula wichurae Steph., Sp. Hepat. (Stephani) 4: 168, 1910.

*** subg. Volutoradula Devos, M.A.M.Renner, Gradst., A.J.Shaw et Vanderp., Taxon 60 (6): 1629, 2011.

** Radula ankefinensis Gottsche, Hedwigia 23 (10): 152, 1884.

** Radula antilleana Castle, J. Hattori Bot. Lab. 21: 48, 1959.

** Radula comorensis Steph., Hedwigia 23 (9): 132, 1884.

** Radula cubensis K.Yamada, J. Hattori Bot. Lab. 54: 241, 1983.

** Radula diversifolia Steph., Sp. Hepat. (Stephani) 4: 212, 1910.

*** Radula eggersii K.Yamada, J. Hattori Bot. Lab. 82: 339, 1997.

** Radula episcia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 318, 1885.

** Radula floridana Castle, Rev. Bryol. Lichénol. 36 (1/2): 1, 1968 [1969].

** Radula fulvifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 261, 1845. Bas.: Jungermannia fulvifolia Hook.f. et Taylor, London J. Bot. 4: 85, 1845.

*** Radula hastata Steph., Sp. Hepat. (Stephani) 4: 163, 1910.

* Radula holstiana Steph., Bot. Jahrb. Syst. 20 (3): 320, 1895.

*** Radula holtii Spruce, J. Bot. 25: 209, 1887.

** Radula husnotii Castle, J. Hattori Bot. Lab. 21: 45, 1959.

** Radula inflexa Gottsche, Hedwigia 23 (10): 148, 1884.

** Radula kegelii Gottsche ex Steph., Hedwigia 23 (10): 152, 1884.

** Radula macroloba Steph., Bull. Soc. Roy. Bot. Belgique 31: 121, 1892.

** Radula mazarunensis K.Yamada, Trop. Bryol. 1: 38, 1989.

** Radula mexicana Lindenb. et Gottsche, Mexik. Leverm.: 150, 1863.

** Radula microloba Gottsche, Syn. Hepat. 2: 259, 1845.

** Radula neotropica Castle, J. Hattori Bot. Lab. 21: 31, 1959.

*** Radula nudicaulis Steph., Sp. Hepat. (Stephani) 4: 174, 1910.

* Radula nudicaulis var. delicatula P.Allorge et V.Allorge, Rev. Bryol. Lichénol. 19 (1/2): 106, 1950.

** Radula pocsii K.Yamada, J. Hattori Bot. Lab. 54: 245, 1983.

** Radula recubans Taylor, London J. Bot. 5: 376, 1846.

** Radula saccatiloba Steph., Hedwigia 23 (8): 129, 1884.

** Radula schaefer-verwimpii K.Yamada, J. Jap. Bot. 65 (1): 3, 1990.

** Radula schofieldiana K.Yamada, J. Hattori Bot. Lab. 82: 337, 1997.

** Radula stipatiflora Steph., Sp. Hepat. (Stephani) 4: 159, 1910.

** Radula striata Mitt., Hedwigia 23 (10): 155, 1884.

** Radula subinflata Lindenb. et Gottsche, Syn. Hepat. 5: 724, 1847.

** Radula sullivantii Austin, Hepat. bor.-amer.: 22, 1873.

** Radula tenera Mitt., Hedwigia 23 (10): 149, 1884.

*** Radula voluta Taylor, Syn. Hepat. 2: 255, 1845.

Incertae sedis

*** Radula acutangula Steph., Bull. Herb. Boissier 5 (10): 848, 1897.

** Radula angulata Steph., Hedwigia 23 (8): 114, 1884.

** Radula bogotensis Steph., Hedwigia 23 (8): 115, 1884.

*** Radula bolanderi Gottsche, Hedwigia 23 (10): 145, 1884.

** Radula boninensis Furuki et K.Yamada, J. Jap. Bot. 61 (10): 312, 1986.

** Radula brasilica K.Yamada, J. Hattori Bot. Lab. 74: 35, 1993.

** Radula caespitosa Steph., Hedwigia 27 (3/4): 107, 1888.

** Radula campanulata Lindenb. et Gottsche, Syn. Hepat. 2: 256, 1845.

** Radula castlei Grolle, Bryologist 73 (4): 662, 1970.

*** Radula cavifolia Hampe ex Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 259, 1845.

** Radula cochabambaensis K.Yamada, J. Hattori Bot. Lab. 74: 37, 1993.

** Radula conferta Lindenb. et Gottsche, Syn. Hepat. 5: 729, 1847.

*** Radula cordata Mitt., Fl. vit.: 410, 1871 [1873].

** Radula costaricensis Gottsche, J. Bot. 15: 226, 1877.

*** Radula curvilobula M.L.So, J. Hattori Bot. Lab. 98: 176, 2005.

* Radula decurrens Mitt., Fl. vit.: 410, 1871 [1873]. 387

** Radula densifolia Castle, Rev. Bryol. Lichénol. 33 (3/4): 385, 1965.

** Radula diaphana K.I.Goebel, Organogr. Pfl., ed. 2, 2 (1): 677, 1915.

** Radula dolabrata K.Yamada, J. Jap. Bot. 60 (9): 257, 1985.

** Radula elliottii Castle, J. Hattori Bot. Lab. 21: 12, 1959.

** Radula falcata Steph., Hedwigia 23 (8): 115, 1884.

** Radula fauriana Steph., Sp. Hepat. (Stephani) 4: 207, 1910.

** Radula fernandezana Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 84, 1911.

** Radula flavifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 259, 1845. Bas.: Jungermannia flavifolia Hook.f. et Taylor, London J. Bot. 3: 476, 1844.

** Radula fujitae Furuki, Bryol. Res. 9 (5): 143, 2007.

** Radula galapagona Steph., Sp. Hepat. (Stephani) 4: 176, 1910.

** Radula gedena Gottsche, Hedwigia 23 (10): 146, 1884.

** Radula gracilis Mitt., Hedwigia 23 (10): 147, 1884.

** Radula gradsteinii K.Yamada, Trop. Bryol. 1: 37, 1989.

** Radula grandifolia Steph., Sp. Hepat. (Stephani) 4: 184, 1910.

** Radula grollei K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 379, 1989.

** Radula guyanensis K.Yamada, Trop. Bryol. 1: 38, 1989.

** Radula hattorii K.Yamada, J. Jap. Bot. 60 (9): 259, 1985.

*** Radula hawaiica M.L.So, J. Hattori Bot. Lab. 98: 177, 2005.

* Radula hedingeri K.I.Goebel, Ann. Jard. Bot. Buitenzorg 7 (1): 51, 1888.

*** Radula inouei K.Yamada, J. Hattori Bot. Lab. 45: 262, 1979.

*** Radula involvens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 325, 1885.

*** Radula iwatsukiana K.Yamada, J. Hattori Bot. Lab. 58: 114, 1985.

*** Radula jamaicensis Pearson, Ann. Bryol. 4: 103, 1931.

** Radula jamesonii Taylor, London J. Bot. 5: 375, 1846.

** Radula kinabaluensis K.Yamada, Misc. Bryol. Lichenol. 6 (6): 97, 1973.

*** Radula kitagawae K.Yamada, J. Hattori Bot. Lab. 58: 116, 1985.

** Radula koponenii K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 364, 1989.

** Radula kurzii Steph., Hedwigia 23 (10): 153, 1884.

** Radula laxiramea Steph., Sp. Hepat. (Stephani) 4: 178, 1910.

** Radula leiboldii Steph., Hedwigia 23 (8): 116, 1884.

** Radula lewisii K.Yamada, J. Hattori Bot. Lab. 74: 39, 1993.

*** Radula ligula Steph., Sp. Hepat. (Stephani) 4: 228, 1910.

*** Radula lingulata Gottsche, Syn. Hepat. 2: 260, 1845.

** Radula longiloba K.Yamada, J. Hattori Bot. Lab. 54: 243, 1983.

* Radula longispica Steph., Sp. Hepat. (Stephani) 4: 183, 1910. 388

*** Radula marginata Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 261, 1845. Nom. nov. pro Jungermannia marginata Hook.f. et Taylor, London J. Bot. 3: 566, 1844, nom. illeg.

*** Radula mauiensis M.L.So, J. Hattori Bot. Lab. 98: 178, 2005.

** Radula microlobula Castle, J. Hattori Bot. Lab. 21: 35, 1959.

** Radula minutilobula K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 377, 1989.

** Radula mizutanii K.Yamada, J. Jap. Bot. 48 (5): 134, 1973.

*** Radula nigra Pearson, J. Linn. Soc., Bot. 46 (305): 31, 1922.

** Radula nilgiriensis Udar et D.Kumar, J. Indian Bot. Soc. 61: 177, 1982.

** Radula norrisii K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 374, 1989.

** Radula novivrieseana K.Yamada, J. Hattori Bot. Lab. 51: 326, 1982.

*** Radula novocaledonica Hürl. et K.Yamada, J. Jap. Bot. 54 (8): 238, 1979.

*** Radula novocaledoniensis K.Yamada, J. Hattori Bot. Lab. 58: 120, 1985.

** Radula obovata Castle, J. Hattori Bot. Lab. 21: 16, 1959.

** Radula obscura Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861].

** Radula okamurana Steph., Sp. Hepat. (Stephani) 4: 209, 1910.

** Radula onraedtii K.Yamada, Misc. Bryol. Lichenol. 8 (6): 113, 1979.

** Radula opaciuscula (Spruce) Castle, J. Hattori Bot. Lab. 21: 22, 1959. Bas.: Radula episcia var. opaciuscula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 319, 1885.

** Radula ovalilobula K.Yamada, J. Hattori Bot. Lab. 45: 257, 1979.

** Radula oyamensis Steph., Hedwigia 23 (10): 149, 1884.

* Radula paganii Castle, J. Hattori Bot. Lab. 21: 33, 1959. 389

*** Radula pallens (Sw.) Nees ex Mont., Voy. Amér. Mérid. 7 (2): 71, 1839. Bas.: Jungermannia pallens Sw., Prodr. (Swartz): 143, 1788.

*** Radula pandei Udar et Dh.Kumar, Lindbergia 9 (2): 133, 1983.

** Radula patens K.Yamada, Cryptog. Bryol. Lichénol. 5 (1/2): 197, 1984.

** Radula peruviana K.Yamada, Beih. Nova Hedwigia 88: 79, 1987.

** Radula philippinensis K.Yamada, J. Hattori Bot. Lab. 45: 299, 1979.

* Radula pinnulata Mitt., Fl. vit.: 410, 1871 [1873]. 390

** Radula pseudostachya Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 319, 1885.

** Radula punctata Steph., Hedwigia 23 (8): 135, 1884.

** Radula pusilla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 320, 1885.

*** Radula rhombiloba Steph., Sp. Hepat. (Stephani) 4: 204, 1910.

* Radula rupicola K.Yamada, J. Hattori Bot. Lab. 58: 124, 1985. 391

** Radula santacruziana K.Yamada et Gradst., Trop. Bryol. 4: 67, 1991.

** Radula silvestris Gottsche, Abh. Naturwiss. Vereins Bremen 7: 349, 1882.

** Radula sinskeana K.Yamada, J. Hattori Bot. Lab. 74: 41, 1993.

** Radula sinuata Gottsche ex Steph., Sp. Hepat. (Stephani) 4: 161, 1910.

* Radula socorana Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 475, 1947.

** Radula sonsonensis Steph., Sp. Hepat. (Stephani) 4: 201, 1910.

** Radula stellatogemmipara C.Gao et Y.H.Wu, Nova Hedwigia 80 (1/2): 239, 2005.

** Radula subsimplex Steph., Hedwigia 23 (8): 130, 1884.

** Radula subsquarrosa S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 15, 1957.

** Radula tabularis Steph., Hedwigia 23 (9): 131, 1884.

** Radula taylorii Steph., Hedwigia 23 (9): 133, 1884.

** Radula tectiloba Steph., Hedwigia 27 (11/12): 298, 1888.

** Radula tenuis K.Yamada, J. Hattori Bot. Lab. 54: 247, 1983.

** Radula underwoodii Castle, J. Hattori Bot. Lab. 21: 37, 1959.

** Radula venezuelensis K.Yamada, Misc. Bryol. Lichenol. 9 (6): 122, 1982.

*** Radula vieillardii Gottsche, Hedwigia 23 (10): 150, 1884.

** Radula visianica C.Massal., Ann. Bot. (Rome) 1 (4): 298, 1904.

** Radula vrieseana Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 305, 1864.

** Radula wrightii Castle, J. Hattori Bot. Lab. 21: 15, 1959.

** Radula xalapensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 56, 1836.

** Herzogianthaceae Stotler et Crand.-Stotl.

** Herzogianthus R.M.Schust., J. Hattori Bot. Lab. 23: 71, 1960 [1961].

* Herzogianthus sanguineus R.M.Schust., Phytologia 56 (7): 457, 1985. 392

*** Herzogianthus vaginatus (Herzog) R.M.Schust., J. Hattori Bot. Lab. 23: 71, 1960 [1961]. Bas.: Blepharostoma vaginatum Herzog, Trans. & Proc. Roy. Soc. New Zealand 65 (3): 355, 1936.

** Neotrichocoleaceae Inoue

** Neotrichocolea S.Hatt., J. Hattori Bot. Lab. 2: 9, 1947 [1948].

** Neotrichocolea bissetii (Mitt.) S.Hatt., J. Hattori Bot. Lab. 2: 10, 1947 [1948]. Bas.: Mastigophora bissetii Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 200, 1891.

** Trichocoleopsis S.Okamura, Bot. Mag. (Tokyo) 25 (293): 159, 1911.

** Trichocoleopsis sacculata (Mitt.) S.Okamura, Bot. Mag. (Tokyo) 25 (293): 159, 1911. Bas.: Blepharozia sacculata Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 200, 1891.

*** Ptilidiaceae H.Klinggr.

by L. Söderström

Ptilidiaceae was recently studied molecularly by showing the occurrence of a possibly undescribed cryptic species from the Himalayas.

*** Ptilidium Nees, Naturgesch. Eur. Leberm. 1: 95, 1833.

*** Ptilidium californicum (Austin) Pearson, List. Canad. Hepat.: 7, 1890. Bas.: Lepidozia californica Austin, Bull. Torrey Bot. Club 6 (3): 19, 1875.

*** Ptilidium ciliare (L.) Hampe, Prod. fl. hercyn.: 76, 1836. Bas.: Jungermannia ciliaris L., Sp. Pl. 1: 1134, 1753.

*** Ptilidium pulcherrimum (Weber) Vain., Meddel. Soc. Fauna Fl. Fenn. 3: 88, 1878. Bas.: Jungermannia pulcherrima Weber, Spic. Fl. Goett.: 150, 1778.

*** Aneuraceae H.Klinggr.

by M. Nebel

A molecular phylogeny of Aneuraceae was recently published by. Their study revealed Verdoorniaceae to be nested within Aneuraceae and Cryptothallus within Aneura (see also ). Nomenclatural and taxonomic notes can also be found in, and.

*** Aneura Dumort., Commentat. Bot. (Dumortier): 115, 1822. 393

* Aneura amboinensis Steph., Bull. Herb. Boissier 7 (9): 678 (219), 1899.

* Aneura augustae Steph., Sp. Hepat. (Stephani) 6: 430, 1923.

* Aneura biflora Colenso, Trans. & Proc. New Zealand Inst. 17: 262, 1885.

*** Aneura blasioides (Horik.) Furuki, J. Hattori Bot. Lab. 70: 311, 1991. Bas.: Riccardia blasioides Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 197, 1933.

** Aneura brasiliensis (Ångstr.) Steph., Hedwigia 32 (3): 137, 1893. Bas.: Pseudoneura brasiliensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 91, 1876 [1877].

* Aneura brevissima Steph., Sp. Hepat. (Stephani) 6: 21, 1917.

** Aneura cerebrata Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 185, 1970.

*** Aneura crateriformis Furuki et D.G.Long, J. Bryol. 18 (2): 281, 1994.

* Aneura crinita C.Massal., Bull. Soc. Bot. Ital. 1917 (8/9): 81, 1917. 394

** Aneura crumii L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 43, 2012. Nom. nov. pro Cryptothallus hirsutus H.A.Crum, Bryologist 99 (4): 438, 1996.

* Aneura densa Steph., Sp. Hepat. (Stephani) 6: 24, 1917.

* Aneura denticulata Mitt. ex Thurn, Timehri 5: 222, 1886. 395

** Aneura eachamensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 184, 1970.

** Aneura erronea Steph., Sp. Hepat. (Stephani) 6: 20, 1917.

** Aneura eskuchei Hässel, Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 294, 1986.

** Aneura gemmifera Furuki, J. Hattori Bot. Lab. 70: 309, 1991.

* Aneura giangena Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 190, 1970.

** Aneura gibbsiana Steph., Sp. Hepat. (Stephani) 6: 28, 1917.

** Aneura glaucescens Steph., Sp. Hepat. (Stephani) 6: 28, 1917.

* Aneura goebeliana Steph., Sp. Hepat. (Stephani) 6: 28, 1917.

*** Aneura hirsuta Furuki, J. Hattori Bot. Lab. 70: 317, 1991.

* Aneura hunsteinii Steph., Sp. Hepat. (Stephani) 6: 431, 1923.

** Aneura imbricata Colenso, Trans. & Proc. New Zealand Inst. 16: 359, 1884.

** Aneura kaguaensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 190, 1970.

** Aneura keniae Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 3, 1916.

* Aneura latemultifida Steph., Sp. Hepat. (Stephani) 6: 431, 1923.

** Aneura latissima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 544, 1885.

* Aneura ledermannii Steph., Sp. Hepat. (Stephani) 6: 431, 1923.

** Aneura macrostachya Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 550, 1885.

*** Aneura marianensis Furuki, Bryologist 97 (1): 87, 1994.

*** Aneura maxima (Schiffn.) Steph., Bull. Herb. Boissier 7 (10): 760 (270), 1899. Bas.: Riccardia maxima Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 177, 1898.

*** Aneura mirabilis (Malmb.) Wickett et Goffinet, Bot. J. Linn. Soc. 156 (1): 11, 2008. Bas.: Cryptothallus mirabilis Malmb., Ann. Bryol. 6: 122, 1933.

** Aneura novaecaledoniae R.M.Schust., Phytologia 56 (7): 451, 1985.

*** Aneura novaguineensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 189, 1970.

* Aneura nymannii Steph., Sp. Hepat. (Stephani) 6: 35, 1917.

** Aneura pellucida Colenso, Trans. & Proc. New Zealand Inst. 18: 252, 1886.

*** Aneura pinguis (L.) Dumort., Syll. Jungerm. Europ.: 86, 1831. Bas.: Jungermannia pinguis L., Sp. Pl. 1: 1136, 1753.

** Aneura polyantha Colenso, Trans. & Proc. New Zealand Inst. 17: 262, 1885.

** Aneura punctata Colenso, Trans. & Proc. New Zealand Inst. 18: 254, 1886.

** Aneura rodwayi Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 188, 1970.

* Aneura roraimensis Steph., Trans. Linn. Soc. London, Bot. 6 (1): 94, 1901.

** Aneura rotangicola Steph., Sp. Hepat. (Stephani) 6: 432, 1923.

* Aneura serrulata Gottsche ex Steph., Sp. Hepat. (Stephani) 6: 42, 1917.

** Aneura sharpii Inoue et N.G.Mill., Bull. Natl. Sci. Mus. Tokyo, B 11 (3): 96, 1985. 396

* Aneura singalangana (Schiffn.) Steph., Bull. Herb. Boissier 7 (10): 751 (261), 1899. Bas.: Riccardia singalangana Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 174, 1898. 397

** Aneura subcanaliculata R.M.Schust., J. Hattori Bot. Lab. 67: 60, 1989.

* Aneura subledermannii Steph., Sp. Hepat. (Stephani) 6: 432, 1923.

* Aneura subtenerrima Steph., Sp. Hepat. (Stephani) 6: 432, 1923.

* Aneura vincentina Steph., Sp. Hepat. (Stephani) 6: 45, 1917.

** Lobatiriccardia (Mizut. et S.Hatt.) Furuki, J. Hattori Bot. Lab. 70: 319, 1991. Bas.: Riccardia subg. Lobatiriccardia Mizut. et S.Hatt., J. Hattori Bot. Lab. 18: 38, 1957.

*** Lobatiriccardia alterniloba (Hook.f. et Taylor) Furuki, J. Hattori Bot. Lab. 70: 319, 1991. Bas.: Jungermannia alterniloba Hook.f. et Taylor, London J. Bot. 3: 572, 1844.

** Lobatiriccardia alterniloba var. gigantea (Steph.) Nebel, Phytotaxa 81 (1): 10, 2013. Bas.: Aneura gigantea Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 95, 1914.

** Lobatiriccardia alterniloba var. robusta (Rodway) Nebel, Phytotaxa 81 (1): 10, 2013. Bas.: Aneura alterniloba f. robusta Rodway, Tasm. Bryoph.: 12, 1917.

*** Lobatiriccardia athertonensis (Hewson) Furuki, J. Hattori Bot. Lab. 70: 319, 1991. Bas.: Aneura athertonensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 188, 1970.

*** Lobatiriccardia coronopus (De Not.) Furuki, J. Hattori Bot. Lab. 100: 90, 2006. Bas.: Aneura coronopus De Not., Hedwigia 32 (1): 19, 1893.

** Lobatiriccardia coronopus subsp. australis (R.M.Schust.) Nebel, Preussing, Schäf.-Verw. et D.Quandt, Taxon 59 (5): 1434, 2010. Bas.: Aneura lobata subsp. australis R.M.Schust., Phytologia 56 (7): 451, 1985.

** Lobatiriccardia oberwinkleri Nebel, Preussing, Schäf.-Verw. et D.Quandt, Taxon 59 (5): 1435, 2010.

** Lobatiriccardia subaquatica (R.M.Schust.) Nebel, Phytotaxa 81 (1): 10, 2013. Bas.: Aneura subaquatica R.M.Schust., Phytologia 56 (7): 450, 1985.

** Lobatiriccardia verdoornioides Nebel, Preussing, Schäf.-Verw. et D.Quandt, Taxon 59 (5): 1437, 2010.

*** Lobatiriccardia yakusimensis (S.Hatt.) Furuki, J. Hattori Bot. Lab. 70: 321, 1991. Bas.: Riccardia lobata var. yakusimensis S.Hatt., J. Hattori Bot. Lab. 6: 10, 1951 [1952].

*** Lobatiriccardia yunnanensis Furuki et D.G.Long, J. Bryol. 29 (3): 161, 2007.

*** Riccardia Gray, Nat. Arr. Brit. Pl. 1: 679, 1821 nom. conserv. 398

** Riccardia pectinata var. fasciculata (Steph.) Hürl., Bauhinia 5 (4): 208, 1976. Bas.: Aneura fasciculata Steph., Sp. Hepat. (Stephani) 6: 25, 1917.

** subg. Arceoneura Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 204, 1972.

** Riccardia marionensis R.M.Schust., J. Hattori Bot. Lab. 67: 65, 1989.

*** Riccardia prehensilis (Hook.f. et Taylor) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 255, 1885. Bas.: Jungermannia prehensilis Hook.f. et Taylor, London J. Bot. 3: 480, 1844.

** subg. Corioneura Furuki, J. Hattori Bot. Lab. 70: 394, 1991.

** Riccardia argentolimbata Hewson et Grolle, J. Hattori Bot. Lab. 29: 70, 1966.

*** Riccardia hattorii Furuki, J. Hattori Bot. Lab. 75: 257, 1994.

** subg. Hyaloneura R.M.Schust., Phytologia 56 (7): 452, 1985.

* Riccardia albomarginata (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 53, 1898. Bas.: Aneura albomarginata Steph., Hedwigia 32 (1): 18, 1893. 399

** Riccardia canaliculata (Nees) Kuntze, Revis. Gen. Pl. 2: 838, 1891. Bas.: Jungermannia canaliculata Nees, Enum. Pl. Crypt. Javae: 10, 1830.

** Riccardia pindensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 87, 1970.

** subg. Lophoneura Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 218, 1972.

** Riccardia fuegiensis C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 255, 1885.

** subg. Neoneura Furuki, J. Hattori Bot. Lab. 70: 385, 1991.

*** Riccardia spongiosa Furuki, J. Hattori Bot. Lab. 70: 385, 1991.

** subg. Phycaneura R.M.Schust., J. Hattori Bot. Lab. 26: 294, 1963.

** Riccardia aequicellularis (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 79, 1970. Bas.: Aneura aequicellularis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 95, 1914.

** Riccardia asperulata R.M.Schust., J. Hattori Bot. Lab. 27: 209, 1964.

** subg. Riccardia

*** Riccardia aeruginosa Furuki, J. Hattori Bot. Lab. 70: 345, 1991.

*** Riccardia arcuata Furuki, J. Hattori Bot. Lab. 70: 361, 1991.

*** Riccardia chamedryfolia (With.) Grolle, Trans. Brit. Bryol. Soc. 5 (4): 772, 1969. Bas.: Jungermannia chamedryfolia With., Bot. arr. veg. Gr. Brit. 2: 699, 1776.

*** Riccardia cochleata (Hook.f. et Taylor) Kuntze, Revis. Gen. Pl. 2: 838, 1891. Bas.: Riccia cochleata Hook.f. et Taylor, London J. Bot. 4: 96, 1845.

*** Riccardia eriocaula (Hook.) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 256, 1885. Bas.: Jungermannia eriocaula Hook., Musci Exot. 1: tab. 72, 1818.

*** Riccardia flavovirens Furuki, J. Hattori Bot. Lab. 70: 333, 1991.

*** Riccardia fruticosa (Steph.) Furuki, Nat. Hist. Res. 5 (1): 1, 1998. Bas.: Aneura fruticosa Steph., Sp. Hepat. (Stephani) 6: 27, 1917.

*** Riccardia glauca Furuki, J. Hattori Bot. Lab. 70: 352, 1991.

** Riccardia kodamae Mizut. et S.Hatt., J. Hattori Bot. Lab. 18: 57, 1957.

*** Riccardia latifrons (Lindb.) Lindb., Acta Soc. Sci. Fenn. 10: 513, 1875. Bas.: Aneura latifrons Lindb., Bot. Not. 26: 62, 1873.

** Riccardia latifrons subsp. arctica R.M.Schust. et Damsh., J. Hattori Bot. Lab. 62: 303, 1987.

** Riccardia latifrons var. miyakeana (Schiffn.) Furuki, J. Hattori Bot. Lab. 70: 375, 1991. Bas.: Riccardia miyakeana Schiffn., Österr. Bot. Z. 49 (11): 388, 1899.

** Riccardia nagasakiensis (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944. Bas.: Aneura nagasakiensis Steph., Sp. Hepat. (Stephani) 6: 34, 1917.

*** Riccardia palmata (Hedw.) Carruth., J. Bot. 3: 302, 1865. Bas.: Jungermannia palmata Hedw., Theoria generat.: 87, 1784, nom. conserv.

** Riccardia planiflora (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944. Bas.: Aneura planiflora Steph., Sp. Hepat. (Stephani) 6: 38, 1917.

** Riccardia planiflora var. aequatorialis Furuki, Nat. Hist. Res. 4 (2): 77, 1997.

** Riccardia pseudodendroceros R.M.Schust., Phytologia 56 (7): 452, 1985.

*** Riccardia pumila Furuki, J. Hattori Bot. Lab. 70: 361, 1991.

** Riccardia pusilla Grolle, J. Jap. Bot. 41 (8): 231, 1966. Nom. nov. pro Riccardia nana Mizut. et S.Hatt., J. Hattori Bot. Lab. 18: 53, 1957, nom. illeg.

*** Riccardia subalpina Furuki, J. Hattori Bot. Lab. 70: 367, 1991.

** Riccardia tamariscina (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 58, 1898. Bas.: Aneura tamariscina Steph., Hedwigia 32 (1): 27, 1893.

*** Riccardia vitrea Furuki, J. Hattori Bot. Lab. 70: 327, 1991.

** sect. Alcicornia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 17, 1972.

** Riccardia alcicornis (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877. Bas.: Jungermannia alcicornis Hook.f. et Taylor, London J. Bot. 3: 479, 1844.

** Riccardia conimitra (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 156, 1921. Bas.: Aneura conimitra Steph., Bull. Herb. Boissier 7 (10): 749 (259), 1899.

** Riccardia corralensis (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 143, 1921. Bas.: Aneura corralensis Steph., Bull. Herb. Boissier 7 (10): 742 (252), 1899.

*** Riccardia furtiva E.A.Br. et Braggins, J. Hattori Bot. Lab. 66: 35, 1989.

** Riccardia fuscobrunnea (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 152, 1921. Bas.: Aneura fuscobrunnea Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 7, 1911.

** Riccardia longioleata Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 30, 1972.

*** Riccardia multicorpora E.A.Br., J. Hattori Bot. Lab. 66: 40, 1989.

* Riccardia umida E.A.Br., J. Hattori Bot. Lab. 66: 38, 1989.

** sect. Crassantia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 41, 1972.

*** Riccardia aequitexta (Steph.) E.A.Br., J. Hattori Bot. Lab. 66: 78, 1989. Bas.: Aneura aequitexta Steph., J. Linn. Soc., Bot. 29 (201): 263, 1892.

*** Riccardia alba (Colenso) E.A.Br., J. Hattori Bot. Lab. 66: 66, 1989. Bas.: Aneura alba Colenso, Trans. & Proc. New Zealand Inst. 16: 357, 1884.

** Riccardia amnicola Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 92, 1972.

** Riccardia bipinnatifida (Colenso) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 93, 1970. Bas.: Aneura bipinnatifida Colenso, Trans. & Proc. New Zealand Inst. 16: 358, 1884.

** Riccardia calva (Schiffn.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 134, 1921. Bas.: Aneura calva Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 42, 1890.

*** Riccardia colensoi (Steph.) W.Martin, Trans. & Proc. Roy. Soc. New Zealand 78 (4): 499, 1950. Bas.: Aneura colensoi Steph., J. Linn. Soc., Bot. 29 (201): 264, 1892.

*** Riccardia crassa (Schwägr.) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 254, 1885. Bas.: Jungermannia crassa Schwägr., Hist. Musc. Hepat. Prodr.: 31, 1814.

** Riccardia crassicrispa (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 194, 1921. Bas.: Aneura crassicrispa Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 6, 1911.

** Riccardia diderma Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 56, 1972.

** Riccardia diversiflora A.Evans, Trans. Connecticut Acad. Arts 25 (2): 167, 1921.

** Riccardia diversiflora subsp. paucigyna R.M.Schust., J. Hattori Bot. Lab. 67: 97, 1989.

** Riccardia duriuscula Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 82, 1972.

** Riccardia falsifloribunda Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 119, 1972.

** Riccardia floribunda (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 182, 1921. Bas.: Aneura floribunda Steph., Bull. Herb. Boissier 7 (10): 749 (259), 1899.

** Riccardia fluvigena Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 73, 1972.

** Riccardia georgiensis (Steph.) Hässel, Lindbergia 1 (1/2): 80, 1971 [1972]. Bas.: Aneura georgiensis Steph., Wiss. Ergebn. Schwed. Südpolar-Exped. [1901–1903] 4 (1): 2, 1905.

** Riccardia georgiensis subsp. sympodea R.M.Schust., J. Hattori Bot. Lab. 67: 83, 1989.

*** Riccardia graeffei (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 118, 1970. Bas.: Aneura graeffei Steph., Hedwigia 32 (1): 21, 1893.

*** Riccardia marginata (Colenso) Pearson, Univ. Calif. Publ. Bot. 10 (4): 309, 1923. Bas.: Aneura marginata Colenso, Trans. & Proc. New Zealand Inst. 18: 253, 1886.

*** Riccardia marginata var. pacifica Furuki, J. Hattori Bot. Lab. 70: 380, 1991.

** Riccardia mycophora A.Evans, Trans. Connecticut Acad. Arts 25 (2): 175, 1921.

** Riccardia negeri (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 172, 1921. Bas.: Aneura negeri Steph., Bull. Herb. Boissier 7 (10): 747 (257), 1899.

** Riccardia nitida (Colenso) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 92, 1965. Bas.: Aneura nitida Colenso, Trans. & Proc. New Zealand Inst. 18: 253, 1886.

** Riccardia papillosa (C.Massal. et Steph.) Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 60, 1972. Bas.: Aneura papillosa C.Massal. et Steph., Atti Reale Ist. Veneto Sci. Lett. Arti 87 (2): 241, 1928.

** Riccardia regularis (Steph.) Kühnem., Revista Centro Estud. Doct. Ci. Nat. 1: 171, 1937. Bas.: Aneura regularis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 9, 1911.

** Riccardia rivularis Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 52, 1972.

** Riccardia saxicola Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 137, 1972.

** Riccardia spectabilis (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 140, 1921. Bas.: Aneura spectabilis Steph., Bull. Herb. Boissier 7 (10): 746 (256), 1899.

** Riccardia spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 254, 1885.

** Riccardia tenax (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 186, 1921. Bas.: Aneura tenax Steph., Bull. Herb. Boissier 7 (10): 755 (265), 1899.

** Riccardia tenerrima (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 164, 1921. Bas.: Aneura tenerrima Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 9, 1911.

** Riccardia theliophora Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 96, 1972.

** sect. Pallidevirida Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 142, 1972.

** Riccardia granulata (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 192, 1921. Bas.: Aneura granulata Steph., Hedwigia 32 (1): 21, 1893.

** Riccardia intercellula E.A.Br., J. Hattori Bot. Lab. 66: 58, 1989.

** Riccardia opuntiiformis S.W.Arnell, Arch. Soc. Zool. Bot. Fenn. “Vanamo” 9: 52, 1954.

** Riccardia pallidevirens (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 189, 1921. Bas.: Aneura pallidevirens Steph., Bull. Herb. Boissier 7 (10): 762 (272), 1899.

*** Riccardia pennata E.A.Br., J. Hattori Bot. Lab. 66: 53, 1989.

*** Riccardia perspicua E.A.Br., J. Hattori Bot. Lab. 66: 55, 1989.

** Riccardia xylophila Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 156, 1972.

** sect. Riccardia

*** Riccardia australis (Lehm.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 110, 1970. Bas.: Sarcomitrium australe Lehm., Nov. Stirp. Pug. 10: 19, 1857.

** Riccardia autoica (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 159, 1921. Bas.: Aneura autoica Steph., Bull. Herb. Boissier 7 (9): 691 (232), 1899.

** Riccardia breviala E.A.Br., J. Hattori Bot. Lab. 66: 46, 1989.

** Riccardia breviramosa (Steph.) A.Evans, Nat. Hist. Juan Fernandez (Botany) 2 (20): 559, 1930. Bas.: Aneura breviramosa Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 6, 1911.

** Riccardia filicina (Colenso) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 129, 1962. Bas.: Aneura filicina Colenso, Trans. & Proc. New Zealand Inst. 16: 358, 1884.

** Riccardia leptostachya A.Evans, Nat. Hist. Juan Fernandez (Botany) 2 (20): 570, 1930.

** Riccardia lobulata (Colenso) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 92, 1965. Bas.: Zoopsis lobulata Colenso, Trans. & Proc. New Zealand Inst. 18: 250, 1886.

** Riccardia mejlandii S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 30, 1958.

*** Riccardia multifida (L.) Gray, Nat. Arr. Brit. Pl. 1: 684, 1821. Bas.: Jungermannia multifida L., Sp. Pl. 1: 1136, 1753.

*** Riccardia multifida subsp. decrescens (Steph.) Furuki, J. Hattori Bot. Lab. 70: 341, 1991. Bas.: Aneura decrescens Steph., Bull. Herb. Boissier 7 (9): 686 (227), 1899.

** Riccardia multifida subsp. synoica R.M.Schust., J. Hattori Bot. Lab. 62: 319, 1987.

** Riccardia multioleata Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 195, 1972.

*** Riccardia papulosa (Steph.) E.A.Br., J. Hattori Bot. Lab. 66: 49, 1989. Bas.: Aneura papulosa Steph., Hedwigia 32 (1): 25, 1893.

** Riccardia patens Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 165, 1972.

** Riccardia polyclada (Mitt.) Hässel, Bryologist 109 (1): 34, 2006. Bas.: Aneura polyclada Mitt., Trans. Linn. Soc. London, Bot. 2 (13): 297, 1887.

** Riccardia statensis Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 177, 1972.

** Riccardia thaxteri A.Evans, Trans. Connecticut Acad. Arts 25 (2): 126, 1921.

** Riccardia tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 31, 1958.

** subg. Spinella (Schiffn.) Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 214, 1972. Bas.: Spinella Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 41, 1890.

** Riccardia spinulifera C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 254, 1885.

** subg. Thornoneura Furuki, J. Hattori Bot. Lab. 70: 382, 1991.

* Riccardia baumannii Hürl., Bauhinia 5 (4): 203, 1976. 400

** Riccardia deguchii Furuki et K.T.Yong, Hikobia 16 (3): 285, 2013.

*** Riccardia grossitexta (Steph.) Furuki, J. Hattori Bot. Lab. 70: 382, 1991. Bas.: Aneura grossitexta Steph., Sp. Hepat. (Stephani) 6: 29, 1917.

** Riccardia inconspicua (Steph.) Reeb et Bardat, Cryptog. Bryol. 35 (1): 61, 2014. Bas.: Aneura inconspicua Steph., Hedwigia 32 (1): 23, 1893.

** subg. Trichothallia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 224, 1972.

** Riccardia hyalitricha Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 225, 1972.

Incertae sedis

** Riccardia aberrans (Steph.) Gradst., J. Hattori Bot. Lab. 45: 129, 1979. Bas.: Aneura aberrans Steph., Hedwigia 32 (1): 18, 1893.

** Riccardia agumana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 82, 1970.

** Riccardia algoides (Taylor) Meenks, J. Hattori Bot. Lab. 62: 168, 1987. Bas.: Metzgeria algoides Taylor, London J. Bot. 5: 410, 1846.

*** Riccardia amazonica (Spruce) Schiffn. ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 129, 1979. Bas.: Aneura amazonica Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 545, 1885.

*** Riccardia andina (Spruce) Herzog, Svensk Bot. Tidskr. 46 (1): 65, 1952. Bas.: Aneura andina Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 548, 1885.

** Riccardia angustata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 126, 1934.

** Riccardia angustealata (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 111, 1970. Bas.: Aneura angustealata Steph., Sp. Hepat. (Stephani) 6: 20, 1917.

** Riccardia angustissima (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981. Bas.: Aneura angustissima Steph., Sp. Hepat. (Stephani) 6: 20, 1917.

** Riccardia aspera (Steph.) Grolle, J. Hattori Bot. Lab. 30: 117, 1967. Bas.: Aneura aspera Steph., Sp. Hepat. (Stephani) 6: 21, 1917.

* Riccardia baldwinii (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963. Bas.: Aneura baldwinii Steph., Bull. Herb. Boissier 7 (10): 743 (253), 1899. 401

** Riccardia barbiflora (Steph.) Piippo, J. Hattori Bot. Lab. 68: 134, 1990. Bas.: Aneura barbiflora Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 209, 1894.

** Riccardia bogotensis (Gottsche) Pagán, Bryologist 42 (1): 6, 1939. Bas.: Pseudoneura bogotensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 184, 1864.

** Riccardia boliviensis (Steph.) Meenks, J. Hattori Bot. Lab. 62: 170, 1987. Bas.: Aneura boliviensis Steph., Biblioth. Bot. 87 (2): 176, 1916.

** Riccardia bongeriana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 101, 1970.

** Riccardia brunnea (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944. Bas.: Aneura brunnea Steph., Sp. Hepat. (Stephani) 6: 21, 1917.

** Riccardia calcarea (Steph.) Meenks, J. Hattori Bot. Lab. 62: 170, 1987. Bas.: Aneura calcarea Steph., Bull. Herb. Boissier 7 (10): 756 (266), 1899.

** Riccardia capillacea (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 6, 1985. Bas.: Aneura capillacea Steph., Biblioth. Bot. 87 (2): 174, 1916.

** Riccardia capillacea var. dentata Meenks, J. Hattori Bot. Lab. 62: 170, 1987.

** Riccardia cardotii (Steph.) Pandé et S.C.Srivast., Biol. Mem. 1 (1/2): 131, 1976. Bas.: Aneura cardotii Steph., Bull. Soc. Roy. Bot. Belgique, Mém. 41 (1): 118, 1904 [1905].

** Riccardia cataractarum (Spruce) Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 18, 1964. Bas.: Aneura cataractarum Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcv, 1889 [1890].

*** Riccardia cervicornis (Spruce) Herzog ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 129, 1979. Bas.: Aneura cervicornis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 550, 1885.

** Riccardia changbaishanensis C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 209, 1981.

** Riccardia chinensis C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 209, 1981.

** Riccardia ciliolata (Spruce) Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 198, 1933. Bas.: Aneura ciliolata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 547, 1885.

** Riccardia columbica (Steph.) Hässel ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 130, 1979. Bas.: Aneura columbica Steph., Sp. Hepat. (Stephani) 6: 22, 1917.

** Riccardia comata (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981. Bas.: Aneura comata Steph., Sp. Hepat. (Stephani) 6: 22, 1917.

** Riccardia compacta (Steph.) S.W.Arnell, Bot. Not. 105: 141, 1952. Bas.: Aneura compacta Steph., Hedwigia 32 (1): 19, 1893.

** Riccardia comptonii (Pearson) H.A.Mill., Phytologia 47 (4): 323, 1981. Bas.: Aneura comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 17, 1922.

** Riccardia costata (Steph.) Hürl., Bauhinia 5 (4): 206, 1976. Bas.: Aneura costata Steph., Sp. Hepat. (Stephani) 6: 23, 1917.

*** Riccardia crassicaulis (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 8, 1985. Bas.: Aneura crassicaulis Steph., Biblioth. Bot. 87 (2): 174, 1916.

** Riccardia crassiretis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 173, 1898.

** Riccardia crenulata Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 173, 1898.

** Riccardia crenuliformis R.M.Schust., J. Hattori Bot. Lab. 67: 72, 1989.

* Riccardia decolyana Schiffn., J. Indian Bot. Soc. 38 (4): 538, 1959 [1960]. 402

** Riccardia densiramea (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944. Bas.: Aneura densiramea Steph., Sp. Hepat. (Stephani) 6: 24, 1917.

** Riccardia devexa Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 12, 1964.

** Riccardia diablotina (Spruce) Pagán, Bryologist 45 (4): 80, 1942. Bas.: Aneura diablotina Spruce, J. Linn. Soc., Bot. 30 (210): 366, 1895.

*** Riccardia digitiloba (Spruce) Pagán, Bryologist 42 (1): 6, 1939. Bas.: Aneura digitiloba Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cci, 1889 [1890].

** Riccardia dilatata (Spruce) Schäf.-Verw. et Pócs, Cryptog. Bryol. 31 (4): 389, 2010. Bas.: Aneura dilatata Spruce, J. Linn. Soc., Bot. 30 (210): 368, 1895.

* Riccardia diminuta Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 170, 1898. 403

* Riccardia diminuta var. thermarum Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898.

** Riccardia distans (Spruce) Pagán, Bryologist 45 (4): 80, 1942. Bas.: Aneura distans Spruce, J. Linn. Soc., Bot. 30 (210): 367, 1895.

** Riccardia elata (Steph.) Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 169, 1898. Bas.: Aneura elata Steph., Hedwigia 32 (1): 19, 1893.

* Riccardia elata var. flaccida Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 170, 1898.

* Riccardia elata var. intercedens Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 170, 1898.

** Riccardia elegans (Steph.) Hürl., Bauhinia 5 (4): 196, 1976. Bas.: Aneura elegans Steph., Sp. Hepat. (Stephani) 6: 25, 1917.

** Riccardia elisabethae Thouvenot et Reeb, Telopea 17: 229, 2014.

*** Riccardia emarginata (Steph.) K.G.Hell, Bol. Fac. Filos. Univ. São Paulo, Bot. 25: 100, 1969. Bas.: Aneura emarginata Steph., Hedwigia 32 (1): 20, 1893.

** Riccardia erosa (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 3 (1): 83, 1956. Bas.: Aneura erosa Steph., Hedwigia 30 (6): 269, 1891.

** Riccardia fastigiata (Lehm.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877. Bas.: Jungermannia fastigiata Lehm., Linnaea 4: 370, 1829.

** Riccardia fendleri (Steph.) Pagán, Bryologist 45 (4): 80, 1942. Bas.: Aneura fendleri Steph., Hedwigia 32 (1): 20, 1893.

** Riccardia flaccida (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944. Bas.: Aneura flaccida Steph., Sp. Hepat. (Stephani) 6: 26, 1917.

** Riccardia flaccidissima Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 167, 1898.

** Riccardia flagellaris (A.Gepp) H.A.Mill., Phytologia 47 (4): 323, 1981. Bas.: Aneura flagellaris A.Gepp, J. Linn. Soc., Bot. 39 (270): 194, 1909.

** Riccardia flagellifrons C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 209, 1981.

** Riccardia fleischeri (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963. Bas.: Aneura fleischeri Steph., Sp. Hepat. (Stephani) 6: 26, 1917.

** Riccardia foliacea Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 8, 1985.

** Riccardia formosensis (Steph.) Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 125, 1934. Bas.: Aneura formosensis Steph., Sp. Hepat. (Stephani) 6: 27, 1917.

*** Riccardia fucoidea (Sw.) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 256, 1885. Bas.: Jungermannia fucoidea Sw., Prodr. (Swartz): 145, 1788.

** Riccardia geniana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 76, 1970.

*** Riccardia glaziovii (Spruce) Meenks, J. Hattori Bot. Lab. 62: 173, 1987. Bas.: Aneura glaziovii Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cci, 1889 [1890].

** Riccardia gogolensis (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 102, 1970. Bas.: Aneura gogolensis Steph., Bull. Herb. Boissier 7 (9): 689 (230), 1899.

** Riccardia gracilis (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 295, 1963. Bas.: Aneura gracilis Steph., Bull. Herb. Boissier 7 (10): 752 (262), 1899.

** Riccardia grandiflora (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963. Bas.: Aneura grandiflora Steph., Sp. Hepat. (Stephani) 6: 29, 1917.

*** Riccardia grollei Furuki, Haussknechtia, Beih. 9: 139, 1999.

** Riccardia grossidens (Steph.) Pagán, Bryologist 45 (4): 80, 1942. Bas.: Aneura grossidens Steph., Hedwigia 32 (1): 23, 1893.

** Riccardia gunniana (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981. Bas.: Aneura gunniana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 96, 1914.

** Riccardia hamatiflora (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 526, 1963. Bas.: Aneura hamatiflora Steph., Bull. Herb. Boissier 5 (10): 844, 1897.

*** Riccardia hans-meyeri (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 12, 1985. Bas.: Aneura hans-meyeri Steph., Sp. Hepat. (Stephani) 6: 29, 1917.

** Riccardia hans-meyeri var. dentata Meenks, J. Hattori Bot. Lab. 62: 173, 1987.

** Riccardia hawaica (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963. Bas.: Aneura hawaica Steph., Sp. Hepat. (Stephani) 6: 30, 1917.

** Riccardia hebridensis (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981. Bas.: Aneura hebridensis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 96, 1914.

** Riccardia herzogiana (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 12, 1985. Bas.: Aneura herzogiana Steph., Biblioth. Bot. 87 (2): 175, 1916.

** Riccardia heteroclada Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 175, 1898.

* Riccardia hirtiflora (Steph.) Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 19, 1964. Bas.: Aneura hirtiflora Steph., Arch. Mus. Nac. Rio de Janeiro 13: 116, 1905. 404

** Riccardia humilis (Gottsche) O.Yano, J. Hattori Bot. Lab. 56: 530, 1984. Bas.: Pseudoneura humilis Gottsche, Mexik. Leverm.: 260, 1863.

** Riccardia hyalina (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981. Bas.: Aneura hyalina Steph., Sp. Hepat. (Stephani) 6: 31, 1917.

** Riccardia hydra Hürl., Bauhinia 5 (4): 210, 1976.

** Riccardia hymenophylloides Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 175, 1898.

* Riccardia hymenophylloides var. flaccida Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 176, 1898.

*** Riccardia hymenophytoides (Spruce) Meenks, Beih. Nova Hedwigia 88: 101, 1987. Bas.: Aneura hymenophytoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 549, 1885.

** Riccardia hypipamensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 97, 1970.

** Riccardia ibana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 78, 1970.

*** Riccardia incurvata Lindb., Helsingf. Dagbl. 1878 (315, 18 Nov.): 2, 1878.

** Riccardia innovans (Steph.) Pagán, Bryologist 45 (4): 80, 1942. Bas.: Aneura innovans Steph., Symb. Antill. 2: 470, 1901.

** Riccardia insularis Schiffn., Deutsche Südpolar-Exped. 1901-1903, 8 (bot.) 1: 66, 1906.

** Riccardia intricata (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981. Bas.: Aneura intricata Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 301, 1896.

** Riccardia jackii Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898.

* Riccardia jackii var. densa Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898.

** Riccardia judithae Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 12, 1985.

*** Riccardia jugata R.M.Schust., J. Hattori Bot. Lab. 62: 305, 1987.

* Riccardia karstenii (Steph.) Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 177, 1898. Bas.: Aneura karstenii Steph., Hedwigia 32 (1): 23, 1893.

** Riccardia laticostata (Spruce) Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 12, 1964. Bas.: Aneura laticostata Spruce, J. Linn. Soc., Bot. 30 (210): 367, 1895.

** Riccardia latifrondoides Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 168, 1898.

** Riccardia lepidomitra (Spruce) Gradst., J. Hattori Bot. Lab. 45: 130, 1979. Bas.: Aneura lepidomitra Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 549, 1885.

** Riccardia leptophylla (Spruce) Herzog, Svensk Bot. Tidskr. 46 (1): 65, 1952. Bas.: Aneura leptophylla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 544, 1885.

** Riccardia leptothallus R.M.Schust., J. Hattori Bot. Lab. 67: 87, 1989.

** Riccardia levieri Schiffn., Österr. Bot. Z. 49 (4): 130, 1899.

** Riccardia lichenoides (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981. Bas.: Aneura lichenoides Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 301, 1896.

** Riccardia ligulata (Steph.) Pócs et Schäf.-Verw., Cryptog. Bryol. 31 (4): 390, 2010. Bas.: Aneura ligulata Steph., Sp. Hepat. (Stephani) 6: 32, 1917.

** Riccardia lilliena (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963. Bas.: Aneura lilliena Steph., Sp. Hepat. (Stephani) 6: 33, 1917.

** Riccardia limbata (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 3 (1): 79, 1956. Bas.: Aneura limbata Steph., Hedwigia 30 (5): 203, 1891.

** Riccardia loefgrenii Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 14, 1964.

** Riccardia longiflora (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 88, 1970. Bas.: Aneura longiflora Steph., Bull. Herb. Boissier 7 (10): 746 (256), 1899.

** Riccardia longispica (Steph.) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 4, 1893. Bas.: Aneura longispica Steph., Bot. Gaz. 15 (11): 281, 1890.

** Riccardia loriana (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963. Bas.: Aneura loriana Steph., Bull. Herb. Boissier 7 (10): 733 (243), 1899.

** Riccardia macdonaldiana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 93, 1970.

** Riccardia macrantha (Pearson) H.A.Mill., Phytologia 47 (4): 324, 1981. Bas.: Aneura macrantha Pearson, J. Linn. Soc., Bot. 46 (305): 17, 1922.

*** Riccardia magnicellularis Furuki, J. Hattori Bot. Lab. 100: 93, 2006.

*** Riccardia metzgeriiformis (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 295, 1963. Bas.: Aneura metzgeriiformis Steph., Bull. Herb. Boissier 7 (10): 753 (263), 1899.

** Riccardia microscopica (Nees) Kuntze, Revis. Gen. Pl. 2: 838, 1891. Bas.: Aneura microscopica Nees, Syn. Hepat. 4: 500, 1846.

** Riccardia minuta (Steph.) W.Martin, Trans. & Proc. Roy. Soc. New Zealand 78 (4): 499, 1950. Bas.: Aneura minuta Steph., Sp. Hepat. (Stephani) 6: 34, 1917.

* Riccardia multifidoides Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 166, 1898. 405

** Riccardia multispica (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944. Bas.: Aneura multispica Steph., Sp. Hepat. (Stephani) 6: 34, 1917.

** Riccardia nadeaudii (Steph.) Hürl., Bauhinia 5 (4): 201, 1976. Bas.: Aneura nadeaudii Steph., Bull. Herb. Boissier 7 (10): 750 (260), 1899.

** Riccardia newellana (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963. Bas.: Aneura newellana Steph., Sp. Hepat. (Stephani) 6: 35, 1917.

** Riccardia nigra (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963. Bas.: Aneura nigra Steph., Sp. Hepat. (Stephani) 6: 35, 1917.

** Riccardia nobilis (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 56, 1898. Bas.: Aneura nobilis Steph., Hedwigia 32 (1): 24, 1893.

** Riccardia novo-amstelodamensis Schiffn., Deutsche Südpolar-Exped. 1901-1903, 8 (bot.) 1: 65, 1906.

** Riccardia nudiflora (Steph.) Grolle, Bryophyt. Biblioth. 48: 130, 1995. Bas.: Aneura nudiflora Steph., Bot. Gaz. 15 (11): 282, 1890.

** Riccardia obtusa S.W.Arnell, Bot. Not. 105: 142, 1952.

** Riccardia obtusifrons (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963. Bas.: Aneura obtusifrons Steph., Sp. Hepat. (Stephani) 6: 36, 1917.

** Riccardia omkaliensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 97, 1970.

*** Riccardia pallida (Spruce) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 15, 1985. Bas.: Aneura pallida Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 547, 1885.

** Riccardia palmatifida (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981. Bas.: Aneura palmatifida Steph., Sp. Hepat. (Stephani) 6: 36, 1917.

* Riccardia palmatiformis Schiffn., J. Indian Bot. Soc. 38 (4): 538, 1959 [1960]. 406

** Riccardia papillata (Gottsche) Hässel ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 130, 1979. Bas.: Pseudoneura papillata Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 185, 1864.

** Riccardia paramorum Meenks, J. Hattori Bot. Lab. 62: 176, 1987.

*** Riccardia parasitans (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 17, 1985. Bas.: Aneura parasitans Steph., Biblioth. Bot. 87 (2): 175, 1916.

** Riccardia parvula Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898.

** Riccardia pauciramea (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963. Bas.: Aneura pauciramea Steph., Bull. Herb. Boissier 5 (10): 845, 1897.

** Riccardia paulensis Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 16, 1964.

** Riccardia pectinata (Austin) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 526, 1963. Bas.: Aneura pectinata Austin, Bull. Torrey Bot. Club 5 (3): 15, 1874.

** Riccardia pellucida Piippo, J. Hattori Bot. Lab. 68: 134, 1990. Nom. nov. pro Aneura pellucida Steph., Sp. Hepat. (Stephani) 6: 37, 1917, nom. illeg.

** Riccardia pembaiensis (Steph.) Hürl., Bauhinia 5 (4): 205, 1976. Bas.: Aneura pembaiensis Steph., Sp. Hepat. (Stephani) 6: 37, 1917.

** Riccardia pengagensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 108, 1970.

** Riccardia perssonii S.C.Srivast. et Udar, Lindbergia 4 (1/2): 127, 1977.

*** Riccardia philippinensis Furuki, J. Hattori Bot. Lab. 100: 94, 2006.

** Riccardia phleganiana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 80, 1970.

** Riccardia plana (Steph.) Hürl., Bauhinia 5 (4): 205, 1976. Bas.: Aneura plana Steph., Sp. Hepat. (Stephani) 6: 38, 1917.

* Riccardia plana var. minor (Pearson) Hürl. ex H.A.Mill., H.Whittier et B.Whittier, Bryophyt. Biblioth. 25: 303, 1983. Bas.: Aneura plana var. minor Pearson, J. Linn. Soc., Bot. 46 (305): 16, 1922.

** Riccardia planifrons (Spruce) Pagán, Bryologist 45 (4): 80, 1942. Bas.: Aneura planifrons Spruce, J. Linn. Soc., Bot. 30 (210): 368, 1895.

*** Riccardia plumiformis (Spruce) Hässel ex Meenks, Beih. Nova Hedwigia 88: 101, 1987. Bas.: Aneura plumiformis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 548, 1885.

** Riccardia plumosa (Mitt.) E.O.Campb., J. Roy. Soc. New Zealand 1 (1): 24, 1971. Bas.: Sarcomitrium plumosum Mitt., Bonplandia 10 (2): 19, 1862.

*** Riccardia poeppigiana (Lehm. et Lindenb.) Hässel ex Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 17, 1985. Bas.: Jungermannia poeppigiana Lehm. et Lindenb., Nov. Stirp. Pug. 6: 23, 1834.

** Riccardia porcina (Hewson) L.Söderstr., Phytotaxa 202 (1): 70, 2015. Bas.: Riccardia bliklika var. porcina Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 84, 1970.

** Riccardia portoricensis (Steph.) Pagán, Bryologist 42 (1): 7, 1939. Bas.: Aneura portoricensis Steph., Bull. Herb. Boissier 7 (10): 739 (249), 1899.

** Riccardia punahuina (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 526, 1963. Bas.: Aneura punahuina Steph., Sp. Hepat. (Stephani) 6: 39, 1917.

** Riccardia ramosissima (Steph.) Grolle, Bryophyt. Biblioth. 48: 130, 1995. Bas.: Aneura ramosissima Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 196, 1891 [1892].

** Riccardia regina Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 17, 1985.

*** Riccardia regnellii (Ångstr.) K.G.Hell, Bol. Fac. Filos. Univ. São Paulo, Bot. 25: 110, 1969. Bas.: Pseudoneura regnellii Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 90, 1876 [1877].

** Riccardia reyesiana Meenks, Acta Bot. Hung. 32 (1/4): 207, 1986.

** Riccardia riccioides Pearson, Univ. Calif. Publ. Bot. 10 (4): 310, 1923.

* Riccardia rigida Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898.

** Riccardia robbinsii Hewson et Grolle, J. Hattori Bot. Lab. 29: 72, 1966.

** Riccardia robusta (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981. Bas.: Aneura robusta Steph., Sp. Hepat. (Stephani) 6: 40, 1917.

** Riccardia rockii (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963. Bas.: Aneura rockii Steph., Sp. Hepat. (Stephani) 6: 41, 1917.

** Riccardia rupicola (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 99, 1970. Bas.: Aneura rupicola Steph., Sp. Hepat. (Stephani) 6: 41, 1917.

** Riccardia russellii R.M.Schust., J. Hattori Bot. Lab. 67: 93, 1989.

** Riccardia saccatiflora (Steph.) S.W.Arnell, Bot. Not. 105: 144, 1952. Bas.: Aneura saccatiflora Steph., Bot. Gaz. 15 (11): 282, 1890.

** Riccardia santapaui Udar et S.C.Srivast., Rev. Bryol. Lichénol. 39 (1): 155, 1973.

*** Riccardia schwaneckei (Steph.) Pagán, Bryologist 42 (1): 7, 1939. Bas.: Aneura schwaneckei Steph., Hedwigia 27 (11/12): 278, 1888.

** Riccardia singapurensis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898.

** Riccardia smaragdina Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 20, 1985.

*** Riccardia sprucei (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 22, 1985. Bas.: Aneura sprucei Steph., Bull. Herb. Boissier 5 (10): 844, 1897.

** Riccardia squamifera Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 19, 1964.

* Riccardia stipatiflora (Steph.) Pagán, Bryologist 45 (4): 81, 1942. Bas.: Aneura stipatiflora Steph., Hedwigia 32 (1): 27, 1893.

** Riccardia stricta R.M.Schust., J. Hattori Bot. Lab. 62: 326, 1987.

** Riccardia subantarctica Grolle et L.Söderstr., Phytotaxa 202 (1): 70, 2015. Nom. nov. pro Riccardia pauciramea R.M.Schust., J. Hattori Bot. Lab. 67: 102, 1989, nom. illeg.

** Riccardia subexalata Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 163, 1898.

* Riccardia subexalata var. procera Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 164, 1898.

* Riccardia submersa (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877. Bas.: Jungermannia multifida var. submersa Hook.f. et Taylor, London J. Bot. 4: 94, 1845.

** Riccardia submultifida Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 128, 1934.

** Riccardia subpalmata (Steph.) Hürl., Bauhinia 5 (4): 194, 1976. Bas.: Aneura subpalmata Steph., Sp. Hepat. (Stephani) 6: 43, 1917.

** Riccardia subsimplex (Steph.) Pagán, Bryologist 45 (4): 81, 1942. Bas.: Aneura subsimplex Steph., Hedwigia 32 (1): 27, 1893.

** Riccardia sumatrana Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 173, 1898.

** Riccardia tahitensis (Steph.) Hürl., Bauhinia 5 (4): 201, 1976. Bas.: Aneura tahitensis Steph., Bull. Herb. Boissier 7 (10): 728 (238), 1899.

* Riccardia tenella Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 95, 1970.

*** Riccardia tenuicula (Spruce) Meenks, Beih. Nova Hedwigia 88: 101, 1987. Bas.: Aneura tenuicula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 545, 1885.

** Riccardia tenuis (Steph.) Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898. Bas.: Aneura tenuis Steph., Hedwigia 32 (1): 28, 1893.

* Riccardia tjibodensis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898.

*** Riccardia trichomanoides (Spruce) Hässel ex Meenks, Beih. Nova Hedwigia 88: 101, 1987. Bas.: Aneura trichomanoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 547, 1885.

** Riccardia trukensis H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 70, 1963.

** Riccardia tumbareriensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 87, 1970.

** Riccardia upoluna (Steph.) Grolle, J. Hattori Bot. Lab. 36: 77, 1972 [1973]. Bas.: Aneura upoluna Steph., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 91: 165, 1915.

** Riccardia valida (Steph.) J.J.Engel, Bryologist 78 (3): 362, 1975. Bas.: Aneura valida Steph., Sp. Hepat. (Stephani) 6: 44, 1917.

** Riccardia venosa (Steph.) Hürl., Bauhinia 5 (4): 208, 1976. Bas.: Aneura venosa Steph., Sp. Hepat. (Stephani) 6: 45, 1917.

* Riccardia villosa (Steph.) Pandé et S.C.Srivast., Biol. Mem. 1 (1/2): 129, 1976. Bas.: Aneura villosa Steph., Sp. Hepat. (Stephani) 6: 45, 1917.

** Riccardia virens (Steph.) Hürl., Bauhinia 5 (4): 209, 1976. Bas.: Aneura virens Steph., Sp. Hepat. (Stephani) 6: 45, 1917.

** Riccardia virgata (Gottsche) Pagán, Bryologist 42 (1): 7, 1939. Bas.: Aneura virgata Gottsche, Hedwigia 27 (11/12): 277, 1888.

** Riccardia wallisii (Steph.) Gradst., J. Hattori Bot. Lab. 45: 130, 1979. Bas.: Aneura wallisii Steph., Hedwigia 32 (1): 28, 1893.

** Riccardia wettsteinii Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 162, 1898.

* Riccardia wettsteinii var. angustilimbia Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 163, 1898.

* Riccardia wettsteinii var. crassa Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 163, 1898.

* Riccardia wettsteinii var. procera Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 162, 1898.

* Riccardia wettsteinii var. tenuiretis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 162, 1898.

** Riccardia womersleyana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 98, 1970.

** Verdoornia R.M.Schust., J. Hattori Bot. Lab. 26: 291, 1963.

*** Verdoornia succulenta R.M.Schust., J. Hattori Bot. Lab. 26: 291, 1963.

*** Metzgeriaceae H.Klinggr.

by D. P. Costa

The treatment of Metzgeriaceae follows what was outlined in,,, and.

*** Metzgeria Raddi, Jungermanniogr. Etrusca: 34, 1818. 407

*** Metzgeria acuminata Steph., Bull. Herb. Boissier 7 (12): 934 (282), 1899.

*** Metzgeria adscendens Steph. ex K.I.Goebel, Flora 77: 427, 1893.

*** Metzgeria agnewiae Kuwah., Bryologist 76 (4): 569, 1973. 408

*** Metzgeria albinea Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cci, 1889 [1890].

*** Metzgeria albinea var. aberrans Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 27, 1964.

*** Metzgeria albinea var. angusta (Steph.) D.P.Costa et Gradst., Bryologist 103 (4): 757, 2000 [2001]. Bas.: Metzgeria angusta Steph., Bull. Herb. Boissier 7 (12): 944 (292), 1899.

*** Metzgeria allionii Steph., Sp. Hepat. (Stephani) 6: 47, 1917.

*** Metzgeria alpina R.M.Schust. et J.J.Engel, Brittonia 40 (2): 203, 1988.

*** Metzgeria americana Masuzaki, Hikobia 15 (4): 441, 2010.

*** Metzgeria attenuata Steph., Biblioth. Bot. 87 (2): 177, 1916.

*** Metzgeria aurantiaca Steph., Bull. Herb. Boissier 7 (12): 938 (286), 1899.

*** Metzgeria auriculata Grolle et Kuwah., Bryophyt. Biblioth. 28: 194, 1986.

*** Metzgeria bahiensis Schiffn., Österr. Bot. Z. 61 (7/8): 262, 1911.

*** Metzgeria bartlettii Kuwah., Mem. New York Bot. Gard. 45: 561, 1987.

*** Metzgeria bischlerae Kuwah., J. Hattori Bot. Lab. 40: 264, 1976.

*** Metzgeria bracteata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 553, 1885.

*** Metzgeria brasiliensis Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 22, 1964.

*** Metzgeria chilensis Steph., Bull. Herb. Boissier 7 (12): 937 (285), 1899.

*** Metzgeria ciliata Raddi, Critt. Brasil.: 17, 1822. 409

*** Metzgeria claviflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 556, 1885.

*** Metzgeria cleefii Kuwah., Proc. Kon. Ned. Akad. Wetensch. C 85 (3): 360, 1982.

** Metzgeria comata Steph., Bull. Herb. Boissier 7 (12): 939 (287), 1899.

*** Metzgeria conjugata Lindb., Acta Soc. Sci. Fenn. 10: 495, 1875. 410

*** Metzgeria consanguinea Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 271, 1893.

*** Metzgeria convoluta Steph., Bull. Herb. Boissier 7 (12): 940 (288), 1899.

** Metzgeria coorgensis S.C.Srivast. et S.Srivast., Phytotaxonomy 4: 81, 2004.

** Metzgeria corralensis Steph., Bull. Herb. Boissier 7 (12): 933 (281), 1899.

*** Metzgeria crassipilis (Lindb.) A.Evans, Rhodora 11 (130): 188, 1909. Bas.: Metzgeria furcata subsp. crassipilis Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 42, 1877.

*** Metzgeria cratoneura Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 24, 1964.

*** Metzgeria cylindra Kuwah., Bryologist 81 (3): 405, 1978.

** Metzgeria decrescens Steph., Bull. Herb. Boissier 7 (12): 932 (280), 1899.

*** Metzgeria dichotoma (Sw.) Nees, Syn. Hepat. 4: 504, 1846. Bas.: Jungermannia dichotoma Sw., Prodr. (Swartz): 145, 1788.

** Metzgeria divaricata A.Evans, Proc. Amer. Acad. Arts 58 (7): 288, 1923.

*** Metzgeria dorsipara (Herzog) Kuwah., J. Hattori Bot. Lab. 40: 269, 1976. Bas.: Metzgeria violacea var. dorsipara Herzog, Svensk Bot. Tidskr. 51 (1): 187, 1957.

*** Metzgeria duricosta Steph., Sp. Hepat. (Stephani) 6: 50, 1917.

** Metzgeria engelii Kuwah., Hikobia 8 (3/4): 275, 1980.

** Metzgeria epiphylla A.Evans, Proc. Amer. Acad. Arts 58 (7): 303, 1923.

*** Metzgeria filicina Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 361, 1851.

** Metzgeria flavovirens Colenso, Trans. & Proc. New Zealand Inst. 21: 79, 1889.

* Metzgeria foliicola Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 181, 1898. 411

*** Metzgeria francana Steph., Sp. Hepat. (Stephani) 6: 51, 1917.

** Metzgeria frontipilis Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 14, 1877.

*** Metzgeria fruticola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 554, 1885.

*** Metzgeria furcata (L.) Corda, Gen. hepat.: 654, 1829. Bas.: Jungermannia furcata L., Sp. Pl. 1: 1136, 1753. 412

* Metzgeria furcata var. expansa Douin, Rev. Bryol. 30 (3): 47, 1903.

* Metzgeria furcata var. pacifica Brinkm., Bryologist 34 (2): 15, 1931.

*** Metzgeria grandiflora A.Evans, Torreya 16 (3): 68, 1916.

** Metzgeria hasselii Kuwah., J. Hattori Bot. Lab. 40: 509, 1976.

** Metzgeria hebridensis Steph., Sp. Hepat. (Stephani) 6: 51, 1917.

*** Metzgeria hegewaldii Kuwah., Nova Hedwigia 34: 784, 1981.

*** Metzgeria herminieri Schiffn., Österr. Bot. Z. 61 (7/8): 261, 1911.

* Metzgeria heteroramea Steph., Biblioth. Bot. 87 (2): 178, 1916.

** Metzgeria imberbis J.B.Jack et Steph., Hedwigia 34 (6): 316, 1895.

*** Metzgeria inflata Steph., Bull. Herb. Boissier 7 (12): 936 (284), 1899.

*** Metzgeria jamesonii Kuwah., Bryophyt. Biblioth. 28: 157, 1986.

** Metzgeria kanaii Kuwah., Fl. E. Himalaya 2: 240, 1971.

*** Metzgeria kinabaluensis Masuzaki, Hikobia 16 (1): 59, 2011. Based on: Apometzgeria pubescens var. kinabaluensis Kuwah., J. Hattori Bot. Lab. 28: 166, 1965, nom. inval.

** Metzgeria kuwaharae Piippo, Acta Bot. Fenn. 143: 8, 1991.

** Metzgeria laciniata Kuwah., Bryologist 81 (3): 406, 1978.

*** Metzgeria lechleri Steph., Bull. Herb. Boissier 7 (12): 942 (290), 1899.

*** Metzgeria leptoneura Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 555, 1885.

** Metzgeria leptoneura var. breviseta (Schiffn.) O.Yano, J. Hattori Bot. Lab. 56: 532, 1984. Bas.: Metzgeria hamata var. breviseta Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 27, 1964.

** Metzgeria leptoneura var. polychaeta R.M.Schust., Phytotaxa 202 (1): 70, 2015. Based on: Metzgeria leptoneura var. polychaeta R.M.Schust., J. Hattori Bot. Lab. 70: 150, 1991, nom. inval.

*** Metzgeria liebmaniana Lindenb. et Gottsche, Syn. Hepat. 4: 505, 1846.

*** Metzgeria lindbergii Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 182, 1898.

** Metzgeria litoralis J.J.Engel et Kuwah., Bryologist 76 (2): 293, 1973.

*** Metzgeria longitexta Steph., Bull. Herb. Boissier 7 (12): 940 (288), 1899.

** Metzgeria macrospora Kuwah., J. Hattori Bot. Lab. 32: 17, 1969.

** Metzgeria macveanii Kuwah., Rev. Bryol. Lichénol. 36 (3/4): 539, 1969 [1970].

*** Metzgeria maegdefraui Kuwah., Hikobia 8 (3/4): 269, 1980.

** Metzgeria magellanica Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 43, 1890.

*** Metzgeria metaensis Kuwah., Proc. Kon. Ned. Akad. Wetensch. C 85 (3): 375, 1982.

*** Metzgeria mexicana Steph., Sp. Hepat. (Stephani) 6: 55, 1917. 413

** Metzgeria monoica Kuwah. et J.J.Engel, Hikobia 8 (3/4): 284, 1980.

*** Metzgeria myriopoda Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 22, 1877.

*** Metzgeria neotropica Kuwah., Nova Hedwigia 34: 779, 1981.

*** Metzgeria nudifrons Steph., Hedwigia 31 (3): 126, 1892.

*** Metzgeria parviinvolucrata Kuwah., Nova Hedwigia 34: 774, 1981.

** Metzgeria patagonica Steph., Bull. Herb. Boissier 7 (12): 940 (288), 1899.

*** Metzgeria polytricha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 553, 1885.

*** Metzgeria procera Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 166, 1855.

*** Metzgeria psilocraspeda Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 25, 1964.

*** Metzgeria pubescens (Schrank) Raddi, Jungermanniogr. Etrusca: 35, 1818. Bas.: Jungermannia pubescens Schrank, Prim. Fl. Salisb.: 231, 1792.

*** Metzgeria pulvinata Steph., Biblioth. Bot. 87 (2): 179, 1916.

** Metzgeria quadrifaria Steph., Bull. Herb. Boissier 7 (12): 953 (301), 1899.

** Metzgeria raoi S.C.Srivast. et S.Srivast., Phytotaxonomy 4: 83, 2004.

** Metzgeria rigida Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 43, 1877.

** Metzgeria robinsonii Steph., Sp. Hepat. (Stephani) 6: 60, 1917.

** Metzgeria roivainenii Kuwah., J. Hattori Bot. Lab. 40: 516, 1976.

*** Metzgeria rufula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 555, 1885.

** Metzgeria saccata Mitt., J. Linn. Soc., Bot. 22 (145): 241, 1886.

** Metzgeria saxbyi Pearson, Ann. Cryptog. Exot. 4 (2): 70, 1931.

** Metzgeria scobina Mitt., J. Linn. Soc., Bot. 22 (145): 243, 1886.

*** Metzgeria scyphigera A.Evans, Trans. Connecticut Acad. Arts 18 (5): 299, 1914.

*** Metzgeria senjoana Masuzaki, Hikobia 15 (4): 445, 2010.

** Metzgeria setigera R.M.Schust. ex Crand.-Stotl. et L.Söderstr., Phytotaxa 202 (1): 69, 2015. Based on: Metzgeria furcata var. setigera R.M.Schust., J. Hattori Bot. Lab. 70: 149, 1991, nom. inval.

** Metzgeria sikkimensis S.C.Srivast. et K.K.Rawat, Geophytology 31 (1/2): 71, 2001 [2003].

* Metzgeria simplex Lorb. ex Müll.Frib., Hedwigia 80 (1/2): 115, 1941.

*** Metzgeria sinuata Loitl., Diagn. pl. nov.: 25, 1894.

** Metzgeria sparrei Kuwah., Hikobia 8 (3/4): 278, 1980.

*** Metzgeria spindleri Steph., Biblioth. Bot. 87 (2): 179, 1916.

*** Metzgeria subaneura Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 22, 1964.

** Metzgeria submarginata M.L.So, New Zealand J. Bot. 40 (2): 201, 2002.

** Metzgeria subundulata (Lindb.) Kuwah., Bryologist 86 (3): 276, 1983 [1984]. Bas.: Metzgeria furcata subsp. subundulata Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 42, 1877.

** Metzgeria temperata Kuwah., J. Hattori Bot. Lab. 40: 219, 1976.

*** Metzgeria uncigera A.Evans, Ann. Bot. (Oxford) 24 (2): 276, 1910.

*** Metzgeria undulata Kuwah., Nova Hedwigia 34: 792, 1981.

*** Metzgeria violacea (Ach.) Dumort., Recueil Observ. Jungerm.: 26, 1835. Bas.: Jungermannia violacea Ach., Beitr. Naturk. (Weber & Mohr) 1: 77, 1805.

** Steereella Kuwah., Amer. J. Bot. 60 (6): 602, 1973.

*** Steereella lilliana (Steph.) Kuwah., Bryophyt. Biblioth. 28: 179, 1986. Bas.: Metzgeria lilliana Steph., Sp. Hepat. (Stephani) 6: 53, 1917.

*** Steereella linearis (Sw.) Kuwah., Amer. J. Bot. 60 (6): 604, 1973. Bas.: Jungermannia linearis Sw., Prodr. (Swartz): 145, 1788.

** Vandiemenia Hewson, J. Hattori Bot. Lab. 52: 163, 1982.

** Vandiemenia ratkowskiana Hewson, J. Hattori Bot. Lab. 52: 163, 1982.

*** Pleuroziaceae Müll.Frib.

by B. Thiers

*** Pleurozia Dumort., Recueil Observ. Jungerm.: 15, 1835.

** subg. Constantifolia B.M.Thiers, Bryologist 96 (4): 526, 1993.

*** Pleurozia conchifolia (Hook. et Arn.) Austin, Bull. Torrey Bot. Club 5 (3): 16, 1874. Bas.: Jungermannia conchifolia Hook. et Arn., Bot. Beechey Voy. 3: 110, 1832.

** Pleurozia conchifolia var. papillosa B.M.Thiers, Bryologist 96 (4): 528, 1993.

*** Pleurozia purpurea Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 27, 1877. Based on: Jungermannia purpurea Lightf., Fl. Scot. 2: 778, 1777, nom. illeg.

** subg. Diversifolia B.M.Thiers, Bryologist 96 (4): 531, 1993.

*** Pleurozia acinosa (Mitt.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 412, 1877. Bas.: Physiotium acinosum Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 102, 1860 [1861].

*** Pleurozia articulata (Lindb.) Lindb. et Lackström, Hepat. Scand. Exsicc.: 5 (adnot.), 1874. Bas.: Physiotium articulatum Lindb., Öfvers. Förh. Finska Vetensk.-Soc. 12 (2): 78, 1870.

*** Pleurozia caledonica (Gottsche) Steph., Rev. Bryol. 33 (2): 29. Bas.: Physiotium caledonicum Gottsche, Hedwigia 25 (2/3): 81, 1886.

*** Pleurozia curiosa B.M.Thiers, Bryologist 96 (4): 537, 1993.

*** Pleurozia heterophylla Steph. ex Fulford, Mem. New York Bot. Gard. 23: 842, 1972.

*** Pleurozia johannis-winkleri Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 195, 1931.

*** Pleurozia paradoxa (J.B.Jack) Schiffn., Hepat. (Engl.-Prantl): 115, 1893. Bas.: Physiotium paradoxum J.B.Jack, Hedwigia 25 (2/3): 85, 1886.

*** Pleurozia subinflata (Austin) Austin, Bull. Torrey Bot. Club 5 (3): 16, 1874. Bas.: Physiotium subinflatum Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 224, 1869.

** subg. Pleurozia

*** Pleurozia gigantea (F.Weber) Lindb., Hepat. Scand. Exsicc.: no. 5, 1874. Bas.: Jungermannia gigantea F.Weber, Hist. Musc. Hepat. Prodr.: 57, 1815.

Incertae sedis

** Pleurozia pocsii Frank Müll., Polish Bot. J. 58 (1): 50, 2013.

*** Calyculariaceae He-Nygrén, Juslén, Ahonen, Glenny et Piippo

*** Calycularia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 122, 1860 [1861].

*** Calycularia crispula Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 122, 1860 [1861].

*** Calycularia laxa Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 66, 1889.

** Allisoniaceae Schljakov

*** Allisonia Herzog, Hedwigia 80 (1/2): 77, 1941.

*** Allisonia cockaynei (K.I.Goebel) R.M.Schust., J. Hattori Bot. Lab. 26: 294, 1963. Bas.: Moerckia cockaynei K.I.Goebel, Flora 96: 190, 1906.

*** Fossombroniaceae Hazsl. nom. conserv.

by R. Stotler, B. J. Crandall-Stotler and D. C. Cargill

*** Fossombronia Raddi, Jungermanniogr. Etrusca: 29, 1818.

*** Fossombronia alaskana Steere et Inoue, Bryologist 77 (1): 66, 1974.

*** Fossombronia alata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 340, 1984.

*** Fossombronia altilamellosa G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003. Based on: Fossombronia altilamellosa G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 367, 1987, nom. inval.

*** Fossombronia angulifolia Perold, Bothalia 28 (2): 159, 1998.

*** Fossombronia angulosa (Dicks.) Raddi, Jungermanniogr. Etrusca: 29, 1818. Bas.: Jungermannia angulosa Dicks., Fasc. Pl. Crypt. Brit. 1: 7, 1785.

** Fossombronia areolata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 368, 1987.

** Fossombronia auricolor G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003. Based on: Fossombronia auricolor G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 370, 1987, nom. inval.

*** Fossombronia australis Mitt., J. Linn. Soc., Bot. 15 (82): 73, 1876.

*** Fossombronia caespitiformis (Raddi) De Not. ex Rabenh., Hepat. Eur., Leberm. 13-14: no. 123, 1860 [1861]. Bas.: Fossombronia angulosa var. caespitiformis Raddi, Jungermanniogr. Etrusca: 30, 1818.

*** Fossombronia caespitiformis subsp. multispira (Schiffn.) J.R.Bray et Cargill, Bryologist 106 (1): 131, 2003. Bas.: Fossombronia caespitiformis var. multispira Schiffn., Österr. Bot. Z. 67 (4/5): 152, 1918.

*** Fossombronia caledonica Steph., Sp. Hepat. (Stephani) 6: 71, 1917.

*** Fossombronia cederbergensis Perold, Bothalia 28 (1): 1, 1998.

*** Fossombronia cerebriformis G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 343, 1984.

*** Fossombronia crassifolia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 527, 1885.

*** Fossombronia crispa Nees, Syn. Hepat. 4: 469, 1846.

* Fossombronia crispula (Brot.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 383, 1992. Bas.: Jungermannia crispula Brot., Fl. lusit. 2: 422, 1804 [1805]. 414

*** Fossombronia cristula Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 228, 1869.

*** Fossombronia cultriformis G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 371, 1987.

*** Fossombronia densa G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 372, 1987.

*** Fossombronia densilamellata S.W.Arnell, Bot. Not. 105: 317, 1952.

*** Fossombronia echinata Macvicar, Rev. Bryol. 38 (4): 73, 1911.

*** Fossombronia elsieae Perold, Bothalia 29 (1): 25, 1999.

*** Fossombronia fernandeziensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 15, 1911.

*** Fossombronia fimbriata Paton, J. Bryol. 8 (1): 1, 1974.

*** Fossombronia fleischeri Osterwald ex Loeske, Verh. Bot. Vereins Prov. Brandenburg 70: 125, 1928.

*** Fossombronia foveolata Lindb., Helsingf. Dagbl. 1873 (353, 28 Dec): 2, 1873.

** Fossombronia fuhreri G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 374, 1987.

*** Fossombronia gemmifera Perold, Bothalia 27 (1): 19, 1997.

*** Fossombronia glenii Perold, Bothalia 27 (1): 20, 1997.

** Fossombronia grandis Steph., Mém. Herb. Boissier 16: 29 (383), 1900.

* Fossombronia gregaria Colenso, Trans. & Proc. New Zealand Inst. 20: 252, 1888. 415

* Fossombronia grossepapillata Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 105, 1914. 416

*** Fossombronia hamatohirta Steph., Hedwigia 33 (1): 8, 1894.

** Fossombronia hewsoniae G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 375, 1987.

*** Fossombronia himalayensis Kashyap, New Phytol. 14 (1): 4, 1915.

*** Fossombronia hyalorhiza Perold, Bothalia 29 (1): 83, 1999.

*** Fossombronia incurva Lindb., Helsingf. Dagbl. 1873 (273, 7 Oct): 2, 1873.

*** Fossombronia indica Steph., Sp. Hepat. (Stephani) 6: 73, 1917.

* Fossombronia integerrima Steph., Mém. Herb. Boissier 16: 40 (394), 1900. 417

* Fossombronia integrifolia Steph., Sp. Hepat. (Stephani) 6: 73, 1917. 418

*** Fossombronia intestinalis Taylor, London J. Bot. 5: 408, 1846.

*** Fossombronia japonica Schiffn., Österr. Bot. Z. 49 (11): 389, 1899.

** Fossombronia laciniata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 375, 1987.

*** Fossombronia lamellata Steph., Hedwigia 33 (1): 9, 1894.

*** Fossombronia leucoxantha (Lehm.) Lehm. et Lindenb., Syn. Hepat. 4: 469, 1846. Bas.: Jungermannia leucoxantha Lehm., Linnaea 4: 368, 1829.

*** Fossombronia longiseta (Austin) Austin, Hepat. bor.-amer.: 30, 1873. Bas.: Androcryphia longiseta Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 228, 1869.

*** Fossombronia lophoclada Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 529, 1885.

** Fossombronia lophoscypha Hässel, Beih. Nova Hedwigia 131: 16, 2007.

*** Fossombronia luetzelburgiana K.I.Goebel, Flora 105: 55, 1912.

*** Fossombronia macrocalyx Steph., Sp. Hepat. (Stephani) 6: 74, 1917.

* Fossombronia macrophylla Colenso, Trans. & Proc. New Zealand Inst. 18: 285, 1886.

** Fossombronia magnaspora G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003. Based on: Fossombronia magnaspora G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 377, 1987, nom. inval.

*** Fossombronia marindae Perold, Bothalia 29 (1): 86, 1999.

** Fossombronia maritima (Paton) Paton, J. Bryol. 18 (2): 367, 1994. Bas.: Fossombronia pusilla var. maritima Paton, J. Bryol. 7 (3): 244, 1973.

*** Fossombronia marshii J.R.Bray et Stotler, Phytologia 92 (2): 230, 2010.

** Fossombronia microlamellata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 347, 1984.

*** Fossombronia mittenii Tind., J. Bot. 36: 44, 1898.

*** Fossombronia montaguensis S.W.Arnell, Bot. Not. 105: 316, 1952.

*** Fossombronia monticola Perold, Bothalia 29 (1): 87, 1999.

*** Fossombronia mylioides Inoue, J. Hattori Bot. Lab. 37: 296, 1973.

* Fossombronia nigricaulis Colenso, Trans. & Proc. New Zealand Inst. 18: 248, 1886. 419

** Fossombronia nyikaensis Perold, Bothalia 31 (1): 48, 2001.

*** Fossombronia papillata Steph., Hedwigia 28 (3): 157, 1889.

*** Fossombronia paranapanemae Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 34, 1964.

*** Fossombronia peruviana Gottsche et Hampe, Linnaea 27 (5): 555, 1854.

*** Fossombronia porphyrorhiza (Nees) Prosk., Bryologist 58 (3): 197, 1955. Bas.: Jungermannia porphyrorhiza Nees, Fl. Bras. (Martius) 1 (1): 343, 1833.

** Fossombronia pulvinata Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 113, 1911.

*** Fossombronia punctata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 341, 1984.

*** Fossombronia purpureospora G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003. Based on: Fossombronia purpureospora G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 379, 1987, nom. inval.

*** Fossombronia pusilla (L.) Nees, Naturgesch. Eur. Leberm. 3: 319, 1838. Bas.: Jungermannia pusilla L., Sp. Pl. 1: 1136, 1753.

*** Fossombronia renateae Perold, Bothalia 29 (1): 77, 1999.

*** Fossombronia reticulata Steph., Hedwigia 33 (1): 9, 1894.

* Fossombronia rosulata Colenso, Trans. & Proc. New Zealand Inst. 18: 248, 1886.

** Fossombronia rudis G.A.M.Scott et D.C.Pike, Beih. Nova Hedwigia 90: 110, 1988.

*** Fossombronia ruminata Cargill, Phytotaxa 65: 45, 2012. Nom. nov. pro Fossombronia maritima G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003, nom. illeg.

** Fossombronia rupestris G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003. Based on: Fossombronia rupestris G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 381, 1987, nom. inval.

*** Fossombronia rwandaensis Perold, Bothalia 28 (1): 45, 1998.

*** Fossombronia scrobiculata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 341, 1984.

*** Fossombronia spinifolia Steph., Mém. Herb. Boissier 16: 35 (389), 1900.

* Fossombronia spinosa Perold, Bothalia 29 (1): 29, 1999.

*** Fossombronia stephanii Schiffn. ex Steph., Mém. Herb. Boissier 16: 27 (381), 1900.

*** Fossombronia straussiana Perold, Bothalia 27 (1): 24, 1997.

* Fossombronia subsaccata Steph., Sp. Hepat. (Stephani) 6: 75, 1917. 420

*** Fossombronia swaziensis Perold, Bothalia 28 (2): 162, 1998.

** Fossombronia tesselata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 382, 1987.

*** Fossombronia texana Lindb., Acta Soc. Sci. Fenn. 10: 533, 1875.

*** Fossombronia truncata G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003. Based on: Fossombronia truncata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 383, 1987, nom. inval.

*** Fossombronia tumida Mitt., J. Linn. Soc., Bot. 16 (91): 193, 1877.

** Fossombronia valparaisiana Hässel, Beih. Nova Hedwigia 131: 14, 2007.

** Fossombronia vermiculata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 345, 1984.

*** Fossombronia wattsii Steph., Sp. Hepat. (Stephani) 6: 75, 1917.

*** Fossombronia wondraczekii (Corda) Dumort. ex Lindb., Helsingf. Dagbl. 1873 (273, 7 Oct): 2, 1873. Bas.: Jungermannia wondraczekii Corda, Deutschl. Fl. (Sturm), Abt. 2, Cryptog.: 30, 1830.

*** Fossombronia wrightii Austin, Bot. Bull. (Hanover) 1 (8): 36, 1876.

** Fossombronia zuurbergensis Perold, Bothalia 31 (1): 25, 2001.

*** Petalophyllaceae Stotler et Crand.-Stotl.

by R. Stotler and B.J. Crandall-Stotler

*** Petalophyllum Nees et Gottsche, Nov. Stirp. Pug. 8: 29, 1844.

*** Petalophyllum americanum C.H.Ford et Crand.-Stotl., Novon 12 (3): 335, 2002.

*** Petalophyllum hodgsoniae C.H.Ford et Crand.-Stotl., Novon 12 (3): 336, 2002.

*** Petalophyllum indicum Kashyap, J. Indian Bot. Soc. 7: 14, 1928.

*** Petalophyllum preissii Gottsche, Nov. Stirp. Pug. 8: 30, 1844.

*** Petalophyllum ralfsii (Wilson) Nees et Gottsche, Nov. Stirp. Pug. 8: 30, 1844. Bas.: Jungermannia ralfsii Wilson, Suppl. Engl. Bot. 4: tab. 2874, 1849.

*** Sewardiella Kashyap, New Phytol. 14 (1): 5, 1915.

*** Sewardiella tuberifera Kashyap, New Phytol. 14 (1): 5, 1915.

Makinoiineae He-Nygrén, Juslén, Ahonen, Glenny et Piippo

*** Makinoaceae Nakai

by R. Stotler and B.J. Crandall-Stotler

*** Makinoa Miyake, Bot. Mag. (Tokyo) 13 (144): 23, 1899.

*** Makinoa crispata (Steph.) Miyake, Bot. Mag. (Tokyo) 13 (144): 23, 1899. Bas.: Pellia crispata Steph., Bull. Herb. Boissier 5 (2): 103, 1897.

*** Hymenophytaceae R.M.Schust.

by R. Stotler and B.J. Crandall-Stotler

*** Hymenophyton Dumort., Recueil Observ. Jungerm.: 25, 1835.

*** Hymenophyton flabellatum (Labill.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 430, 1877. Bas.: Jungermannia flabellata Labill., Nov. Holl. Pl. 2: 109, 1806.

*** Hymenophyton leptopodum (Hook.f. et Taylor) A.Evans, Trans. Connecticut Acad. Arts 8 (16): 274, 1893. Bas.: Jungermannia leptopoda Hook.f. et Taylor, London J. Bot. 3: 571, 1844.

*** Hymenophyton pedicellatum Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 11, 1911.

*** Moerckiaceae K.I.Goebel ex Stotler et Crand.-Stotl.

by B.J. Crandall-Stotler and R. Stotler

showed that the family Moerckiaceae is heterogeneous and they moved Hattorianthus and Moerckia flotoviana to their new family Cordaeaceae. However, the type of Moerckiaceae (Moerckia hibernica) was not included in the study and only provisionally left in Moerckia. We prefer to follow Crandall-Stotler & Stotler (2007) until Moerckia hibernica is further studied.

*** Hattorianthus R.M.Schust. et Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 103, 1975.

*** Hattorianthus erimonus (Steph.) R.M.Schust. et Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 106, 1975. Bas.: Pallavicinia erimona Steph., Bull. Herb. Boissier 5 (2): 102, 1897.

*** Moerckia Gottsche, Hepat. Eur., Leberm. 13-14: no. 121, 1860 [1861].

*** Moerckia blyttii (Mørch) Brockm., Arch. Vereins Freunde Naturgesch. Mecklenburg 17: 190, 1863. Bas.: Jungermannia blyttii Mørch, Fl. Danica 12: 6, 1830.

*** Moerckia flotoviana (Nees) Schiffn., Österr. Bot. Z. 51 (2): 43, 1901. Bas.: Cordaea flotoviana Nees, Flora 16 (26): 405, 1833.

*** Moerckia hibernica (Hook.) Gottsche, Hepat. Eur., Leberm. 13-14: no. 121, 1860 [1861]. Bas.: Jungermannia hibernica Hook., Brit. Jungermann.: tab. 78, 1816.

*** Pallaviciniaceae Mig.

by R. Stotler and B.J. Crandall-Stotler

** Pallavicinioideae Mig. ex Grolle

*** Jensenia Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 13, 1868.

*** Jensenia connivens (Colenso) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965]. Bas.: Symphyogyna connivens Colenso, Trans. & Proc. New Zealand Inst. 20: 254, 1888.

* Jensenia crassifrons (Steph.) S.Schuette et Stotler, J. Hattori Bot. Lab. 97: 300, 2005. Bas.: Pallavicinia crassifrons Steph., Mém. Herb. Boissier 11: 21 (325), 1900. 421

*** Jensenia decipiens (Mitt.) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965]. Bas.: Steetzia decipiens Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 123, 1860 [1861].

*** Jensenia difformis (Nees) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965]. Bas.: Jungermannia difformis Nees, Fl. Bras. (Martius) 1 (1): 329, 1833.

*** Jensenia florschuetzii Gronde, Proc. Kon. Ned. Akad. Wetensch. C 83 (3): 273, 1980.

*** Jensenia spinosa (Lindenb. et Gottsche) Grolle, Acta Bot. Fenn. 133: 65, 1986. Bas.: Symphyogyna spinosa Lindenb. et Gottsche, Syn. Hepat. 5: 786, 1847.

*** Jensenia wallisii (J.B.Jack et Steph.) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965]. Bas.: Pallavicinia wallisii J.B.Jack et Steph., Hedwigia 31 (1): 23, 1892.

*** Pallavicinia Gray, Nat. Arr. Brit. Pl. 1: 775, 1821 nom. conserv.

*** Pallavicinia ambigua (Mitt.) Steph., Mém. Herb. Boissier 11: 8 (312), 1900. Bas.: Steetzia ambigua Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 123, 1860 [1861].

** Pallavicinia baldwinii (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891. Bas.: Steetzia baldwinii Austin, Bull. Torrey Bot. Club 6 (52): 303, 1879.

* Pallavicinia bipinnata Steph., Sp. Hepat. (Stephani) 6: 62, 1917.

** Pallavicinia camisassai Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 4, 1916.

** Pallavicinia cylindrica (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891. Bas.: Steetzia cylindrica Austin, Bull. Torrey Bot. Club 5 (3): 17, 1874.

** Pallavicinia himalayensis Schiffn., Mém. Herb. Boissier 11: 13 (317), 1900.

*** Pallavicinia indica Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 183, 1898.

*** Pallavicinia levieri Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 184, 1898.

*** Pallavicinia lyellii (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 775, 1821. Bas.: Jungermannia lyellii Hook., Brit. Jungermann.: tab. 77, 1816.

** Pallavicinia pilifera Steph., Hedwigia 30 (6): 271, 1891.

** Pallavicinia purpurea Steph., Sp. Hepat. (Stephani) 6: 64, 1917.

** Pallavicinia ridleyi Steph., Sp. Hepat. (Stephani) 6: 64, 1917.

*** Pallavicinia rubristipa Schiffn., Österr. Bot. Z. 56 (1): 24, 1906.

*** Pallavicinia subciliata (Austin) Steph., Mém. Herb. Boissier 11: 9 (313), 1900. Bas.: Steetzia subciliata Austin, Bull. Torrey Bot. Club 6 (52): 303, 1879.

*** Pallavicinia xiphoides (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 427, 1877. Bas.: Jungermannia xiphoides Hook.f. et Taylor, London J. Bot. 3: 569, 1844.

*** Podomitrium Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 164, 1855.

*** Podomitrium malaccense (Steph.) Campb., Amer. J. Bot. 2 (5): 199, 1915. Bas.: Hymenophyton malaccense Steph., Hedwigia 34 (2): 46, 1895.

*** Podomitrium marginatum (Steph.) Hürl., Bauhinia 4 (1): 78, 1968 [1969]. Bas.: Hymenophyton marginatum Steph., Sp. Hepat. (Stephani) 6: 61, 1917.

*** Podomitrium phyllanthus (Hook.) Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 164, 1855. Bas.: Jungermannia phyllanthus Hook., Musci Exot. 1: tab. 95, 1818.

** Symphyogynoideae R.M.Schust. ex Grolle

** Greeneothallus Hässel, J. Bryol. 11 (1): 115, 1980.

*** Greeneothallus gemmiparus Hässel, J. Bryol. 11 (1): 115, 1980.

*** Seppeltia Grolle, J. Hattori Bot. Lab. 60: 276, 1986.

*** Seppeltia succuba Grolle, J. Hattori Bot. Lab. 60: 276, 1986.

*** Symphyogyna Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 66, 1836.

*** Symphyogyna apiculispina Steph., Biblioth. Bot. 87 (2): 180, 1916.

*** Symphyogyna aspera Steph. ex F.A.McCormick, Bot. Gaz. 58 (5): 403, 1914.

** Symphyogyna atronervia Taylor, London J. Bot. 5: 409, 1846.

** Symphyogyna bogotensis Steph., Mém. Herb. Boissier 11: 42 (346), 1900. Based on: Symphyogyna hymenophyllum var. bogotensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 181, 1864, nom. inval.

* Symphyogyna boliviensis Steph., Biblioth. Bot. 87 (2): 180, 1916.

*** Symphyogyna brasiliensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 67, 1836. Nom. nov. pro Jungermannia brasiliensis Nees, Enum. Pl. Crypt. Javae: 11, 1830, nom. illeg.

** Symphyogyna brasiliensis var. angustior (Gottsche, Lindenb. et Nees) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 183, 1864. Bas.: Symphyogyna brasiliensis β angustior Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 484, 1846.

** Symphyogyna brasiliensis var. subsinuata Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 32, 1964.

*** Symphyogyna brongniartii Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 265, 1843.

*** Symphyogyna circinata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 69, 1836.

*** Symphyogyna digitisquama Steph., Mém. Herb. Boissier 11: 31 (335), 1900.

** Symphyogyna fuscovirens A.Evans, Trans. Connecticut Acad. Arts 28 (6): 320, 1927.

*** Symphyogyna hochstetteri Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 68, 1836.

*** Symphyogyna hymenophyllum (Hook.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 68, 1836. Bas.: Jungermannia hymenophyllum Hook., Musci Exot. 1: tab. 14, 1818.

** Symphyogyna hymenophyllum var. heterogenum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 536, 1885.

** Symphyogyna ignambiensis Hürl., Bauhinia 4 (1): 79, 1968 [1969].

** Symphyogyna interrupta Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1053, 1888.

** Symphyogyna irregularis Steph., Mém. Herb. Boissier 11: 29 (333), 1900.

** Symphyogyna lacerosquama Steph., Sp. Hepat. (Stephani) 6: 67, 1917.

*** Symphyogyna leptothelia Taylor, London J. Bot. 5: 408, 1846.

** Symphyogyna lindmanii A.Evans, Trans. Connecticut Acad. Arts 28 (6): 316, 1927.

*** Symphyogyna luetzelburgii Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 22, 1925.

*** Symphyogyna marginata Steph., Mém. Herb. Boissier 11: 30 (334), 1900.

*** Symphyogyna mexicana Steph., Rev. Bryol. 36 (6): 140, 1909.

** Symphyogyna multiflora Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 133, 1914.

* Symphyogyna paucidens Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 13, 1911. 422

*** Symphyogyna podophylla (Thunb.) Nees et Mont., Flora 29 (9): 135, 1846. Bas.: Jungermannia podophylla Thunb., Prodr. Pl. Cap. 2: 174, 1800.

* Symphyogyna purpureolimbata E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 95, 1965.

*** Symphyogyna rectidens Grolle, Acta Bot. Fenn. 133: 68, 1986.

** Symphyogyna rhodina (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 487, 1846. Bas.: Jungermannia rhodina Hook.f. et Taylor, London J. Bot. 4: 93, 1845.

*** Symphyogyna rubescens Steph., Mém. Herb. Boissier 11: 29 (333), 1900.

*** Symphyogyna rubritincta A.Evans, Trans. Connecticut Acad. Arts 27 (1): 38, 1925.

** Symphyogyna semi-involucrata Austin, Bull. Torrey Bot. Club 5 (3): 15, 1874.

** Symphyogyna similis Grolle, Acta Bot. Fenn. 133: 70, 1986.

*** Symphyogyna sinuata (Sw.) Nees et Mont., Voy. Amér. Mérid., Bot. 7 (1): 61, 1839. Bas.: Jungermannia sinuata Sw., Prodr. (Swartz): 145, 1788.

*** Symphyogyna subsimplex Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 166, 1855.

*** Symphyogyna tenuinervis (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 61: 253, 1986 [1987]. Bas.: Jungermannia tenuinervis Hook.f. et Taylor, London J. Bot. 3: 570, 1844.

*** Symphyogyna trivittata Spruce, J. Linn. Soc., Bot. 30 (210): 365, 1895.

* Symphyogyna ulvoides (Reinw., Blume et Nees) Nees, Syn. Hepat. 4: 487, 1846. Bas.: Jungermannia ulvoides Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 196, 1824 [1825]. 423

*** Symphyogyna undulata Colenso, Trans. & Proc. New Zealand Inst. 16: 356, 1884.

** Symphyogyna volkensii Steph., Mém. Herb. Boissier 11: 35 (339), 1900.

** Symphyogynopsis Grolle, Acta Bot. Fenn. 133: 72, 1986.

*** Symphyogynopsis gottscheana (Mont. et Nees) Grolle, J. Hattori Bot. Lab. 63: 441, 1987. Bas.: Symphyogyna gottscheana Mont. et Nees, Syn. Hepat. 4: 484, 1846.

** Xenothallus R.M.Schust., J. Hattori Bot. Lab. 26: 293, 1963.

*** Xenothallus vulcanicola R.M.Schust., J. Hattori Bot. Lab. 26: 293, 1963.

** Sandeothallaceae R.M.Schust.

** Sandeothallus R.M.Schust., Nova Hedwigia 36: 10, 1982.

** Sandeothallus japonicus (Inoue) Crand.-Stotl. et Stotler, Beih. Nova Hedwigia 131: 58, 2007. Bas.: Moerckia japonica Inoue, Bull. Natl. Sci. Mus. Tokyo, B 11 (1): 8, 1985.

** Sandeothallus radiculosus (Schiffn.) R.M.Schust., Nova Hedwigia 36: 11, 1982. Bas.: Moerckia radiculosa Schiffn., Österr. Bot. Z. 51 (2): 48, 1901.

*** Phyllothalliaceae E.A.Hodgs. ex T.Katag.

*** Phyllothallia E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (19): 247, 1964.

*** Phyllothallia fuegiana R.M.Schust., Trans. Brit. Bryol. Soc. 5 (2): 284, 1967.

*** Phyllothallia nivicola E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (19): 247, 1964.

*** Noterocladaceae W.Frey et M.Stech

proposed Noterocladaceae as a monogeneric family based on molecular and morphological evidence, which was later corroborated by.

*** Noteroclada Taylor ex Hook.f. et Wilson, London J. Bot. 3: 166, 1844.

*** Noteroclada confluens Taylor, London J. Bot. 3: 166, 1844.

Excluded from the genus

* Noteroclada longiuscula Colenso, Trans. & Proc. New Zealand Inst. 19: 299, 1887. 424

*** Pelliaceae H.Klinggr.

recognized Pelliaceae as a monogeneric family, which was later supported by.

** Pellia Raddi, Jungermanniogr. Etrusca: 38, 1818 nom. conserv.

*** subg. Apopellia Grolle, J. Bryol. 12 (3): 427, 1983.

** Pellia alpicola R.M.Schust. ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 76 (3): 39, 2013. Based on: Pellia endiviifolia subsp. alpicola R.M.Schust., J. Hattori Bot. Lab. 70: 145, 1991, nom. inval.

*** Pellia endiviifolia (Dicks.) Dumort., Recueil Observ. Jungerm.: 27, 1835. Bas.: Jungermannia endiviifolia Dicks., Fasc. Pl. Crypt. Brit. 4: 19, 1801.

** Pellia megaspora R.M.Schust., J. Bryol. 11 (3): 419, 1981.

*** subg. Pellia

** Pellia appalachiana R.M.Schust., Phytotaxa 76 (3): 39, 2013. Based on: Pellia appalachiana R.M.Schust., J. Hattori Bot. Lab. 70: 145, 1991, nom. inval.

*** Pellia epiphylla (L.) Corda, Gen. hepat.: 654, 1829. Bas.: Jungermannia epiphylla L., Sp. Pl. 1: 1135, 1753.

** Pellia epiphylla subsp. borealis (Lorb.) Messe, Bull. Soc. Roy. Bot. Belgique 114 (1): 13, 1981. Bas.: Pellia borealis Lorb., Jahrb. Wiss. Bot. 80: 697, 1934.

*** Pellia neesiana (Gottsche) Limpr., Hedwigia 15 (2): 18, 1876. Bas.: Pellia epiphylla f. neesiana Gottsche, Hedwigia 6 (5): 69, 1867.

Incertae sedis

* Pellia cordaeana Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 433, 1877.

* Pellia crispa P.Kumm., Führer Leberm.: 60, 1875.

* Pellia gottscheana Kreh, Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 90 (4): 237, 1909.

* Pellia longifolia P.Kumm., Führer Leberm.: 60, 1875.

* Pellia undulata P.Kumm., Führer Leberm.: 60, 1875.

*** Blasiaceae H.Klinggr.

by L. Söderström

*** Blasia L., Sp. Pl. 1: 1138, 1753.

*** Blasia pusilla L., Sp. Pl. 1: 1138, 1753.

*** Cavicularia Steph., Bull. Herb. Boissier 5 (2): 87, 1897.

*** Cavicularia densa Steph., Bull. Herb. Boissier 5 (2): 87, 1897.

*** Lunulariaceae H.Klinggr.

by D.G. Long

*** Lunularia Adans., Fam. Pl. (Adanson) 2: 15, 1763.

*** Lunularia cruciata (L.) Dumort. ex Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 298, 1868. Bas.: Marchantia cruciata L., Sp. Pl. 1: 1137, 1753.

* Lunularia cruciata subsp. thaxteri (A.Evans et Herzog) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 91, 1992. Bas.: Lunularia thaxteri A.Evans et Herzog, Arch. Esc. Fárm. Fac. Ci. Méd. Córdoba 7: 5, 1938. 425

*** Aytoniaceae Cavers

by D.G. Long

The genera of Aytoniaceae follow the treatment of with the exception of Asterella and Mannia which were re-defined by.

*** Asterella P.Beauv., Dict. Sci. Nat. [F. Cuvier] 3: 257, 1805 nom. conserv. 426

** subg. Asterella

** sect. Asterella

*** Asterella tenella (L.) P.Beauv., Dict. Sci. Nat. [F. Cuvier] 3: 258, 1805. Bas.: Marchantia tenella L., Sp. Pl. 1: 1137, 1753.

** sect. Brachyblepharis (Nees) D.G.Long, J. Bryol. 22 (2): 113, 2000. Bas.: Fimbraria subg. Brachyblepharis Nees, Syn. Hepat. 4: 569, 1846.

*** Asterella abyssinica (Gottsche) Grolle, Explor. Hydrobiol. Lac Bangweolo Luapula: 170, 1972. Bas.: Fimbraria abyssinica Gottsche, Syn. Hepat. 4: 569, 1846.

*** Asterella africana (Mont.) Underw. ex A.Evans, Contr. U.S. Natl. Herb. 20: 250, 1920. Bas.: Fimbraria africana Mont., Hist. Nat. Îles Canaries 3 (2): 61, 1840.

*** Asterella blumeana (Nees) Kachroo, J. Gauhati India Univ. 3: 130, 1952. Bas.: Fimbraria blumeana Nees, Syn. Hepat. 4: 564, 1846.

*** Asterella chilensis (Nees et Mont.) A.Evans, Bull. Torrey Bot. Club 46 (12): 469, 1919. Bas.: Fimbraria chilensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 41, 1838.

*** Asterella cruciata (Steph.) Horik., Hikobia 1 (2): 79, 1951. Bas.: Fimbraria cruciata Steph., Sp. Hepat. (Stephani) 6: 12, 1917.

*** Asterella dissoluta (Steph.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 38 (2): 237, 1989. Bas.: Fimbraria dissoluta Steph., Pflanzenw. Ost-Afrikas C: 62, 1895.

*** Asterella dominicensis S.W.Arnell, Bryologist 61 (2): 140, 1958.

*** Asterella khasyana (Griff.) Grolle, Khumbu Himal 1 (4): 267, 1966. Bas.: Octokepos khasyanus Griff., Not. pl. asiat. 2: 343, 1849.

*** Asterella leptophylla (Mont.) Grolle, Feddes Repert. 87 (3/4): 246, 1976. Bas.: Fimbraria leptophylla Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 16, 1842.

*** Asterella limbata D.G.Long et Grolle, J. Bryol. 18 (2): 287, 1994.

** Asterella shimizuana Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 10 (3): 361, 1967. 427

*** Asterella tenera (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963. Bas.: Fimbraria tenera Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 170, 1855.

* Asterella tenerrima (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981. Bas.: Fimbraria tenerrima Steph., Sp. Hepat. (Stephani) 6: 17, 1917. 428

*** Asterella venosa (Lehm. et Lindenb.) A.Evans, Contr. U.S. Natl. Herb. 20: 286, 1920. Bas.: Fimbraria venosa Lehm. et Lindenb., Nov. Stirp. Pug. 4: 29, 1832.

** subg. Phragmoblepharis Grolle, Feddes Repert. 87 (3/4): 246, 1976.

*** Asterella australis (Hook.f. et Taylor) Verd., Ann. Bryol. 5: 126, 1932. Bas.: Fimbraria australis Hook.f. et Taylor, London J. Bot. 3: 573, 1844.

*** Asterella bachmannii (Steph.) S.W.Arnell, Hepat. South Africa: 62, 1963. Bas.: Fimbraria bachmannii Steph., Hedwigia 33 (1): 7, 1894.

*** Asterella bolanderi (Austin) Underw., Bot. Gaz. 20 (2): 61, 1895. Bas.: Fimbraria bolanderi Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 230, 1869.

* Asterella caucasica (Steph.) H.Buch, A.Evans et Verd., Ann. Bryol. 10: 8, 1937 [1938]. Bas.: Fimbraria caucasica Steph., Bull. Herb. Boissier 7 (3): 206 (132), 1899. 429

*** Asterella conocephala (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963. Bas.: Fimbraria conocephala Steph., Bull. Herb. Boissier 7 (3): 205 (131), 1899.

** Asterella coronata (Steph.) H.A.Mill., Phytologia 47 (4): 319, 1981. Bas.: Fimbraria coronata Steph., Sp. Hepat. (Stephani) 6: 12, 1917.

** Asterella dioica (Steph.) H.A.Mill., Phytologia 47 (4): 319, 1981. Bas.: Fimbraria dioica Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 104, 1914.

* Asterella dognyensis H.A.Mill., Phytologia 47 (4): 319, 1981. Nom. nov. pro Fimbraria umbonata Steph., Sp. Hepat. (Stephani) 6: 17, 1917, nom. illeg. 430

*** Asterella drummondii (Taylor) R.M.Schust. ex D.G.Long, J. Bryol. 21 (1): 76, 1999. Bas.: Fimbraria drummondii Taylor, London J. Bot. 5: 412, 1846.

*** Asterella echinella (Gottsche) Underw., Bot. Gaz. 20 (2): 62, 1895. Bas.: Fimbraria echinella Gottsche, Mexik. Leverm.: 271, 1863.

*** Asterella elegans (Spreng.) Trevis., Rendiconti Reale Ist. Lombardo Sci. (ser. 2) 7: 785, 1874. Bas.: Fimbraria elegans Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (1): 235, 1827.

*** Asterella heteroflora (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981. Bas.: Fimbraria heteroflora Steph., Sp. Hepat. (Stephani) 6: 14, 1917.

*** Asterella innovans (Austin) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 529, 1963. Bas.: Marchantia innovans Austin, Bull. Torrey Bot. Club 5 (3): 14, 1874.

*** Asterella lateralis M.Howe, Bull. Torrey Bot. Club 25 (4): 189, 1898.

*** Asterella lindenbergiana (Corda ex Nees) Lindb. ex Arnell, Lebermoosstud. nördl. Norwegen: 2, 1892. Bas.: Fimbraria lindenbergiana Corda ex Nees, Naturgesch. Eur. Leberm. 4: 266, 1838.

*** Asterella linearis (Steph.) M.Howe, Bull. Torrey Bot. Club 25 (4): 191, 1898. Bas.: Fimbraria linearis Steph., Bot. Jahrb. Syst. 20 (3): 302, 1895.

*** Asterella longebarbata (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981. Bas.: Fimbraria longebarbata Steph., Hedwigia 28 (3): 156, 1889.

*** Asterella macropoda (Spruce) A.Evans, Bull. Torrey Bot. Club 46 (12): 472, 1919. Bas.: Fimbraria macropoda Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 564, 1885.

*** Asterella marginata (Nees) S.W.Arnell, Hepat. South Africa: 63, 1963. Bas.: Fimbraria marginata Nees, Horae Phys. Berol.: 44, 1820.

** Asterella muelleri (Gottsche) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963. Bas.: Fimbraria muelleri Gottsche, Bull. Herb. Boissier 7 (3): 203 (129), 1899.

*** Asterella multiflora (Steph.) Kachroo, J. Hattori Bot. Lab. 19: 3, 1958. Bas.: Fimbraria multiflora Steph., Bull. Herb. Boissier 7 (3): 198 (124), 1899.

*** Asterella mussuriensis (Kashyap) Verd., Ann. Bryol. 8: 156, 1935. Bas.: Fimbraria mussuriensis Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 345, 1916.

*** Asterella mussuriensis subsp. crassa (Shimizu et S.Hatt.) D.G.Long, Lindbergia 26 (1): 44, 2001. Bas.: Asterella crassa Shimizu et S.Hatt., J. Hattori Bot. Lab. 8: 48, 1952.

*** Asterella pappii (Gola) Grolle, Feddes Repert. 87 (3/4): 246, 1976. Bas.: Fimbraria pappii Gola, Ann. Bot. (Rome) 13 (1): 65, 1914.

*** Asterella persica (Steph.) M.Howe, Bull. Torrey Bot. Club 25 (4): 191, 1898. Bas.: Fimbraria persica Steph., Hedwigia 33 (1): 7, 1894.

* Asterella preussii (Schiffn.) M.Howe, Bull. Torrey Bot. Club 25 (4): 191, 1898. Bas.: Fimbraria preussii Schiffn., Bot. Jahrb. Syst. 20 (3): 303, 1895. 431

** Asterella setisquama (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963. Bas.: Fimbraria setisquama Steph., Hedwigia 28 (3): 156, 1889.

*** Asterella syngenesica (Bory) Grolle, Lindbergia 2 (3/4): 230, 1974. Bas.: Marchantia syngenesica Bory, Voy. îles Afrique 2: 95, 1804.

** Asterella tasmanica (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963. Bas.: Fimbraria tasmanica Steph., Bull. Herb. Boissier 7 (3): 206 (132), 1899.

*** Asterella versicolor A.Evans, Contr. U.S. Natl. Herb. 20: 307, 1920.

*** Asterella vulcanica (Schiffn.) Kachroo et Bapna, J. Indian Bot. Soc. 56 (1): 75, 1977. Bas.: Hypenantron vulcanicum Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 155, 1898.

*** Asterella whiteleggeana (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963. Bas.: Fimbraria whiteleggeana Steph., Hedwigia 28 (3): 155, 1889.

*** Asterella wilmsii (Steph.) S.W.Arnell, Hepat. South Africa: 62, 1963. Bas.: Fimbraria wilmsii Steph., Hedwigia 31 (3): 122, 1892.

** subg. Saccatae (Grolle) D.G.Long, J. Bryol. 22 (2): 113, 2000. Bas.: Asterella subg. Phragmoblepharis sect. Saccatae Grolle, Feddes Repert. 87 (3/4): 346, 1976.

*** Asterella alpina (Steph.) D.G.Long, J. Hattori Bot. Lab. 93: 9, 2003. Bas.: Fimbraria alpina Steph., Bull. Herb. Boissier 7 (3): 211 (137), 1899.

*** Asterella grollei D.G.Long, Bryologist 102 (2): 169, 1999.

*** Asterella muscicola (Steph.) S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 263, 1957. Bas.: Fimbraria muscicola Steph., Hedwigia 31 (3): 121, 1892.

*** Asterella palmeri (Austin) Underw., Bot. Gaz. 20 (2): 63, 1895. Bas.: Fimbraria palmeri Austin, Bull. Torrey Bot. Club 6 (7): 47, 1875.

*** Asterella pringlei Underw., Bot. Gaz. 20 (2): 64, 1895.

*** Asterella rugosa A.Evans, Contr. U.S. Natl. Herb. 20: 289, 1920.

*** Asterella saccata (Wahlenb.) A.Evans, Contr. U.S. Natl. Herb. 20: 276, 1920. Bas.: Marchantia saccata Wahlenb., Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 296, 1811.

** subg. Wallichianae D.G.Long, Lindbergia 26 (1): 43, 2001.

** sect. Californicae D.G.Long, J. Hattori Bot. Lab. 97: 257, 2005.

*** Asterella californica (Hampe ex Austin) Underw., Bot. Gaz. 20 (2): 60, 1895. Bas.: Fimbraria californica Hampe ex Austin, Hepat. bor.-amer.: 33, 1873.

** sect. Wallichianae D.G.Long, Phytotaxa 173 (1): 87, 2014.

*** Asterella wallichiana (Lehm. et Lindenb.) Grolle, Khumbu Himal 1 (4): 262, 1966. Bas.: Fimbraria wallichiana Lehm. et Lindenb., Nov. Stirp. Pug. 4: 4, 1832.

** Cryptomitrium Austin ex Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 36, 1884.

*** Cryptomitrium himalayense Kashyap, New Phytol. 14 (1): 2, 1915.

*** Cryptomitrium oreades Perold, Bothalia 24 (2): 149, 1994.

*** Cryptomitrium tenerum (Hook.) Austin ex Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 36, 1884. Bas.: Marchantia tenera Hook., Syn. Pl. (Kunth) 1: 45, 1822.

*** Mannia Corda, Gen. hepat.: 646, 1829 nom. conserv.

** subg. Mannia

*** Mannia androgyna (L.) A.Evans, Chron. Bot. 4: 224, 1938. Bas.: Marchantia androgyna L., Sp. Pl. 1: 1138, 1753.

*** Mannia californica (Gottsche) L.C.Wheeler, Bryologist 37 (5): 88, 1934 [1935]. Bas.: Grimaldia californica Underw., Bot. Gaz. 13 (5): 114, 1888.

*** Mannia controversa (Meyl.) D.B.Schill, Edinburgh J. Bot. 65 (1): 36, 2008. Bas.: Grimaldia controversa Meyl., Beitr. Kryptogamenfl. Schweiz 6 (4): 87, 1924.

*** Mannia controversa subsp. asiatica D.B.Schill et D.G.Long, Edinburgh J. Bot. 65 (1): 45, 2008.

*** Mannia fragrans (Balb.) Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (1): 62, 1937. Bas.: Marchantia fragrans Balb., Mem. Acad. Sci. Turin, Sci. Phys. 15: 76, 1804.

** Mannia fragrans subsp. orientalis R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 201, 1992. Nom. nov. pro Mannia barbifrons Shimizu et S.Hatt., J. Hattori Bot. Lab. 10: 49, 1953.

* Mannia perssonii Udar et V.Chandra, Canad. J. Bot. 43 (1): 150, 1965. 432

*** Mannia sibirica (Müll.Frib.) Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (1): 66, 1937. Bas.: Grimaldia pilosa var. sibirica Müll.Frib., Lebermoose 1 (5): 265, 1907.

** subg. Neesiella (Schiffn.) D.B.Schill et D.G.Long, Bryologist 113 (1): 175, 2010. Bas.: Neesiella Schiffn., Hepat. (Engl.-Prantl): 32, 1893.

*** Mannia gracilis (F.Weber) D.B.Schill et D.G.Long, Bryologist 113 (1): 173, 2010. Bas.: Marchantia gracilis F.Weber, Hist. Musc. Hepat. Prodr.: 105, 1815.

* Mannia hegewaldii Bischl., Fl. Neotrop. Monogr. 97: 184, 2005. 433

*** Mannia pilosa (Hornem.) Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (1): 64, 1937. Bas.: Marchantia pilosa Hornem., Fl. Danica 8 (24): 7, tab. 1426, 1810.

*** Mannia triandra (Scop.) Grolle, J. Bryol. 8 (4): 487, 1975. Bas.: Marchantia triandra Scop., Fl. Carniol. (ed. 2) 2: 354, 1772.

Incertae sedis

* Mannia paradoxa R.M.Schust., Phytologia 57 (6): 410, 1985. 434

*** Plagiochasma Lehm., Nov. Stirp. Pug. 4: 13, 1832 nom. conserv.

* Plagiochasma megacarpon (Griff.) Steph., Bull. Herb. Boissier 6 (10): 789 (86), 1898. Bas.: Antrocephalus megacarpon Griff., Not. pl. asiat. 2: 338, 1849. 435

** subg. Micropylum Bischl., Rev. Bryol. Lichénol. 43 (1): 103, 1977.

*** Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph., Bull. Herb. Boissier 6 (10): 783 (80), 1898. Bas.: Aytonia rupestris J.R.Forst. et G.Forst., Char. gen. pl., ed. 2: 148, 1776.

*** Plagiochasma rupestre var. volkii Bischl., Rev. Bryol. Lichénol. 44 (3): 289, 1978.

** subg. Plagiochasma

*** Plagiochasma appendiculatum Lehm. et Lindenb., Nov. Stirp. Pug. 4: 14, 1832.

*** Plagiochasma argentinicum Bischl., Rev. Bryol. Lichénol. 45 (3): 301, 1979.

*** Plagiochasma beccarianum Steph., Bull. Herb. Boissier 6 (10): 781 (78), 1898.

*** Plagiochasma cordatum Lehm. et Lindenb., Nov. Stirp. Pug. 4: 13, 1832.

*** Plagiochasma crenulatum Gottsche, Mexik. Leverm.: 266, 1863.

*** Plagiochasma cuneatum A.Evans, Amer. J. Bot. 19 (7): 627, 1932.

*** Plagiochasma eximium (Schiffn.) Steph., Bull. Herb. Boissier 6 (10): 781 (78), 1898. Bas.: Aytonia eximia Schiffn., Bot. Jahrb. Syst. 20 (3): 300, 1895.

*** Plagiochasma intermedium Lindenb. et Gottsche, Syn. Hepat. 4: 513, 1846.

*** Plagiochasma jamaicense (Haynes) A.Evans, Bull. Torrey Bot. Club 42 (5): 292, 1915. Bas.: Aytonia jamaicensis Haynes, Bull. Torrey Bot. Club 34 (2): 58, 1907.

*** Plagiochasma japonicum (Steph.) C.Massal., Hepat. Shen-si: 47, 1897. Bas.: Aytonia japonica Steph., Bull. Herb. Boissier 5 (2): 84, 1897.

*** Plagiochasma landii A.Evans, Bull. Torrey Bot. Club 42 (5): 298, 1915.

*** Plagiochasma microcephalum (Steph.) Steph., Bull. Herb. Boissier 6 (10): 781 (78), 1898. Bas.: Aytonia microcephala Steph., Bot. Jahrb. Syst. 20 (3): 301, 1895.

*** Plagiochasma microcephalum var. tunesicum Bischl., Rev. Bryol. Lichénol. 44 (3): 247, 1978.

*** Plagiochasma muenchianum Steph., Sp. Hepat. (Stephani) 6: 9, 1917.

*** Plagiochasma pterospermum C.Massal., Hepat. Shen-si: 46, 1897.

*** Plagiochasma wrightii Sull., Musc. Hepat. U.S.: 688, 1856.

Incertae sedis

** Plagiochasma udarii A.Alam et S.C.Srivast., Indian J. Forest. 32 (4): 631, 2009.

*** Reboulia Raddi, Opusc. Sci. 2 (6): 357, 1818 nom. conserv.

*** Reboulia hemisphaerica (L.) Raddi, Opusc. Sci. 2 (6): 357, 1818. Bas.: Marchantia hemisphaerica L., Sp. Pl. 1: 1138, 1753.

** Reboulia hemisphaerica subsp. acrogyna (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 168, 1992. Bas.: Asterella bolanderi subsp. acrogyna R.M.Schust., Phytologia 57 (6): 410, 1985.

** Reboulia hemisphaerica subsp. australis R.M.Schust., Phytologia 56 (7): 460, 1985.

** Reboulia hemisphaerica subsp. dioica R.M.Schust., Phytologia 56 (7): 462, 1985.

** Reboulia hemisphaerica var. fissisquama Herzog, Symb. Sin. 5: 5, 1930.

** Reboulia hemisphaerica subsp. orientalis R.M.Schust., Phytologia 56 (7): 461, 1985.

** Reboulia hemisphaerica var. turkestanica C.E.O.Jensen ex Herzog, Symb. Sin. 5: 5, 1930.

*** Cleveaceae Cavers

by D.G. Long

The genera of Cleveaceae were re-defined by based on molecular evidence.

*** Athalamia Falc., Ann. Mag. Nat. Hist. (ser. 2) 1 (5): 375, 1848.

* Athalamia dioica Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 348, 1916.

*** Athalamia pinguis Falc., Ann. Mag. Nat. Hist. (ser. 2) 1 (5): 375, 1848.

* Athalamia pulcherrima (Steph.) S.Hatt., J. Hattori Bot. Lab. 12: 54, 1954. Bas.: Clevea pulcherrima Steph., Bot. Jahrb. Syst. 20 (3): 303, 1895.

*** Clevea Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 289, 1868.

*** Clevea hyalina (Sommerf.) Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 291, 1868. Bas.: Marchantia hyalina Sommerf., Mag. Naturvidensk. 11 (2): 234, 1833.

* Clevea hyalina var. californica M.Howe, Mem. Torrey Bot. Club 7: 38, 1899.

* Clevea pedicellata (Griff.) Lindb., Acta Soc. Fauna Fl. Fenn. 2 (3): 11, 1882. Bas.: Plagiochasma pedicellatum Griff., Not. pl. asiat. 2: 331, 1849.

*** Clevea pusilla (Steph.) Rubas. et D.G.Long, J. Bryol. 33 (2): 167, 2011. Bas.: Gollaniella pusilla Steph., Hedwigia 44 (2): 74, 1905.

*** Clevea spathysii (Lindenb.) Müll.Frib., Hedwigia 79 (1/2): 75, 1940. Bas.: Marchantia spathysii Lindenb., Syn. hepat. eur: 104, 1829.

*** Peltolepis Lindb., Morgonbladet (Helsinki) 1876 (106, 9 May): 1, 1876.

** Peltolepis japonica (Shimizu et S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 14: 103, 1955. Bas.: Peltolepis quadrata var. japonica Shimizu et S.Hatt., J. Hattori Bot. Lab. 12: 69, 1954.

*** Peltolepis quadrata (Saut.) Müll.Frib., Hedwigia 79 (1/2): 74, 1940. Bas.: Sauteria quadrata Saut., Flora 43 (22): 351, 1860.

*** Sauteria Nees, Naturgesch. Eur. Leberm. 4: 139, 1838.

** sect. Sauchia (Kashyap) R.M.Schust., Phytologia 57 (6): 411, 1985. Bas.: Sauchia Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 347, 1916.

* Sauteria japonica (Shimizu et S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 12: 62, 1954. Bas.: Sauchia japonica Shimizu et S.Hatt., J. Hattori Bot. Lab. 9: 32, 1953. 436

*** Sauteria spongiosa (Kashyap) S.Hatt., J. Hattori Bot. Lab. 12: 62, 1954. Bas.: Sauchia spongiosa Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 347, 1916.

** sect. Sauteria

*** Sauteria alpina (Nees) Nees, Naturgesch. Eur. Leberm. 4: 143, 1838. Bas.: Lunularia alpina Nees, Flora 13 (25): 399, 1830.

* Sauteria inflata C.Gao et K.C.Chang, Acta Bot. Yunnan. 3 (4): 389, 1981. 437

Incertae sedis

* Sauteria chilensis (Lindenb.) Grolle, J. Hattori Bot. Lab. 58: 200, 1985. Bas.: Grimaldia chilensis Lindenb., Voy. Amér. Mérid. 7 (2): 53, 1839.

* Sauteria crassipes Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 229, 1869.

* Sauteria nyikaensis Perold, Bothalia 33 (2): 167, 2003.

*** Conocephalaceae Müll.Frib. ex Grolle

by D.G. Long

*** Conocephalum Hill, Gener. Nat. Hist. 2 Hist. pl. (ed. 2): 118, 1773 nom. conserv.

** subg. Conocephalum

*** Conocephalum conicum (L.) Dumort., Commentat. Bot. (Dumortier): 115, 1822. Bas.: Marchantia conica L., Sp. Pl. 1: 1138, 1753.

*** Conocephalum salebrosum Szweyk., Buczk. et Odrzyk., Pl. Syst. Evol. 253 (1/4): 146, 2005.

** subg. Sandea (Lindb.) Inoue, Ill. Jap. Hep. 2: 192, 1976. Bas.: Sandea Lindb., Acta Soc. Fauna Fl. Fenn. 2 (5): 3, 1884.

*** Conocephalum japonicum (Thunb.) Grolle, J. Hattori Bot. Lab. 55: 501, 1984. Bas.: Lichen japonicus Thunb., Fl. Jap. (Thunberg): 344, 1784.

** Corsiniaceae Engl.

by D.G. Long

** Corsinioideae Schiffn.

*** Corsinia Raddi, Opusc. Sci. 2 (6): 354, 1818.

*** Corsinia coriandrina (Spreng.) Lindb., Hepaticol. Utveckl.: 30, 1877. Bas.: Riccia coriandrina Spreng. Anleit. Kenntn. Gew. 3: 320, 1804.

*** Cronisioideae R.M.Schust.

*** Cronisia Berk., Introd. crypt. bot.: 434, 1857.

*** Cronisia fimbriata (Nees) Whittem. et Bischl., Cryptog. Bryol. 22 (3): 170, 2001. Bas.: Riccia fimbriata Nees, Fl. Bras. (Martius) 1 (1): 301, 1833.

*** Cronisia weddellii (Mont.) Grolle, J. Bryol. 9 (4): 532, 1977 [1978]. Bas.: Boschia weddellii Mont., Ann. Sci. Nat. Bot. (sér. 4) 5: 352, 1856.

*** Cyathodiaceae Stotler et Crand.-Stotl.

by D.G. Long

*** Cyathodium Kunze, Nov. Stirp. Pug. 6: 17, 1834.

*** Cyathodium aureonitens (Griff.) Mitten, J. Linn. Soc., Bot. 22 (146): 327. Bas.: Synhymenium aureonitens Griff., Not. pl. asiat. 2: 344, 1849.

*** Cyathodium bischlerianum N.Salazar, Bryologist 104 (1): 141, 2001 (Salazar 2001).

*** Cyathodium cavernarum Kunze, Nov. Stirp. Pug. 6: 18, 1834.

** Cyathodium denticulatum Udar et S.C.Srivast., Geophytology 1 (2): 166, 1971.

*** Cyathodium foetidissimum Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 154, 1898.

*** Cyathodium indicum Udar et D.K.Singh, J. Bryol. 10 (2): 139, 1978 [1979].

*** Cyathodium mehranum D.K.Singh, Misc. Bryol. Lichenol. 9 (8): 173, 1983.

*** Cyathodium smaragdinum Schiffn., Ann. Jard. Bot. Buitenzorg, suppl. 3: 480, 1910.

*** Cyathodium spruceanum Prosk., Bryologist 54 (4): 243, 1951 [1952].

* Cyathodium spurium (Dicks.) Lindb. ex Braithw., J. Bot. 16: 55, 1878. Bas.: Riccia spuria Dicks., Fasc. Pl. Crypt. Brit. 4: 20, 1801. 438

*** Cyathodium steerei Hässel, Rev. Bryol. Lichénol. 30 (3/4): 223, 1961.

*** Cyathodium tuberculatum Udar et D.K.Singh, Bryologist 79 (2): 235, 1976.

*** Cyathodium tuberosum Kashyap, New Phytol. 13 (6/7): 210, 1914.

*** Dumortieraceae D.G.Long

by D.G. Long

*** Dumortiera Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 410, 1824 [1825].

*** Dumortiera hirsuta (Sw.) Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 410, 1824 [1825]. Bas.: Marchantia hirsuta Sw., Prodr. (Swartz): 145, 1788. 439

* Dumortiera hirsuta subsp. nepalensis (Taylor) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 386, 1992. Bas.: Hygrophila nepalensis Taylor, Trans. Linn. Soc. London 17 (3): 392, 1836.

* Dumortiera hirsuta subsp. tatunoi Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 6: 38, 1951.

*** Exormothecaceae Müll.Frib. ex Grolle

by D.G. Long

*** Aitchisoniella Kashyap, New Phytol. 13 (6/7): 219, 1914.

*** Aitchisoniella himalayensis Kashyap, New Phytol. 13 (6/7): 219, 1914.

*** Exormotheca Mitt., Nat. hist. Azores: 325, 1870.

** subg. Corbierella (Douin et Trab.) Schiffn., Hedwigia 81 (1/2): 71, 1942. Bas.: Corbierella Douin et Trab., Rev. Gén. Bot. 31: 326, 1919.

*** Exormotheca bischlerae Furuki et Higuchi, Cryptog. Bryol. 27 (1): 98, 2006.

*** Exormotheca holstii Steph., Bull. Herb. Boissier 7 (3): 219 (145), 1899.

*** Exormotheca welwitschii Steph., Bull. Herb. Boissier 7 (3): 220 (146), 1899.

** subg. Exormotheca

*** Exormotheca pustulosa Mitt., Nat. hist. Azores: 326, 1870.

Incertae sedis

*** Exormotheca bulbigena Bornefeld, O.H.Volk et R.Wolf, Bothalia 26 (2): 159, 1996.

*** Exormotheca ceylonensis Meijer, J. Hattori Bot. Lab. 16: 72, 1956.

* Exormotheca gollanii Steph., Sp. Hepat. (Stephani) 6: 18, 1917.

*** Exormotheca tuberifera Kashyap, New Phytol. 13 (9): 309, 1914.

*** Stephensoniella Kashyap, New Phytol. 13 (9): 312, 1914.

*** Stephensoniella brevipedunculata Kashyap, New Phytol. 13 (9): 312, 1914.

*** Marchantiaceae Lindl.

by D.G. Long

The treatment of Marchantiaceae is mainly following, ) and.

** Bucegioideae R.M.Schust.

*** Bucegia Radian, Bull. Herb. Inst. Bot. Bucarest 3-4: 3, 1903.

*** Bucegia romanica Radian, Bull. Herb. Inst. Bot. Bucarest 3-4: 4, 1903.

** Marchantioideae Schiffn.

*** Marchantia L., Sp. Pl. 1: 1137, 1753.

** subg. Chlamidium (Corda) Bischl., Cryptog. Bryol. Lichénol. 3 (4): 362, 1982. Bas.: Chlamidium Corda, Gen. hepat.: 647, 1829.

*** Marchantia breviloba A.Evans, Trans. Connecticut Acad. Arts 21 (3): 265, 1917.

*** Marchantia inflexa Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 43, 1838.

** sect. Chlamidium (Corda) Nees, Naturgesch. Eur. Leberm. 4: 101, 1838. Bas.: Chlamidium Corda, Gen. hepat.: 647, 1829.

*** Marchantia chenopoda L., Sp. Pl. 1: 1137, 1753.

*** Marchantia crenata Austin, Bull. Torrey Bot. Club 5 (3): 14, 1874.

*** Marchantia foliacea Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 168, 1855.

*** Marchantia formosana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 121, 1934.

*** Marchantia globosa Brid., Hist. Musc. Hepat. Prodr.: 102, 1815.

*** Marchantia hexaptera Reichardt, Verh. K.K. Zool.-Bot. Ges. Wien 16: 957, 1866.

*** Marchantia linearis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 8, 1832.

*** Marchantia miqueliana Lehm., Nov. Stirp. Pug. 10: 20, 1857.

*** Marchantia novoguineensis Bischl., Bryophyt. Biblioth. 38: 130, 1989.

*** Marchantia pappeana Lehm., Nov. Stirp. Pug. 10: 21, 1857.

** Marchantia pappeana subsp. robusta (Steph.) Bischl., Bryophyt. Biblioth. 45: 91, 1993. Bas.: Marchantia robusta Steph., Candollea 14: 111, 1953.

*** Marchantia pileata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 169, 1855.

*** Marchantia pinnata Steph., Candollea 14: 109, 1953.

*** Marchantia rubribarba Steph., Bull. Herb. Boissier 7 (5): 400 (172), 1899.

*** Marchantia vitiensis Steph., Bull. Herb. Boissier 7 (7): 520 (182), 1899.

** sect. Paleaceae Bischl., Bryophyt. Biblioth. 38: 90, 1989.

*** Marchantia paleacea Bertol., Opusc. Sci. 1: 242, 1817.

*** Marchantia paleacea subsp. diptera (Nees et Mont.) Inoue, J. Jap. Bot. 64 (7): 194, 1989. Bas.: Marchantia diptera Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 243, 1843.

** sect. Papillatae Bischl., Cryptog. Bryol. Lichénol. 10 (1): 69, 1989.

*** Marchantia debilis K.I.Goebel, Organogr. Pfl., ed. 2, 2 (1): 901, 1915.

*** Marchantia emarginata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 192, 1824 [1825].

*** Marchantia emarginata subsp. lecordiana (Steph.) Bischl., Cryptog. Bryol. Lichénol. 10 (1): 78, 1989. Bas.: Marchantia lecordiana Steph., Bull. Herb. Boissier 7 (7): 525 (187), 1899.

*** Marchantia emarginata subsp. tosana (Steph.) Bischl., Cryptog. Bryol. Lichénol. 10 (1): 77, 1989. Bas.: Marchantia tosana Steph., Bull. Herb. Boissier 5 (2): 99, 1897.

*** Marchantia papillata Raddi, Critt. Brasil.: 20, 1822.

*** Marchantia papillata subsp. grossibarba (Steph.) Bischl., Cryptog. Bryol. Lichénol. 10 (1): 78, 1989. Bas.: Marchantia grossibarba Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 221, 1894.

** subg. Marchantia

*** Marchantia plicata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 43, 1838.

** sect. Berteroanae R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 310, 1992.

*** Marchantia berteroana Lehm. et Lindenb., Nov. Stirp. Pug. 6: 21, 1834.

** sect. Marchantia

*** Marchantia polymorpha L., Sp. Pl. 1: 1137, 1753.

*** Marchantia polymorpha subsp. montivagans Bischl. et Boissel.-Dub., J. Bryol. 16 (3): 364, 1991.

*** Marchantia polymorpha subsp. ruderalis Bischl. et Boissel.-Dub., J. Bryol. 16 (3): 364, 1991.

** subg. Protomarchantia R.M.Schust., Phytologia 57 (6): 410, 1985.

** sect. Protomarchantia (R.M.Schust.) L.Söderstr., Phytotaxa 202 (1): 69, 2015. Bas.: Marchantia subg. Protomarchantia R.M.Schust., Phytologia 57 (6): 410, 1985.

*** Marchantia acaulis Steph., Bull. Herb. Boissier 7 (7): 533 (195), 1899.

*** Marchantia antiqua Steph., Candollea 14: 103, 1953.

*** Marchantia carrii Bischl., Bryophyt. Biblioth. 38: 256, 1989.

*** Marchantia geminata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 194, 1824 [1825].

*** Marchantia hartlessiana Steph., Candollea 14: 107, 1953.

*** Marchantia macropora Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 169, 1855.

*** Marchantia philippinensis Bischl., Bryophyt. Biblioth. 38: 245, 1989.

*** Marchantia solomonensis Bischl., Bryophyt. Biblioth. 38: 281, 1989.

*** Marchantia streimannii Bischl., Bryophyt. Biblioth. 38: 250, 1989.

*** Marchantia subintegra Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 125, 1860 [1861].

*** Marchantia treubii Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 160, 1898.

*** Marchantia wallisii J.B.Jack et Steph., Bull. Herb. Boissier 7 (7): 520 (182), 1899.

** sect. Subgeminatae Bischl., Bryophyt. Biblioth. 38: 219, 1989.

*** Marchantia subgeminata Steph., Bull. Herb. Boissier 7 (7): 530 (192), 1899.

Incertae sedis

* Marchantia assamica Griff., Not. pl. asiat. 2: 327, 1849. 440

* Marchantia balboi Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 2, 1916. 441

* Marchantia balboi var. acutisquamata Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 472, 1947. 442

* Marchantia cagnii Gola, Ann. Bot. (Rome) 6 (2): 271, 1907. 443

* Marchantia cengiana Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 473, 1947. 444

* Marchantia friedrichsthaliana Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 438, 1877.

* Marchantia keniae Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 3, 1916. 445

* Marchantia papyracea Gola, Ann. Bot. (Rome) 6 (2): 271, 1907. 446

* Marchantia quadriloba Steph., Candollea 14: 110, 1953.

* Marchantia sellae Gola, Ann. Bot. (Rome) 6 (2): 271, 1907. 447

** Marchantia stoloniscyphulus (C.Gao et K.C.Chang) Piippo, J. Hattori Bot. Lab. 68: 134, 1990. Bas.: Marchantiopsis stoloniscyphulus C.Gao et K.C.Chang, Bull. Bot. Res., Harbin 2 (4): 114, 1982.

* Marchantia trilocularis Roth, Tent. Fl. Germ. 1: 487, 1788. 448

* Marchantia tusui Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 3, 1916. 449

** Preissia Corda, Gen. hepat.: 647, 1829.

*** Preissia quadrata (Scop.) Nees, Naturgesch. Eur. Leberm. 4: 135, 1838. Bas.: Marchantia quadrata Scop., Fl. Carniol. (ed. 2) 2: 355, 1772.

** Preissia quadrata subsp. hyperborea R.M.Schust., Phytologia 57 (6): 410, 1985.

*** Monocleaceae A.B.Frank

by D.G. Long

*** Monoclea Hook., Musci Exot. 2: tab. clxxiv, 1820.

*** Monoclea forsteri Hook., Musci Exot. 2: tab. clxxiv, 1820.

*** Monoclea gottschei Lindb., Rev. Bryol. 13 (6): 102, 1886.

*** Monoclea gottschei subsp. elongata Gradst. et Mues, Pl. Syst. Evol. 180 (1/2): 133, 1992.

*** Monosoleniaceae Inoue

by D.G.Long

*** Monosolenium Griff., Not. pl. asiat. 2: 341, 1849.

*** Monosolenium tenerum Griff., Not. pl. asiat. 2: 341, 1849.

*** Oxymitraceae Müll.Frib. ex Grolle

by D.G. Long

*** Oxymitra Bisch. ex Lindenb., Syn. hepat. eur: 124, 1829.

*** Oxymitra cristata Garside, Bothalia 23 (2): 211, 1993. Based on: Oxymitra cristata Garside, J. S. African Bot. 24: 83, 1958, nom. inval.

*** Oxymitra incrassata (Brot.) Sérgio et Sim-Sim, J. Bryol. 15 (4): 662, 1989. Bas.: Riccia incrassata Brot., Fl. lusit. 2: 428, 1804 [1805].

*** Ricciaceae Rchb.

by R. Stotler, B.J. Crandall-Stotler and D.C. Cargill

** Riccia L., Sp. Pl. 1: 1138, 1753 nom. conserv.

** subg. Chartaceae Perold, Bothalia 16 (1): 29, 1986.

*** Riccia schelpei O.H.Volk et Perold, Bothalia 16 (1): 29, 1986.

** subg. Leptoriccia R.M.Schust., Phytologia 56 (2): 72, 1984.

*** Riccia membranacea Gottsche et Lindenb., Syn. Hepat. 4: 608, 1846.

** subg. Riccia

*** Riccia albida Sull. ex Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 231, 1869.

*** Riccia albopunctata Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 237, 1991.

*** Riccia australis Steph., Bull. Herb. Boissier 6 (4): 337 (29), 1898.

*** Riccia boliviensis Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 242, 1991.

*** Riccia brasiliensis Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 6, 1964.

*** Riccia breutelii Hampe, Bull. Herb. Boissier 6 (4): 325 (17), 1898.

*** Riccia brittonii M.Howe, Ann. Missouri Bot. Gard. 2 (1/2): 50, 1915.

* Riccia chudoana Steph., Sp. Hepat. (Stephani) 6: 1, 1917.

*** Riccia coracina Jovet-Ast, Cryptog. Bryol. 24 (3): 212, 2003.

*** Riccia corrugata Jovet-Ast, Cryptog. Bryol. 21 (4): 308, 2000.

*** Riccia crassivenia Jovet-Ast, Cryptog. Bryol. 21 (4): 312, 2000.

*** Riccia cubensis S.W.Arnell, Bryologist 61 (2): 142, 1958.

*** Riccia discolor Lehm. et Lindenb., Nov. Stirp. Pug. 4: 1, 1832.

*** Riccia ekmanii S.W.Arnell, Bryologist 61 (2): 143, 1958.

*** Riccia elliottii Steph., Bull. Herb. Boissier 6 (4): 324 (16), 1898.

*** Riccia enyae Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 230, 1991.

*** Riccia erythrocarpa Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 257, 1991.

*** Riccia fruchartii Steph., Bull. Herb. Boissier 6 (4): 330 (22), 1898.

*** Riccia gangetica Ahmad ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 57, 2012. Based on: Riccia gangetica Ahmad, Curr. Sci. 11 (11): 433, 1942, nom. inval.

*** Riccia grandis Nees, Fl. Bras. (Martius) 1 (1): 300, 1833.

*** Riccia helenae Jovet-Ast, J. Hattori Bot. Lab. 74: 96, 1993.

** Riccia hirta (Austin) Underw., Bot. Gaz. 19 (7): 274, 1894. Bas.: Riccia arvensis var. hirta Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 232, 1869.

*** Riccia horrida Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 226, 1991.

*** Riccia hortorum Bory, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18 (1): 435, 1836 [1837].

*** Riccia howellii M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 202, 1934.

* Riccia ianthina Jovet-Ast, Rev. Bryol. Lichénol. 44 (4): 418, 1978. 450

*** Riccia inflexa Taylor, London J. Bot. 5: 417, 1846.

*** Riccia iodocheila M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 200, 1934.

*** Riccia lanceolata Steph., Hedwigia 27 (3/4): 110, 1888.

*** Riccia lindmanii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 29, 1897.

** Riccia macallisteri M.Howe, Bryologist 20 (3): 35, 1917.

*** Riccia macrospora Steph., Bull. Herb. Boissier 6 (4): 328 (20), 1898.

** Riccia mamillata Trab. ex Steph., Rev. Bryol. 16 (5): 65, 1889.

*** Riccia mauryana Steph., Bull. Herb. Boissier 6 (4): 327 (19), 1898.

*** Riccia olgensis Na-Thalang, Brunonia 3 (1): 100, 1980.

*** Riccia planobiconvexa Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 29, 1897.

*** Riccia ridleyi A.Gepp, J. Linn. Soc., Bot. 27 (181): 74, 1890.

*** Riccia sanguineisporis Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 253, 1991.

*** Riccia squamata Nees, Fl. Bras. (Martius) 1 (1): 302, 1833.

*** Riccia subdepilata Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 228, 1991.

*** Riccia subplana Steph., Symb. Antill. (Urban) 3 (2): 275, 1902.

*** Riccia taeniiformis Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 270, 1991.

*** Riccia viannae Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 261, 1991.

*** Riccia vitalii Jovet-Ast, Mem. New York Bot. Gard. 45: 285, 1987.

*** Riccia weinionis Steph., Bull. Herb. Boissier 6 (4): 326 (18), 1898.

** sect. Pilifer O.H.Volk, Mitt. Bot. Staatssamml. München 19: 455, 1983.

*** Riccia alatospora O.H.Volk et Perold, Bothalia 15 (3/4): 534, 1985.

*** Riccia albomarginata Bisch. ex C.Krauss, Flora 29 (9): 135, 1846.

*** Riccia albovestita O.H.Volk, Mitt. Bot. Staatssamml. München 17: 245, 1981.

*** Riccia ampullacea Perold, Bothalia 20 (2): 168, 1990.

*** Riccia concava Bisch. ex C.Krauss, Flora 29 (9): 135, 1846.

*** Riccia elongata Perold, Bothalia 20 (2): 167, 1990.

*** Riccia furfuracea Perold, Bothalia 20 (2): 176, 1990.

*** Riccia hantamensis Perold, Bothalia 19 (2): 157, 1989.

*** Riccia hirsuta O.H.Volk et Perold, Bothalia 16 (2): 187, 1986.

*** Riccia namaquensis Perold, Bothalia 20 (2): 180, 1990.

*** Riccia parvoareolata O.H.Volk et Perold, Bothalia 15 (1/2): 117, 1984.

*** Riccia pulveracea Perold, Bothalia 20 (2): 185, 1990.

*** Riccia radicosa Pearson, Natuurw. Tijdschr. 4 (5/6): 142, 1922.

*** Riccia simii Perold, Bothalia 20 (1): 36, 1990.

*** Riccia trachyglossa Perold, Bothalia 20 (2): 172, 1990.

*** Riccia villosa Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 724, 1913.

*** Riccia vitrea Perold, Bothalia 20 (2): 178, 1990.

** sect. Riccia

*** Riccia albolimbata S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 264, 1957.

*** Riccia alboporosa Perold, Bothalia 19 (1): 12, 1989.

*** Riccia albornata O.H.Volk et Perold, Bothalia 18 (2): 160, 1988.

*** Riccia angolensis Steph., Bull. Herb. Boissier 6 (4): 323 (15), 1898.

*** Riccia argenteolimbata O.H.Volk et Perold, Bothalia 18 (2): 155, 1988.

** Riccia atlantica Sérgio et Perold, J. Bryol. 17 (1): 127, 1992.

*** Riccia atromarginata Levier, Nuovo Giorn. Bot. Ital. 21 (2): 291, 1889.

** Riccia atromarginata var. jovet-astiae Rauh et Buchloh, Rev. Bryol. Lichénol. 30 (1/2): 77, 1961.

*** Riccia atropurpurea Sim, Trans. Roy. Soc. South Africa 15 (1): 11, 1926.

*** Riccia beyrichiana Hampe, Nov. Stirp. Pug. 7: 1, 1838.

*** Riccia bicarinata Lindb., Rev. Bryol. 4 (3): 41, 1877.

*** Riccia bicolorata Perold, Bothalia 20 (2): 188, 1990.

*** Riccia bifurca Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 95, 1795 [1796].

*** Riccia billardierei Mont. et Nees, Syn. Hepat. 4: 602, 1846.

*** Riccia breidleri Jur. ex Steph., Hedwigia 24 (1): 6, 1885.

*** Riccia californica Austin, Bull. Torrey Bot. Club 6 (7): 46, 1875.

*** Riccia campbelliana M.Howe, Mem. Torrey Bot. Club 7: 26, 1899.

*** Riccia ciliata Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 95, 1795 [1796].

*** Riccia ciliifera Link, Syn. hepat. eur: 119, 1829.

* Riccia congoana Steph., Bull. Herb. Boissier 6 (4): 328 (20), 1898. 451

*** Riccia crinita Taylor, London J. Bot. 5: 415, 1846.

*** Riccia crozalsii Levier, Rev. Bryol. 29 (4): 73, 1902.

* Riccia crustata Trab., Bull. Soc. Hist. Nat. Afrique N. 7: 87, 1916. 452

** Riccia dictyospora M.Howe, Bull. Torrey Bot. Club 28 (3): 163, 1901.

*** Riccia glauca L., Sp. Pl. 1: 1139, 1753.

** Riccia glauca var. ciliaris Warnst., Verh. Bot. Vereins Prov. Brandenburg 27 (1): 87, 1886.

** Riccia gothica Damsh. et Hallingb., Lindbergia 12 (2/3): 100, 1986 [1987].

*** Riccia gougetiana Durieu et Mont., Ann. Sci. Nat. Bot. (sér. 3) 11: 35, 1849.

** Riccia gougetiana var. armatissima Levier ex Müll.Frib., Lebermoose 1 (3): 161, 1907.

*** Riccia lamellosa Raddi, Opusc. Sci. 2 (6): 351, 1818.

*** Riccia ligula Steph., Bull. Herb. Boissier 6 (4): 315 (7), 1898.

*** Riccia limbata Bisch. ex C.Krauss, Flora 29 (9): 135, 1846.

*** Riccia macrocarpa Levier, Bull. Soc. Bot. Ital. 1894: 114, 1894.

*** Riccia mammifera O.H.Volk et Perold, Bothalia 16 (2): 176, 1986.

*** Riccia melitensis C.Massal., Bull. Soc. Bot. Ital. 1913 (2/3): 52, 1913.

*** Riccia michelii Raddi, Opusc. Sci. 2 (6): 352, 1818.

*** Riccia microciliata O.H.Volk et Perold, Bothalia 16 (2): 173, 1986.

*** Riccia montana Perold, Bothalia 19 (1): 9, 1989.

*** Riccia natalensis Sim, Trans. Roy. Soc. South Africa 15 (1): 9, 1926.

*** Riccia nigrella DC., Fl. Franç. (DC. & Lamarck), 5 (6): 193, 1815.

*** Riccia okahandjana S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 268, 1957.

** Riccia ozarkiana McGregor, Bryologist 63 (1): 30, 1960.

*** Riccia papillosa Moris, Stirp. Sard. Elench.: 18, 1829.

*** Riccia pottsiana Sim, Trans. Roy. Soc. South Africa 15 (1): 10, 1926.

*** Riccia rosea O.H.Volk et Perold, Bothalia 16 (2): 181, 1986.

* Riccia runssorensis Steph., Bull. Herb. Boissier 6 (4): 330 (22), 1898. 453

*** Riccia sommieri Levier, Isola Giglio: 119, 1900.

*** Riccia sorocarpa Bisch., Bem. Leberm.: 145, 1835.

** Riccia sorocarpa var. heegii Schiffn., Hedwigia 53 (1/2): 36, 1912.

*** Riccia subbifurca Warnst. ex Croz., Rev. Bryol. 30 (4): 62, 1903.

** Riccia tenella D.L.Jacobs, Bryologist 52 (4): 168, 1949 [1950].

*** Riccia trabutiana Steph., Rev. Bryol. 16 (5): 65, 1889.

*** Riccia violacea M.Howe, Ann. Missouri Bot. Gard. 2 (1/2): 51, 1915.

*** Riccia violacea var. laevis Jovet-Ast, Cryptog. Bryol. Lichénol. 10 (2): 100, 1989.

*** Riccia warnstorfii Limpr. ex Warnst., Verh. Bot. Vereins Prov. Brandenburg 27 (1): 85, 1886.

** subg. Ricciella (A.Braun) Boulay, Musc. France 2: 198, 1904. Bas.: Ricciella A.Braun, Flora 4 (2): 756, 1821.

*** Riccia cancellata Taylor, London J. Bot. 5: 414, 1846.

*** Riccia cincta Jovet-Ast, Cryptog. Bryol. 21 (4): 303, 2000.

*** Riccia cruciata Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 349, 1916.

*** Riccia eburnea Jovet-Ast, Cryptog. Bryol. 21 (4): 300, 2000.

*** Riccia hasskarliana Steph., Bull. Herb. Boissier 6 (5): 374 (49), 1898.

*** Riccia junghuhniana Nees et Lindenb., Syn. Hepat. 4: 609, 1846.

** Riccia junghuhniana var. simplex Schiffn., Hep. Fl. Buitenzorg: 14, 1900.

*** Riccia mangalorica Ahmad ex Jovet-Ast, Cryptog. Bryol. 24 (3): 223, 2003. Based on: Riccia mangalorica Ahmad, Curr. Sci. 11 (11): 433, 1942, nom. inval.

*** Riccia multifida (Steph.) Steph., Bull. Herb. Boissier 6 (5): 365 (40), 1898. Bas.: Ricciella multifida Steph., Hedwigia 28 (4): 273, 1889.

** Riccia polycarpa (Trab.) Jelenc, Bull. Trimestriel Geogr. Archeol. Oran 73 (228): 88, 1950. Bas.: Ricciella polycarpa Trab., Mém. Soc. Hist. Nat. Afrique N. 3: 36, 1933.

*** Riccia porosa Taylor, London J. Bot. 5: 416, 1846.

*** Riccia pullulans Jovet-Ast, Cryptog. Bryol. Lichénol. 18 (3): 183, 1997.

** sect. Ricciella (A.Braun) Bisch., Bem. Leberm.: 160, 1835. Bas.: Ricciella A.Braun, Flora 4 (2): 756, 1821.

* Riccia bahiensis Steph., Bull. Herb. Boissier 6 (5): 375 (50), 1898. 454

*** Riccia canaliculata Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 96, 1795 [1796].

*** Riccia chiapasensis Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 235, 1993.

*** Riccia crassifrons Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 570, 1885.

*** Riccia duplex Lorb. ex Müll.Frib., Hedwigia 80 (1/2): 100, 1941.

** Riccia duplex var. megaspora Na-Thalang, Brunonia 3 (1): 128, 1980.

*** Riccia dussiana Steph., Symb. Antill. (Urban) 3 (2): 275, 1902.

*** Riccia fluitans L., Sp. Pl. 1: 1139, 1753.

*** Riccia frostii Austin, Bull. Torrey Bot. Club 6 (3): 17, 1875.

* Riccia frostii var. crystallinoides Schiffn., Ann. K. K. Naturhist. Hofmus. 27: 503, 1913.

*** Riccia geissleriana Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 236, 1993.

*** Riccia hegewaldiana Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 238, 1993.

*** Riccia huebeneriana Lindenb., Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18 (1): 504d, 1836 [1837].

* Riccia huebeneriana subsp. sullivantii (Austin) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 457, 1992. Bas.: Riccia sullivantii Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 233, 1869.

*** Riccia jovet-astiae E.Vianna, Bol. Inst. Bioci. Univ. Fed. Rio Grande do Sul 38: 165, 1985.

*** Riccia limicola Jovet-Ast, Rev. Bryol. Lichénol. 44 (4): 422, 1978.

*** Riccia paraguayensis Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcvi, 1889 [1890].

*** Riccia paranaensis Hässel, Opera Lilloana 7: 228, 1962 [1963]. 455

*** Riccia perennis Steph., Bull. Herb. Boissier 6 (5): 372 (47), 1898.

*** Riccia purpurascens Lehm., Linnaea 4: 371, 1829.

** Riccia rhenana Lorb. ex Müll.Frib., Hedwigia 80 (1/2): 94, 1941.

** Riccia rhenana var. violacea M.F.Boiko, Chornom. Bot. J. 7: 93, 2011.

*** Riccia stricta (Lindenb.) Perold, Bothalia 20 (2): 197, 1990. Bas.: Riccia fluitans var. stricta Lindenb., Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18 (1): 444, 1836 [1837].

** sect. Spongodes Nees, Naturgesch. Eur. Leberm. 4: 391, 1838.

*** Riccia bullosa Link, Syn. hepat. eur: 119, 1829.

*** Riccia cavernosa Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 95, 1795 [1796].

*** Riccia crystallina L., Sp. Pl. 1: 1138, 1753.

*** Riccia cupulifera A.V.Duthie, Trans. Roy. Soc. South Africa 24 (2): 116, 1936.

*** Riccia garsidei Sim, Trans. Roy. Soc. South Africa 15 (1): 13, 1926.

*** Riccia moenkemeyeri Steph., Bot. Jahrb. Syst. 8 (2): 95, 1886.

*** Riccia rubricollis Garside et A.V.Duthie ex Perold, Bothalia 21 (1): 51, 1991.

*** Riccia volkii S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 271, 1957.

*** Riccia vulcanicola Eb.Fisch., Trop. Bryol. 8: 70, 1993.

** subg. Thallocarpus (Lindb.) Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 220, 1993. Bas.: Thallocarpus Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 377, 1874.

*** Riccia curtisii (Austin) Austin, Bull. Torrey Bot. Club 6 (52): 305, 1879. Bas.: Cryptocarpus curtisii Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 231, 1869.

** Riccia leptothallus R.M.Schust., J. Hattori Bot. Lab. 71: 271, 1992.

*** Riccia perssonii Sultan Khan, Svensk Bot. Tidskr. 49 (3): 433, 1955.

** subg. Triseriata Jovet-Ast, Cryptog. Bryol. Lichénol. 17 (2): 132, 1996.

*** Riccia singularis Jovet-Ast, Cryptog. Bryol. Lichénol. 17 (2): 127, 1996.

Incertae sedis

** Riccia abuensis Bapna, Trans. Brit. Bryol. Soc. 4 (2): 249, 1962.

** Riccia acutisulca Steph., Sp. Hepat. (Stephani) 6: 1, 1917.

* Riccia amboinensis Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 44, 1890.

** Riccia aravalliensis Pandé et Udar, J. Indian Bot. Soc. 36 (3): 249, 1957.

* Riccia arnellii Sultan Khan, Bryologist 60 (1): 29, 1957.

** Riccia asprella Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1059, 1888.

* Riccia asservanda De Not. ex Lamothe, Rech. Anat. Taxinom. Gamét. Marchantiales: 138, 1919.

** Riccia attenuata Pandé, Proc. Natl. Inst. Sci. India B 25 (2): 92, 1959.

* Riccia balansae Steph., Bull. Herb. Boissier 6 (5): 370 (45), 1898.

* Riccia bialbistrata Hässel, Opera Lilloana 7: 243, 1962 [1963]. 456

** Riccia biokoensis Perold, Nova Hedwigia 64 (1/2): 244, 1997.

** Riccia blackii Na-Thalang, Brunonia 3 (1): 81, 1980.

*** Riccia caroliniana Na-Thalang, Brunonia 3 (1): 72, 1980.

** Riccia cartilaginosa Steph., Hedwigia 28 (4): 272, 1889.

* Riccia chartacea K.I.Goebel, Organogr. Pfl., ed. 2, 2 (1): 630, 1915.

** Riccia chinensis Herzog, Symb. Sin. 5: 1, 1930.

** Riccia collata Na-Thalang, Brunonia 3 (1): 122, 1980.

** Riccia compacta Garside, Trans. Roy. Soc. South Africa 27 (1): 17, 1939.

** Riccia convexa Steph., Sp. Hepat. (Stephani) 6: 2, 1917.

* Riccia coronata Sim, Trans. Roy. Soc. South Africa 15 (1): 9, 1926. 457

** Riccia crassa Steph., Bull. Herb. Boissier 6 (5): 376 (51), 1898.

** Riccia crenatodentata O.H.Volk, Nova Hedwigia 46 (1/2): 27, 1988.

** Riccia delavayi Steph., Bull. Herb. Boissier 6 (5): 367 (42), 1898.

** Riccia deserticola Steph., Bull. Herb. Boissier 6 (5): 373 (48), 1898.

*** Riccia erubescens Perold, J. Bryol. 16 (3): 371, 1991.

** Riccia esulcata Steph., Sp. Hepat. (Stephani) 6: 2, 1917.

** Riccia fertilissima Steph., Sp. Hepat. (Stephani) 6: 2, 1917.

* Riccia gemmifera O.H.Volk, Nova Hedwigia 39: 131, 1984. 458

** Riccia grollei Udar, Curr. Sci. 34 (4): 126, 1965. Nom. nov. pro Riccia tuberculata Pandé et Udar, Proc. Natl. Inst. Sci. India B 24 (2): 83, 1958, nom. illeg.

** Riccia handelii Schiffn., Symb. sin. 2: 81, 1937.

** Riccia hawaiiensis Hürl., Phytologia 61 (5): 339, 1986.

** Riccia indica Udar et A.Gupta, Proc. V Indian Geophytol. Conf., Special Publ.: 307, 1984.

** Riccia indira-gandhiensis Dabhade et A.Hasan, J. Bombay Nat. Hist. Soc. 83 (2): 400, 1986.

* Riccia intermedia Roum., Mém. Soc. Arts Sci. Carcassonne 5: 198, 1888. 459

** Riccia jodhpurensis Bapna, Bot. Not. 114 (2): 181, 1961.

** Riccia kirinensis C.Gao et K.C.Chang, Acta Phytotax. Sin. 16 (4): 117, 1978.

*** Riccia laxisquamata (Steph.) Steph., Bull. Herb. Boissier 6 (5): 371 (46), 1898. Bas.: Ricciella laxisquamata Steph., Bot. Jahrb. Syst. 20 (3): 299, 1895.

** Riccia liaoningensis C.Gao et K.C.Chang, Acta Phytotax. Sin. 16 (4): 113, 1978.

** Riccia linearis (Schiffn.) Steph., Bull. Herb. Boissier 6 (5): 371 (46), 1898. Bas.: Ricciella linearis Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 43, 1890.

** Riccia luticola Na-Thalang, Brunonia 3 (1): 123, 1980.

** Riccia mamrensis Perold, Cryptog. Bryol. 26 (1): 68, 2005.

* Riccia marginata Lindb., Meddel. Soc. Fauna Fl. Fenn. 1: 106, 1876 [1877]. 460

** Riccia melanospora Kashyap, Liverworts W. Himal. 1: 94, 1929.

** Riccia miyakeana Schiffn., Österr. Bot. Z. 49 (11): 386, 1899.

** Riccia muscicola Steph., Hedwigia 24 (1): 4, 1885.

*** Riccia nigerica E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 225, 1957.

* Riccia nigrescens Mont., Voy. Amér. Mérid., Bot. 7 (1): 15, 1839. 461

** Riccia nipponica S.Hatt., J. Hattori Bot. Lab. 9: 38, 1953.

** Riccia novo-hannoverana Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 44, 1890.

** Riccia numeensis Steph., Bull. Herb. Boissier 6 (4): 343 (35), 1898.

* Riccia obtusa Meijer, J. Hattori Bot. Lab. 20: 113, 1958. 462

*** Riccia oerstediana Lindenb. et Hampe, Linnaea 24 (3): 304, 1851 [1852].

** Riccia pandei Udar, J. Indian Bot. Soc. 38 (1): 149, 1959.

*** Riccia papillispora Steph., Bull. Herb. Boissier 6 (4): 334 (26), 1898.

** Riccia papulosa (Steph.) Steph., Bull. Herb. Boissier 6 (5): 377 (52), 1898. Bas.: Ricciella papulosa Steph., Hedwigia 28 (4): 273, 1889.

** Riccia papulosa var. variabilis Na-Thalang, Brunonia 3 (1): 112, 1980.

** Riccia pathankotensis Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 349, 1916.

* Riccia perthiana Steph. ex K.I.Goebel, Organogr. Pfl., ed. 2, 2 (1): 630, 1915.

* Riccia prominens Meijer, J. Hattori Bot. Lab. 20: 111, 1958. 463

** Riccia pseudofluitans C.Gao et K.C.Chang, Acta Phytotax. Sin. 16 (4): 116, 1978.

** Riccia pubescens S.Hatt., Nat. Sci. Mus. 14 (6): 141, 1943.

*** Riccia radiata Perold, Bothalia 34 (1): 23, 2004.

** Riccia rechingeri Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 288, 1907.

* Riccia reticulatula Udar, Bull. Bot. Soc. Univ. Saugar 13: 49, 1961.

** Riccia rorida Na-Thalang, Brunonia 3 (1): 101, 1980.

* Riccia saharensis Steph. ex Jovet-Ast, Rev. Bryol. Lichénol. 26 (1/2): 62, 1957. 464

** Riccia satoi S.Hatt., Bot. Mag. (Tokyo) 62 (733/734): 109, 1949.

* Riccia schroederi Steph., 52 (5): 304, 1912.

*** Riccia schweinfurthii Steph., Bull. Herb. Boissier 6 (4): 339 (31), 1898.

** Riccia sibayenii Perold, Bothalia 31 (1): 151, 2001.

*** Riccia somaliensis Perold, J. Bryol. 16 (3): 367, 1991.

** Riccia spongiosula Na-Thalang, Brunonia 3 (1): 113, 1980.

* Riccia subtilis (Steph.) Steph., Bull. Herb. Boissier 6 (5): 364 (39), 1898. Bas.: Ricciella subtilis Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 31, 1897.

** Riccia sumatrana Meijer, J. Hattori Bot. Lab. 20: 114, 1958.

*** Riccia symoensii Vanden Berghen, Explor. Hydrobiol. Lac Bangweolo Luapula: 191, 1972.

** Riccia tasmanica Steph. ex Rodway, Tasm. Bryoph.: 4, 1917.

*** Riccia tomentosa O.H.Volk et Perold, Bothalia 20 (1): 25, 1990.

** Riccia treubiana Steph., Bull. Herb. Boissier 6 (4): 323 (15), 1898.

* Riccia treubiana var. subrubescens Schiffn., Hep. Fl. Buitenzorg: 16, 1900.

* Riccia triangularis Steph., Bull. Mus. Natl. Hist. Nat. 18 (2): 116, 1912.

* Riccia tuberculata Poir., Encycl. (Lamarck) 6: 199, 1804. 465

** Riccia udarii Kanwal, J. Indian Bot. Soc. 58 (3): 282, 1979.

* Riccia velenovskyi Kavina, Arch. Přír. Výzk. Čech 16 (2): 75, 1915. 466

** Riccia velimalaiana A.E.D.Daniels et P.Daniel, Bull. Bot. Surv. India 44 (1/4): 139, 2002 [2003].

** Riccia victoriensis Steph., Bull. Herb. Boissier 6 (5): 370, 1898.

** Riccia weymouthiana Steph. ex Rodway, Tasm. Bryoph.: 5, 1917.

** Riccia wichurae Steph., Bull. Herb. Boissier 6 (4): 330 (22), 1898.

** Ricciocarpos Corda, Gen. hepat.: 651, 1829.

*** Ricciocarpos natans (L.) Corda, Gen. hepat.: 651, 1829. Bas.: Riccia natans L., Syst. Nat., ed. 10., 2: 1339, 1759.

*** Targioniaceae Dumort.

by D.G. Long

*** Targionia L., Sp. Pl. 1: 1136, 1753.

** subg. Prototargionia R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 69, 1992.

** Targionia stellaris (Müll.Frib.) Hässel, Opera Lilloana 7: 74, 1962 [1963]. Bas.: Grimaldia stellaris Müll.Frib., Feddes Repert. Spec. Nov. Regni Veg. 58: 61, 1955.

** subg. Targionia

*** Targionia hypophylla L., Sp. Pl. 1: 1136, 1753.

** Targionia hypophylla subsp. linealis W.Frey et Kürschner, Nova Hedwigia 57 (1/2): 127, 1993.

** Targionia lorbeeriana Müll.Frib., Hedwigia 79 (1/2): 78, 1940.

Incertae sedis

* Targionia dioica Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 154, 1898. 467

* Targionia elongata Bisch., Syn. Hepat. 4: 576, 1846.

* Targionia fiorii Gola, Ann. Bot. (Rome) 13 (1): 62, 1914.

* Targionia formosica Horik., J. Jap. Bot. 11: 499, 1935.

* Targionia indica Udar et A.Gupta, Geophytology 13 (1): 83, 1983.

*** Wiesnerellaceae Inoue

by D.G.Long

*** Wiesnerella Schiffn., Österr. Bot. Z. 46 (3): 86, 1896.

*** Wiesnerella denudata (Mitt.) Steph., Bull. Herb. Boissier 7 (5): 382 (154), 1899. Bas.: Dumortiera denudata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 125, 1860 [1861].

* Wiesnerella fasciaria C.Gao et K.C.Chang, Acta Bot. Yunnan. 3 (4): 391, 1981.

*** Neohodgsoniaceae D.G.Long

by D.G. Long

*** Neohodgsonia Perss., Bot. Not. 107 (1): 40, 1954. Nom. nov. pro Hodgsonia Perss., Hodgsonia Leafl. Stockholm: 1, 1953.

*** Neohodgsonia mirabilis (Perss.) Perss., Bot. Not. 107 (1): 40, 1954. Bas.: Hodgsonia mirabilis Perss., Hodgsonia Leafl. Stockholm: 1, 1953.

*** Monocarpaceae D.J.Carr ex Schelpe

by D.G. Long

*** Monocarpus D.J.Carr, Austral. J. Bot. 4 (2): 176, 1956.

*** Monocarpus sphaerocarpus D.J.Carr, Austral. J. Bot. 4 (2): 176, 1956.

*** Riellaceae Engl.

by J. G. Segarra-Moragues and F. Puche

*** Austroriella Cargill et J.Milne, Polish Bot. J. 58 (1): 72, 2013.

*** Austroriella salta J.Milne et Cargill, Polish Bot. J. 58 (1): 72, 2013.

*** Riella Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 11, 1852.

*** subg. Riella

*** Riella alatospora Wigglesw., J. Linn. Soc., Bot. 51 (339): 317, 1937.

*** Riella americana M.Howe et Underw., Bull. Torrey Bot. Club 30 (4): 218, 1903.

* Riella battandieri Trab., Rev. Bryol. 13 (3): 35, 1886.

*** Riella bialata Trab., Rev. Bryol. 35 (4): 96, 1908.

*** Riella capensis Cavers, Rev. Bryol. 30 (5): 81, 1903.

*** Riella choconensis Hässel, Symp. Biol. Hung. 35: 341, 1987.

* Riella cyrenaica Maire, Bull. Soc. Hist. Nat. Afrique N. 30 (5): 312, 1939.

*** Riella echinospora Wigglesw., J. Linn. Soc., Bot. 51 (339): 321, 1937.

* Riella gallica Balansa ex Trab., Rev. Gén. Bot. 3 (35): 450, 1891.

*** Riella halophila Banwell, Trans. Brit. Bryol. Soc. 1 (5): 475, 1951.

*** Riella helicophylla (Bory et Mont.) Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 12, 1852. Bas.: Duriaea helicophylla Bory et Mont., Ann. Sci. Nat. Bot. (sér. 3) 1: 229, 1844.

* Riella helicophylla var. macrocarpa P.Allorge, Sched. Br. Iber. (ser. 2): 4, 1929.

* Riella indica Steph. ex Kashyap, J. Bombay Nat. Hist. Soc. 25 (2): 279, 1917.

*** Riella notarisii (Mont.) Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 12, 1852. Bas.: Sphaerocarpos notarisii Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 39, 1838. 468

** Riella numidica Trab., Bull. Soc. Hist. Nat. Afrique N. 25 (9): 391, 1934 [1935].

*** Riella pampae Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 5 (9): 207, 1979.

*** Riella parisii Gottsche, Hepat. Eur., Leberm. 38-39: no. 375, 1867.

*** Riella purpureospora Wigglesw., J. Linn. Soc., Bot. 51 (339): 312, 1937.

* Riella reuteri Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 12, 1852.

* Riella sersuensis Trab., Bull. Soc. Hist. Nat. Afrique N. 25 (9): 392, 1934 [1935].

** Riella spiculata J.Taylor, Kew Bull. 9 (1): 45, 1954.

*** Riella trigonospora Segarra et Puche, S. African J. Bot. 94: 175, 2014.

*** Riella undulata Hässel, Symp. Biol. Hung. 35: 341, 1987.

*** subg. Trabutiella Porsild, Bot. Tidsskr. 24 (3): 327, 1902.

*** Riella affinis M.Howe et Underw., Bull. Torrey Bot. Club 30 (4): 221, 1903.

*** Riella cossoniana Trab., Atlas fl. Alger 1: 6, 1886.

*** Riella echinata (Müll.Frib.) Segarra, Puche et Sabovlj., Phytotaxa 159 (3): 165, 2014. Bas.: Riella cossoniana var. echinata Müll.Frib., Rev. Bryol. Lichénol. 22 (3/4): 132, 1953 [1954].

*** Riella gamundiae Hässel, Rev. Bryol. Lichénol. 38 (3/4): 580, 1972 [1973].

*** Riella heliospora Segarra, Puche et Sabovlj., Syst. Bot. 37 (2): 315, 2012.

*** Riella mediterranea Segarra, Puche, Sabovlj., M.Infante et Heras, Phytotaxa 159 (3): 170, 2014.

*** Sphaerocarpaceae Heeg

by D. Long

*** Geothallus Campb., Bot. Gaz. 21 (1): 13, 1896.

*** Geothallus tuberosus Campb., Bot. Gaz. 21 (1): 13, 1896.

*** Sphaerocarpos Boehm., Def. gen. pl., ed. 3: 501, 1760.

** subg. Austrosphaerocarpos R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 813, 1992.

*** Sphaerocarpos stipitatus Bisch. ex Lindenb., Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18 (1): 504i, 1836 [1837].

** subg. Sphaerocarpos

*** Sphaerocarpos michelii Bellardi, App. fl. pedem.: 52, 1792.

** Sphaerocarpos texanus Austin, Bull. Torrey Bot. Club 6 (30): 158, 1877.

Incertae sedis

*** Sphaerocarpos cristatus M.Howe, Mem. Torrey Bot. Club 7: 66, 1899.

*** Sphaerocarpos donnellii Austin, Bull. Torrey Bot. Club 6 (30): 157, 1877.

*** Sphaerocarpos drewiae Wigglesw., Univ. Calif. Publ. Bot. 16 (3): 129, 1929.

*** Sphaerocarpos europaeus Lorb., Jahrb. Wiss. Bot. 80: 665, 1934. 469

*** Sphaerocarpos hians Haynes, Bull. Torrey Bot. Club 37 (5): 225, 1910.

** Sphaerocarpos muccilloi E.Vianna, Lindbergia 7 (1): 58, 1981.

Names in genera not currently accepted

The following taxa in unsupported genera are all poorly understood. We list them here rather than making new combinations for names we do not know the status of.

Acrostolia Dumort., Recueil Observ. Jungerm.: 26, 1835. 470

* Acrostolia alata (Gottsche et Rabenh.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877. Bas.: Pseudoneura alata Gottsche et Rabenh., Hepat. Eur., Leberm. 56-57: no. 560, 1873.

* Acrostolia brevifolia (Gottsche et Rabenh.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877. Bas.: Pseudoneura brevifolia Gottsche et Rabenh., Hepat. Eur., Leberm. 56-57: no. 560, 1873.

Aphanolejeunea A.Evans, Bull. Torrey Bot. Club 38 (6): 272, 1911.

* Aphanolejeunea lancifera R.M.Schust., Phytologia 45 (5): 434, 1980. 471

* Aphanolejeunea minima Tixier, Ann. Fac. Sci. Yaoundé 20: 7, 1975. 472

Aspiromitus Steph., Sp. Hepat. (Stephani) 5: 957, 1916. 473

* Aspiromitus asper Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 402, 1955.

* Aspiromitus bullosus Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 403, 1955.

* Aspiromitus crenatifrons Steph., Sp. Hepat. (Stephani) 5: 968, 1916.

* Aspiromitus lobatus Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 405, 1955.

* Aspiromitus squamulosus Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 407, 1955.

Crossotolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893. Bas.: Lejeunea subg. Crossotolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 161, 1884. 474

* Crossotolejeunea curvifolia Steph., Hedwigia 35 (3): 75, 1896. 475

Eulejeunea Steph., Hedwigia 27 (2): 60, 1888. 476

* Eulejeunea setulosa Steph., Sp. Hepat. (Stephani) 6: 421, 1923.

* Eulejeunea subpililoba Steph., Sp. Hepat. (Stephani) 6: 420, 1923. 477

Euosmolejeunea (Spruce) Steph., Hedwigia 28 (3): 170, 1889. Bas.: Lejeunea subg. Euosmolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 241, 1884. 478

* Euosmolejeunea parvistipula (Lindenb. et Gottsche) Steph., Hedwigia 29 (1): 80, 1890. Bas.: Lejeunea parvistipula Lindenb. et Gottsche, Syn. Hepat. 5: 761, 1847.

* Euosmolejeunea tenerrima (Nees) Steph., Sp. Hepat. (Stephani) 5: 589, 1914. Bas.: Jungermannia sordida var. tenerrima Nees, Fl. Bras. (Martius) 1 (1): 363, 1833. 479

Fimbraria Nees, Horae Phys. Berol.: 44, 1820 nom. illeg. 480

* Fimbraria gigantea Steph., Bull. Herb. Boissier 7 (2): 93 (106), 1899. 481

* Fimbraria incrassata Steph., Bull. Herb. Boissier 7 (2): 87 (100), 1899. 482

* Fimbraria kamerunensis Steph., Sp. Hepat. (Stephani) 6: 14, 1917. 483

* Fimbraria pirottae Gola, Ann. Bot. (Rome) 13 (1): 64, 1914.

Hygrolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 124, 1893. Bas.: Lejeunea subg. Hygrolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 230, 1884. 484

* Hygrolejeunea cubensis Steph., Sp. Hepat. (Stephani) 5: 533, 1914.

* Hygrolejeunea harpaphylla Steph., Sp. Hepat. (Stephani) 5: 552, 1914.

* Hygrolejeunea pacifica Steph., Sp. Hepat. (Stephani) 6: 411, 1923.

* Hygrolejeunea parvicalycina Steph., Hedwigia 35 (3): 103, 1896.

* Hygrolejeunea parvistipula Steph., Sp. Hepat. (Stephani) 5: 568, 1914.

* Hygrolejeunea patellirostris Steph., Hedwigia 35 (3): 103, 1896. 485

* Hygrolejeunea staudtiana Steph., Sp. Hepat. (Stephani) 5: 528, 1914. 486

Hypenantron Corda, Gen. hepat.: 648, 1829. 487

* Hypenantron brachypus Steph. ex Lamothe, Rech. Anat. Taxinom. Gamét. Marchantiales: 107, 1919.

* Hypenantron brasiliense Steph. ex Lamothe, Rech. Anat. Taxinom. Gamét. Marchantiales: 104, 1919.

Jamesoniella (Spruce) Carrington, Cat. Brit. Moss. Hepat.: 25, 1881. Bas.: Jungermannia subg. Jamesoniella Spruce, J. Bot. 14: 230, 1876. 488

* Jamesoniella convoluta Steph., Sp. Hepat. (Stephani) 6: 433, 1924.

Kingiolejeunea H.Rob., Bryologist 70 (1): 53, 1967. 489

* Kingiolejeunea guayanensis H.Rob., Bol. Soc. Venez. Ci. Nat. 32 (132/133): 259, 1976.

Leptocolea (Spruce) A.Evans, Bull. Torrey Bot. Club 38 (6): 261, 1911. Bas.: Lejeunea sect. Leptocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 294, 1884. 490

* Leptocolea sumatrana Herzog, Ann. Bryol. 5: 96, 1932. 491

Mastigobryum (Nees) Lindenb. et Gottsche, Syn. Hepat. 2: 214, 1845 nom. illeg. Bas.: Herpetium sect. Mastigobryum Nees, Naturgesch. Eur. Leberm. 3: 43, 1838. 492

* Mastigobryum aberrans Steph., Sp. Hepat. (Stephani) 6: 486, 1924.

* Mastigobryum asperum Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 121, 1914.

* Mastigobryum deningeri Herzog, Beih. Bot. Centralbl. 38 (2): 322, 1921.

* Mastigobryum karstenii Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 952 (502), 1908.

* Mastigobryum ledermannii Steph., Sp. Hepat. (Stephani) 6: 486, 1924.

* Mastigobryum londbergii Steph., Sp. Hepat. (Stephani) 6: 487, 1924.

* Mastigobryum longifolium Steph., Sp. Hepat. (Stephani) 6: 487, 1924.

* Mastigobryum minutitextum Steph., Sp. Hepat. (Stephani) 6: 487, 1924.

* Mastigobryum multidens Steph., Sp. Hepat. (Stephani) 6: 487, 1924.

* Mastigobryum muscicola Steph., Sp. Hepat. (Stephani) 6: 486, 1924.

* Mastigobryum nigricans Herzog, Beih. Bot. Centralbl. 38 (2): 322, 1921.

* Mastigobryum nipuranum Steph., Sp. Hepat. (Stephani) 6: 485, 1924.

* Mastigobryum palmicola Steph., Sp. Hepat. (Stephani) 6: 488, 1924.

* Mastigobryum rajanum Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 190, 1931.

* Mastigobryum ribehanum Steph., Sp. Hepat. (Stephani) 6: 488, 1924.

* Mastigobryum ruficaule Beauverd, Sp. Hepat. (Stephani) 6: 485, 1924.

* Mastigobryum schraderbergii Steph., Sp. Hepat. (Stephani) 6: 489, 1924.

* Mastigobryum squamulistipum Steph., Sp. Hepat. (Stephani) 6: 480, 1924.

* Mastigobryum subhyalinum Steph., Sp. Hepat. (Stephani) 6: 482, 1924.

* Mastigobryum venezuelanum Molk., Syn. hepat. jav.: 104, 1856 [1857].

* Mastigobryum vermiculare Herzog, Hedwigia 66 (6): 339, 1926.

Nemoursia Mérat, Ann. Agric. Franç. (ser. 4) 2 (7): 10, 1840.

* Nemoursia tuberculata Mérat, Ann. Agric. Franç. (ser. 4) 2 (7): 10, 1840. 493

Physocolea (Spruce) Steph., Sp. Hepat. (Stephani) 5: 863, 1916. Bas.: Lejeunea sect. Physocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 292, 1884. 494

* Physocolea tambillensis (Loitl.) Steph., Sp. Hepat. (Stephani) 5: 885, 1916. Bas.: Lejeunea tambillensis Loitl., Diagn. pl. nov.: 20, 1894. 495

Plectocolea (Mitt.) Mitt., Fl. vit.: 405, 1871 [1873]. Bas.: Solenostoma subg. Plectocolea Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864 [1865]. 496

* Plectocolea subamoena S.Winkl., Rev. Bryol. Lichénol. 42 (3): 821, 1976. 497

Polyotus Gottsche, Syn. Hepat. 2: 244, 1845 nom. illeg. 498

* Polyotus peckianus Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 224, 1869.

Schisma Dumort., Commentat. Bot. (Dumortier): 114, 1822 nom. illeg. 499

* Schisma orizabense (Gottsche) Steph., Sp. Hepat. (Stephani) 4: 19, 1909. Bas.: Sendtnera orizabensis Gottsche, Mexik. Leverm.: 139, 1863.

* Schisma uleanum Steph., Hedwigia 44 (4): 225, 1905.

Strepsilejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893. Bas.: Lejeunea sect. Strepsilejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 168, 1884. 500

* Strepsilejeunea apollinea (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 291, 1913. Bas.: Lejeunea apollinea Gottsche, Fragm. (Mueller): 64, 1880.

* Strepsilejeunea durelii Schiffn., Österr. Bot. Z. 49 (6): 206, 1899.

* Strepsilejeunea hamatifolia Steph., Sp. Hepat. (Stephani) 6: 396, 1923.

* Strepsilejeunea lanceolata (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 283, 1913. Bas.: Lejeunea lanceolata Gottsche, Syn. Hepat. 3: 353, 1845.

* Strepsilejeunea muscicola Herzog, Hedwigia 74 (2): 96, 1934.

* Strepsilejeunea novae-guineae Steph., Sp. Hepat. (Stephani) 6: 397, 1923.

* Strepsilejeunea obtusistipula Steph., Biblioth. Bot. 87 (2): 258, 1916.

* Strepsilejeunea papulifolia Steph., Biblioth. Bot. 87 (2): 259, 1916.

* Strepsilejeunea pectiniformis (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 285, 1913. Bas.: Lejeunea pectiniformis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 156, 1864.

* Strepsilejeunea renistipula Steph., Sp. Hepat. (Stephani) 5: 289, 1913.

* Strepsilejeunea vatovae Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 479, 1947.

Thysanolejeunea (Spruce) Steph., Hedwigia 31 (1): 20, 1892. Bas.: Lejeunea subg. Thysanolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 105, 1885.

* Thysanolejeunea africana Sim, Trans. Roy. Soc. South Africa 15 (1): 50, 1926. 501

Trachylejeunea (Spruce) Steph., Hedwigia 28 (4): 262, 1889 nom. rejic. Bas.: Lejeunea subg. Trachylejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 180, 1884.

* Trachylejeunea conifera Steph., Sp. Hepat. (Stephani) 5: 302, 1913.

* Trachylejeunea cristuliflora Steph., Hedwigia 35 (3): 137, 1896.

* Trachylejeunea englishii Steph., Bull. Misc. Inform. Kew 1899 (151/152): 126, 1899. 502

* Trachylejeunea jamaicensis Pearson, Ann. Bryol. 4: 98, 1931.

* Trachylejeunea kusaiensis Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 147, 1965.

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